Crassignatha thamphra Y. Lin & S. Li, 2020

Li, Ya, Lin, Yucheng & Li, Shuqiang, 2020, A review of Crassignatha (Araneae, Symphytognathidae), ZooKeys 988, pp. 63-128 : 63

Crassignatha thamphra Y. Lin & S. Li sp. nov. Figs 32 , 38

Type material.

Holotype ♀ (IZCAS-Ar 41035), Thailand: Khon Kaen Province, Phu Pha Man District, Phu Pha Man Subdistrict, Tham Phra Cave (16.66603°N, 101.89623°E; 262 m), 10.XI.2016, H. Zhao, Y. Li and Z. Chen leg.; 1♀ (NHMSU-HA089) used for sequencing, GenBank: MT992003, same data as for preceding.

Diagnosis.

This species differs other Crassignatha species, except C. yinzhi , by the copulatory ducts diagonally connected to the copulatory opening and not fused before reaching copulatory opening. It can be easily distinguished from C. yinzhi by the larger spermathecae separated by less than their diameter and the tighter turns of the copulatory ducts at center of the vulva (Fig. 32G vs. Fig. 37G ).

Description.

Female (holotype). Total length 0.64. Carapace 0.32 long, 0.32 wide, 0.28 high. Clypeus 0.10 high. Sternum 0.20 long, 0.20 wide. Abdomen 0.44 long, 0.48 wide, 0.48 high. Length of legs: I 0.92 (0.28, 0.12, 0.20, 0.16, 0.16); II 0.74 (0.22, 0.10, 0.14, 0.12, 0.16); III 0.64 (0.16, 0.10, 0.12, 0.12, 0.14); IV 0.84 (0.24, 0.12, 0.18, 0.12, 0.18).

Somatic characters (Fig. 32A-C ). Coloration: prosoma yellowish brown. Legs dark brownish. Abdomen black with faint, sclerotized dots. Prosoma: carapace nearly pear shaped, surface indistinctly textured. Cephalic area elevated. ALE slightly protruded. PER straight. Chelicerae bears sparse, short setae anteriorly. Labium tongue shaped. Sternum heart shaped, slightly plump, surface smooth, truncated posteriorly. Legs: covered with setae and bristles. Abdomen: abdominal shape irregular and surface rugose (caused by alcohol immersion). Spinnerets weakly sclerotized.

Epigyne (Fig. 32D-G ): epigynal area slightly sclerotized, bears a few long setae. Scape protruded, longer slightly than wide. Internal structures faintly visible via translucent tegument. Spermathecae separated by ~ ½ their diameter. Fertilization ducts thin, slender, originating from posteromedial margin of spermathecae, forming a U-shape. Copulatory ducts long, connected to the posterior surface of spermathecae, curving twice below spermathecae, linked diagonally, and fused to copulatory opening.

Male. Unknown.

Etymology.

The specific name is derived from the type locality; noun in apposition.

Distribution.

Thailand (Fig. 38 ).

Figures

Figure 32. Crassignatha thamphra sp. nov. A female habitus, dorsal B female habitus, ventral C female habitus, lateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.50 mm (A-C); 0.10 mm (D-G).

Figure 37. Crassignatha yinzhi Miller, Griswold & Yin, 2009 A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G).

Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi.