Ephydrini

Mathis, Wayne N., 2008, Two new neotropical genera of the shore-fly tribe Ephydrini Zetterstedt (Diptera: Ephydridae), Zootaxa 1874, pp. 1-15 : 3-4

Key to genera and subgenera of Ephydrini Zetterstedt

1. Prosternum setulose on at least posterior portion. Pulvilli much reduced or absent; tarsal claws long and nearly straight..............................................................................................................................................2

- Prosternum bare. Pulvilli well developed; tarsal claws short and distinctly curved..................................11

2. Basal flagellomere bearing a large lateral seta just below insertion of arista.................... Setacera Cresson

- Basal flagellomere without a lateral seta.....................................................................................................3

3. 3 or more well-developed fronto-orbital setae.............................................................................................4

- 2 well-developed fronto-orbital setae present. Anterior presutural supra-alar seta absent or much reduced, much smaller than posterior notopleural seta..............................................................................................7

4. 5–6 well-developed fronto-orbital setae ................................................................... Neoephydra , gen. nov.

- 3–4 well-developed fronto-orbital setae......................................................................................................5

5. 2 presutural dorsocentral setae, anterior seta sometimes rather short. Face very thickly setulose; arista short, weakly haired, thicker on its basal half; usually 2 posteriorly directed rows of well-developed cruciate interfrontal setae present, with these rows closer to orbital setae than to each other. Male with basitarsomere of foreleg bearing ventral tuft of long setulae near tip ................... Ephydra (Hydropyrus Cresson)

- 1 presutural dorsocentral seta. Face with 1 well-developed row of facial setae and 1 of oral setae, otherwise thinly short-haired; arista without markedly swollen basal region; usually not more than 1 pair of interfrontal setae situated as close to each other as to orbital setae. Male with basitarsomere of foreleg without ventral tuft of setulae......................................................................................................................6

6. 1 well-developed interfrontal seta present; palpus well developed. Crossvein dm-cu making nearly a right angle with vein CuA l .......................................................................................... Ephydra ( Ephydra Fallén)

- Interfrontal seta weak or absent; palpus small. Crossvein dm-cu forming an acute angle with vein CuA l... ........................................................................................................................ Ephydra (Halephydra Wirth)

7. Cruciate intrafrontal setae present...............................................................................................................8

- Cruciate intrafrontal setae absent...............................................................................................................10

8. Dorsocentral setae 4 (1+3). Arista bearing subpectinate dorsally-branching rays on basal 1/2.................... ................................................................................................................................ Paraephydra , gen. nov.

- Dorsocentral setae 5 (1+4). Arista at most minutely haired on basal 1/2....................................................9

9. Aristal rays long, length subequal to width of pedicel. Hindfemur of male not differing markedly from fore- or midfemur, lacking stout setae as above; hindtibia of male lacking tuft of setulae; hindtarsi of male cylindrical, normal....................................................................................................... Dimecoenia Cresson

- Aristal rays short, length approximately 1/2 width of pedicel. Hindfemur of male conspicuously swollen, bearing short row of 4-5 stout setae along anteroventral surface toward base; hindtibia of male with ventroapical tuft of setulae; hindtarsi of male variously modified ............................................ Cirrula Cresson

10. A well-developed prescutellar acrostichal seta; 2 postpronotal setae, dorsal seta about ½ length of ventral seta; prosternal setulae few (Australasian) ............................................... Ephydrella Tonnoir and Malloch

- Lacking a well-developed prescutellar acrostichal seta; 1 postpronotal seta; prosternal setulae numerous (Neotropical) ............................................................................................................. Neoephydra , gen. nov.

11. Dorsocentral setae 4 (1+3) present; postpronotal seta(e) either weak, at most 1/4 length of posterior notopleural seta, or lacking. Hindcoxa bare posteriorly............................................................................12

- Dorsocentral setae 5 (1+4) present (anterior setae reduced in Austrocoenia ); postpronotal seta distinct, at least 1/2 as long as posterior notopleural seta. Hindcoxa bare or with a row of setae posteriorly............13

12. Arista bearing long hairs dorsally, length of longest hairs subequal to height of basal flagellomere; frons only moderately to sparsely microtomentose, especially subshiny mesofrons (Holarctic)........................... .......................................................................................................................... Coenia Robineau-Desvoidy

- Arista either almost bare or bearing short hairs dorsally, length of longest hairs about 1/2 height of basal flagellomere; frons uniformly and densely microtomentose, dull, mesofrons little differentiated from parafrons (Neotropical) .................................................................................................. Notiocoenia Mathis

13. Paravertical seta large, at least 1/3 length of medial vertical seta.............................................................14

- Paravertical seta small, generally subequal to setae of postocular row.....................................................16

14. R stem vein bearing 1–2 setulae dorsally, inserted beyond transverse septum. Scutellar disc convex. Hindcoxa with row of setae along posteroventral margin .................................................... Paracoenia Cresson

- R stem vein bare dorsally. Scutellar disc almost flat. Hindcoxa bare posteriorly along ventral margin (Genus Calocoenia Mathis ).......................................................................................................................15

15. Larger species, body length over 3.25 mm. Gena-to-eye ratio 0.25 or larger. Costal setulae well developed, projected anteriorly from ventral and dorsal surfaces.......................... Subgenus Calocoenia Mathis

- Smaller species, body length under 2.75 mm. Gena-to-eye ratio 0.20 or smaller. Costal setulae weakly developed, only on dorsal surface ................................................................ Subgenus Leptocoenia Mathis

16. Frons mostly lacking setulae and with coloration and vestiture generally uniform, microtomentose, dull; arista long, nearly double length of basal flagellomere. 1 interalar seta inserted just posterior of transverse suture .............................................................................................................................. Notiocoenia Mathis

- Mesofrons tan to brown, generally distinct from grayer parafrons, microtomentum denser, conspicuously setulose, especially laterally; arista short, subequal to length of basal flagellomere. Interalar seta lacking ....................................................................................................................................... Austrocoenia Wirth