Desmoxytes cervaria ( Attems, 1953 )

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2015, Two new species of dragon millipedes, genus Desmoxytes Chamnerlin, 1923, from Laos (Diplopoda: Polydesmida: Paradoxosomatidae), with redescriptions of all four species of Attems from Vietnam, Zootaxa 3931 (4), pp. 483-504 : 500-502

publication ID

https://doi.org/ 10.11646/zootaxa.3931.4.2

publication LSID

lsid:zoobank.org:pub:2F14AF9A-3E4C-4A30-960B-8C612220D4E4

DOI

https://doi.org/10.5281/zenodo.6118685

persistent identifier

https://treatment.plazi.org/id/90032A20-FFCB-B570-66F0-6759AA9A21A3

treatment provided by

Plazi

scientific name

Desmoxytes cervaria ( Attems, 1953 )
status

 

Desmoxytes cervaria ( Attems, 1953) View in CoL

Figs 13–14 View FIGURE 13 View FIGURE 14

Centrodesmus cervarius Attems, 1953: 175 (D).

Pratinus cervarius — Jeekel, 1964: 61 (D, M); 1968: 61 (M).

Hylomus cervarius View in CoL — Jeekel, 1980: 657 (M, R); Golovatch, 1983: 181 (M).

Desmoxytes cervaria View in CoL — Golovatch & Enghoff, 1994: 48, 60 (D, M); Enghoff et al., 2004: 37 (R); Nguyen et al., 2006: 257 (D); Nguyen & Sierwald, 2013: 1241 (R).

Lectotype male (NHMW-3525), Vietnam, Laocai Province, Fanxipan Mountain, 1938–1939, leg. C. Dawydoff. Paralectotypes. 1 male, 1 female (NHMW-3525), same locality, together with lectotype.

Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male.

Description. Length ca 23–24 mm (male) or 28 mm (female), width of midbody pro- and metazonae 1.3–1.7 and 3.8–4.0 mm (male) or 2.3 and 4.0 mm (female), respectively (versus length ca 25 mm (unknown sex), width 2.3 mm (unknown sex), respectively, as given in the original description (Attems, 1953)).

Coloration of alcohol material upon long-term preservation red-brown to yellow-brown ( Fig. 13 View FIGURE 13 ); paraterga and epiproct pale yellow-brown (versus head, antennae and collum dark chestnut brown, metaterga sometimes lighter, paraterga brown, venter yellow-brown, as given in the original description (Attems, 1953)).

Clypeolabral region and vertex densely setose, epicranial suture distinct. Antennae very long, reaching segment 6 (both sexes) when stretched dorsally. In width, head <collum <segment 2 <3 <4 <5–17 (both sexes); thereafter body gently and gradually tapering. Collum with three transverse rows of setiferous spines: 3+3 anterior, 1+1 intermediate, and 3+3 posterior rows, posterior mesal ones strongest; paraterga antler-shaped, directed dorsolaterad, with two setigerous incisions anteriorly ( Fig. 13 View FIGURE 13 A & B). Tegument dull, prozonae finely shagreened, metazonae coarsely granulate, surface below paraterga coarsely microgranulate, sterna delicately microgranulate ( Fig. 13 View FIGURE 13 A–G, K & L). Metaterga with two transverse rows of setigerous spines, 2+2 small anterior and 3+3 thornshaped posterior, lateral spines of posterior rows being much longer than others, intermeditate ones shorter, mesal ones shortest except for segments 2–4 with 2+2 anterior and 2(3)+2(3) posterior spines, mesal ones being largest ( Fig. 13 View FIGURE 13 A, C, F, K & L). Axial line faint. Paraterga antler-shaped, especially well so in male, upturned, directed dorsolaterad, tips bent posteriad; with a prominent anterior spine at 1/3–1/2 extent of paraterga and a small setigerous notch near tip in segments 2–4; a prominent spine located lower to 1/4 extent of paraterga in segments 5–9; with two setigerous notches at about midway and 2/3 extent of paraterga in following segments ( Fig. 13 View FIGURE 13 A–F, K & L). Ozopores lateral, lying distal to base of basal spine ( Fig. 13 View FIGURE 13 D). Transverse sulcus incomplete on segments 5–18, line-shaped, narrow, wide, reaching bases of paraterga; incomplete on segment 4, absent from 19th ( Fig. 13 View FIGURE 13 A, C, F, K & L). Stricture between pro- and metazonae narrow, shallow, beaded at bottom down to base of paraterga ( Fig. 13 View FIGURE 13 A-F, K & L). Pleurosternal carinae complete crests only in segments 2 and 3, with an anteriorly dentate edge, absent thereafter. Epiproct ( Fig. 13 View FIGURE 13 E, F & G) conical, flattened dorsoventrally, apical papillae very small; pre-apical papillae small, but visible, lying close to a convex tip. Hypoproct subtrapeziform, setiferous knobs at a slightly convex caudal edge medium-sized and well-separated.

Sterna sparsely setose, without modifications; cross-impressions shallow; an elevated, subquadrate, setose lamina between male coxae 4, carrying a rounded tubercle at each corner ( Fig. 13 View FIGURE 13 H & I). Legs very long and slender, midbody ones ca 1.4–1.6 (male) or 1.1–1.3 times (female) as long as body height. male prefemora 6 with a very evident, rounded, mesal, densely pilose apophysis in distal 2/3, male prefemora 7 clearly inflated medially ( Fig. 13 View FIGURE 13 J). Male tarsal brushes absent.

Gonopods ( Fig. 14 View FIGURE 14 ) suberect. Prefemur sparsely setose, about half as long as fermorite + postfemoral part. Femorite rather slender, enlarged distad, seminal groove running entirely on mesal face. Solenophore (= lamina medialis) well-developed, falcate and rather condensed, with two apical lobes. Solenomere rather short, flagelliform, well separated at base from solenophore.

Kingdom

Animalia

Phylum

Arthropoda

Class

Diplopoda

Order

Polydesmida

Family

Paradoxosomatidae

Genus

Desmoxytes

Loc

Desmoxytes cervaria ( Attems, 1953 )

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak 2015
2015
Loc

Desmoxytes cervaria

Nguyen 2013: 1241
Nguyen 2006: 257
Enghoff 2004: 37
Golovatch 1994: 48
1994
Loc

Hylomus cervarius

Golovatch 1983: 181
Jeekel 1980: 657
1980
Loc

Pratinus cervarius

Jeekel 1964: 61
1964
Loc

Centrodesmus cervarius

Attems 1953: 175
1953
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