Bernaya grovesi, SQUIRES AND DEMETRION, 1992

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 12-14

BERNAYA GROVESI SQUIRES AND DEMETRION, 1992

Figs. 5A–C

Bernaya (Protocypraea) grovesi Squires and Deme- trion, 1992. p. 31, figs. 74-76. Groves, 1993. p. 11. Groves, 1994b. p. 12. Groves, 1997. p. 7 [as B. (Protocypraea) grovesi ].

Protocypraea grovesi (Squires and Demetrion) . Lorenz, 2017. p. 205.

Holotype and Type Locality— IGM 5172 (Figs. 5A–C), length 17 mm, width 12 mm, height 7.8 mm. LACMIP Lo- cality 41220b [ex CSUN Locality 1220b], Bateque Forma- tion, near Laguna San Ignacio, Baja California Sur, México.

Paratype — IGM 5173 , length 16 mm, width 10.5 mm, height 8 mm; same locality as the holotype .

Referred Specimen— A single, slightly crushed speci- men ( LACMIP 41653.6 , LACMIP Type 14937) measuring length 14.3 mm, width 9.2 mm, height 5.8 mm from LACMIP Locality 16655 [= LACMIP Locality 41653; ex CSUN Locality 1563], Crescent Formation, lower Eocene (“Capay” Stage), Larch Mountain area , Black Hills , Thur- ston County , southwestern Washington that strongly resembles B. grovesi is here noted .

Occurrence— Lower to middle Eocene (“Capay

Figure 5 (previous page) A–C. Bernaya grovesi Squires and Demetrion, 1992 , Bateque Formation (Eocene, “Ca- pay Stage”), Mesa La Salina, Baja California Sur, México. Basal (A), dorsal (B), and right (labral) (C) views, all 2.6x, length 17 mm, width 12 mm, height 7.7 mm, of holotype IGM 5172, LACMIP Locality 401220a (ex CSUN Locality 1220b). D–F. Bernaya kaylinae n. sp., Llajas Formation, (Eocene, “Domengine Stage”), Las Llajas Canyon, north side Simi Valley, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F) views, 3.8x, length 25.6 mm, width 17.3 mm, height 12.9 mm, of holotype, LACMIP 40374.80, LACMIP Type 14938 ( LACMIP Locality 40374). G–I. Bernaya squiresi Groves, 2011 , unnamed Eocene strata (“Capay Stage”), southwest end of Discovery Bay, Jefferson County, Washington. Basal (G), dorsal (H), and right (labral) side (I) views, 0.97x, length 52.7 mm, width 40 mm, height 26.3 mm, of holotype LACMIP 22341.1, LACMIP Type 13644, LACMIP Locality 22341. J–L. Bernaya sp. , unnamed Paleocene strata (Thanetian Stage), near Lower Lake, Lake County, northern California. Basal (J), dorsal (K), and right (labral) side (L) views, 1.7x, length 33.8 mm, width 23 mm, height 18.3 mm, hypotype LACMIP 7047.72, LACMIP Type 14923 ( LACMIP Locality 7047). M–O. Protocypraea? simiensis ( Nelson, 1925) , Santa Susana Formation (Thanetian Stage), Simi Hills, Ventura County, Paleocene, southern Cali- fornia. Basal (M), dorsal (N), and right (labral) side (O) views, 1.6x, length 32.3 mm, width 24.6 mm, height 17.3 mm, of holotype UCMP 30498, UCMP Locality 3818. P. Protocypraea ? sp. 1, Oyster Bay Formation, Paleocene, Appian Way, Central Island District, Vancouver Island, British Columbia. Basal view, 1.8x, length 48.1 mm, width 31.4 mm, height 25.3 mm, of CDM 997.91.3, informal locality AP3.

Stage”), Baja California Sur, México.

Etymology— Originally named for Lindsey T. Groves (Natural History Museum of Los Angeles County, Mala- cology Department).

Description— Shell small (17 mm length), oval, high- est at mid-point of dorsum. Base flattened with promi- nent concavity near anterior end. Spire covered, spire area coincident with small pit. Aperture straight, except for bending leftward in posterior region. Denticulation strong, outer lip with 16 teeth, inner lip with 12 teeth, sculpture extending halfway to shell margin, and ribs slightly stronger on inner lip. Outer lip callus prominent and with dorsal ring circumscribing entire shell ( Squires and Demetrion 1992: p. 31).

Remarks— The two known specimens of Bernaya grovesi have good to excellent preservation.