Alcochera flavoclypeata Sheng & Sun, 2021

Sun, Shu-Ping, Li, Tao, Zong, Shi-Xiang & Sheng, Mao-Ling, 2021, The genus Alcochera Förster (Hymenoptera, Ichneumonidae, Ctenopelmatinae) in China with a key to world species, Zootaxa 5023 (2), pp. 273-283 : 275-277

Alcochera flavoclypeata Sheng & Sun , sp. nov.

Figures 1–11

Material examined. Holotype, Female, CHINA: Labagoumen Natural Reserve, Huairou , Beijing, 5 September 2016, Shi-Xiang Zong ( IT) . Paratypes: 2 females 1 male, same data as holotype except 8–15 August 2016 .

Diagnosis. Malar space approximately 0.4 × as long as basal width of mandible. Postocellar line 0.6–0.7 × as long as ocular-ocellar line. Forewing areolet ( Fig. 8 ) with short stalk, receiving 2m-cu 0.8 × distance from 2rs-m to 3rs-m. Propodeum ( Fig. 9 ) with dense gray white hairs. Area basalis and area superomedia almost smooth, shiny. Subapical portion of ovipositor sheath ( Fig. 11 ) distinctly widened. Basal portion of ovipositor stout, apical slender, dorsal subapical notch ( Fig. 11 ) deep and wide. Head except clypeus, mesosoma and all tergites almost entirely black.

Description. Female ( Fig. 1 ). Body length 8.5–9.0 mm. Fore wing length 7.0– 7.5 mm.

Head. Inner orbits almost parallel, hardly emarginate opposite antennal socket. Face ( Fig. 2 ) 1.7–1.8 × as wide as long, slightly convex medially; shagreened, with dense indistinct punctures, upper margin with small median tubercle. Clypeus ( Fig. 2 ) 2.4–2.6 × as wide as long, smooth, shiny, evenly convex apically, with sparse punctures. Mandible with sparse shallow punctures and yellowish white hairs; lower tooth distinctly longer than upper tooth. Malar space approximately 0.4 × as long as basal width of mandible. Gena ( Fig. 3 ) shagreened, with shallow indistinct punctures; evenly convergent backwardly. Vertex ( Fig. 4 ) with texture as that of gena. Postocellar line 0.6–0.7 × as long as ocular-ocellar line. Frons evenly slightly concave, with dense fine punctures. Antenna with 36–37 flagellomeres; ratio of length from first to fifth flagellomeres approximately: 1.8:1.0:0.9:0.8:0.8. Occipital carina reaching hypostomal carina above base of mandible.

Mesosoma. Along anterior margin of pronotum ( Fig. 5 ) with dense fine punctures and indistinct oblique wrinkles; anterior portion with dense punctures, distance between punctures 0.2–1.0 × diameter of puncture; lateral concavity with strong oblique transverse wrinkles; along posterior margin with short transverse wrinkles; upperposterior portion almost shiny, with distinct punctures, distance between punctures 0.2–2.5 × diameter of puncture. Mesoscutum ( Fig. 6 ) with dense punctures; notauli weak, reaching to middle of mesoscutum. Scutoscutellar groove almost smooth, shiny. Scutellum ( Fig. 9 ) slightly convex, with dense indistinct punctures. Postscutellum with distinct uneven punctures. Mesopleuron ( Fig. 7 ) almost shiny, with dense punctures (except upper posterior portion), distance between punctures 0.2–3.0 × diameter of puncture; speculum smooth, shiny; area beneath speculum slightly rough, with short irregular transverse wrinkles. Upper end of epicnemial carina distant from front edge of mesopleuron, approximately reaching 0.5 distance to subtegular ridge. Metapleuron evenly convex, with relatively dense punctures, distance between punctures 0.2–2.5 × diameter of puncture; lower posterior portion with short oblique wrinkles. Wings brownish, hyaline. Fore wing with 1cu-a distal to 1/M approximately by 0.3 × length of 1cu-a. Areolet ( Fig. 8 ) oblique quadrilateral, with short stalk, receiving 2m-cu approximately 0.8 × distance from 2rs-m to 3rs-m. 2-Cu slightly longer than 2cu-a. Hind wing 1-cu approximately 1.5 × as long as cu-a. Claw simple. Ratio of length of hind tarsomeres from first to fifth approximately: 4.0:2.0:1.7:0.9:1.0. Propodeum ( Fig. 9 ) evenly convex, with dense gray white hairs. Lateromedian longitudinal carinae between basal margin and posterior transverse carina complete, weak medially. Area basalis and area superomedia combined, almost smooth, shiny. Sublateral basal portion (about area externa) shiny, with distinct punctures; basal lateral portion indistinctly punctate; apicomedian portion with irregular short transverse wrinkles and indistinct punctures. Propodeal spiracle almost circular, distance to lateral longitudinal carina longer than to pleural carina.

Metasoma. First two tergites ( Fig. 10 ) and basal portions of tergites 3 and 4 with dense punctures. First tergite approximately 1.4 × as long as posterior width, evenly strongly widened posteriorly, spiracle circular, small, located approximately at middle of first tergite. Second tergite approximately 0.7 × as long as posterior width. Apical portions of tergites 3 and 4 with fine sparse punctures. Third tergite approximately 0.74 × as long as posterior width. Five and subsequent tergites ( Fig. 11 ) with fine indistinct punctures. Subapical portion of ovipositor sheath widened. Basal portion of ovipositor stout, apical portion elongate, dorsal subapical notch ( Fig. 11 ) deep and wide.

Coloration ( Fig. 1 ). Black, except for following: ventrobasal portion of antenna brownish yellow, flagellomeres 14–19 white. Clypeus, mandible except teeth, maxillary palpi, labial palpi, margin of tegula, upper-posterior corner of pronotum, scutellum yellow white. Fore and middle femora blackish brown, apexes, tibiae except apical portions, subbasal portion of hind tibia, all tarsomeres 1–4 yellow. Apical margins of tergites 1–4 slightly yellowish brown narrowly. Pterostigma and veins brownish black.

Male. Body length 8.5 mm. Fore wing length 7.0 mm. Antenna with 37 flagellomeres. Flagellomeres 14–23 white. Face mainly yellow, upper median longitudinal and lateral spots black. Maxillary and labial palpi almost entirely white. Fore and middle femora red brown. Others almost the same as female.

Distribution. CHINA: Beijing.

Etymology. The specific name is derived from the yellow clypeus.

Remarks. The new species is similar to A. albicervicalis Sheng & Fan 1995 , but can be distinguished from the latter by the following combination of characters: forewing areolet ( Fig. 8 ) receiving vein 2m-cu distinctly basal of its lower posterior corner; 2rs-m approximately 0.88 × as long as 3rs-m; lateromedian longitudinal carina of propodeum ( Fig. 9 ) complete from anterior margin to posterior transverse carina; tegula mainly black; hind femur ( Fig. 1 ) entirely black. A. albicervicalis : areolet ( Fig. 12 ) joining vein 2m-cu at its lower posterior corner; 2rs-m approximately 0.45 × as long as 3rs-m; lateromedian longitudinal carina of propodeum mostly entirely absent; tegula yellow; at least basal half of hind femur reddish brown.

Figures

FIGURES 1–5. Alcochera flavoclypeata Sheng & Sun, sp. nov. Holotype. Female. 1 Habitus, lateral view 2 Head, anterior view 3 Head, lateral view 4 Head, dorsal view 5 Pronotum, lateral view.

FIGURES 6–9. Alcochera flavoclypeata Sheng & Sun, sp. nov. Holotype. Female. 6 Mesoscutum 7 Mesosoma, lateral view 8 Areolet 9 Scutellum, postscutellum and propodeum, dorsal view.

FIGURES 10–12. Alcochera flavoclypeata Sheng & Sun, sp. nov. Holotype. Female. 10 Metasoma, dorsal view 11 Apical portion of metasoma, lateral view 12 Alcochera albicervicalis Sheng & Fan 1995. Holotype. Female. Areolet.