Jaguajir rochae ( Borelli, 1910 ) Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini, 2017

Lauren A. Esposito, Humberto Y. Yamaguti, Cláudio A. Souza, Ricardo Pinto-Da-Rocha & Lorenzo Prendini, 2017, Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae), Bulletin of the American Museum of Natural History 415, pp. 1-134 : 84-86

Jaguajir rochae ( Borelli, 1910) , comb. nov.

Figures 1 D , 2D , 8 , 11D , 15F , 17C , 19F , 21F , 22F , 24E–H , 41C , 42C , 43C, F , 44C , 47

Rhopalurus rochae Borelli, 1910: 3–5 ; Mello- Leitão, 1932: 31; Meise, 1934: 42; Lourenço, 1982 a: 122, 135, figs. 47–62, 78, table 1 ; 1982b: 74, figs. 1–3; 1986b: 165, fig. 7; 1990: 161; Kovařík, 1998: 118; Fet and Lowe, 2000: 222; Kamenz and Prendini, 2008: 9, table 2 , pl. 44; Volschenk et al., 2008: 651, 652, 654, 660, 661, 663, 664, 674, fig. 2E, tables 1 , 2 ; Prendini et al., 2009: 222–224.

Centrurus stenochirus Penther, 1913: 240–242 , figs. 1, 2 (synonymized by Lourenço, 1982 a: 122).

Centrurus barythenar Penther, 1913: 242–243 , figs. 1, 2 (synonymized by Lourenço, 1982 a: 122); Werner, 1927: 358; Mello-Leitão, 1945: 286.

Rhopalurus barythenar: Mello-Campos, 1924 a: 252 , 276, 281, 282, pl. 5, fig. 27, pl. 8, fig. 34; 1924b: 318, 341, 347, pl. 5, fig. 27, pl. 8, fig. 34; Mello-Leitão, 1932: 16, 31; Vellard, 1932: 556; Meise, 1934: 42; Prado, 1940: 26, 34, 35.

Rhopalurus rochai: Mello-Campos, 1924 a: 252 , 276, 279–281, pl. 10, figs. 35, 36; 1924b: 318, 341, 345, 346, pl. 10, figs. 35, 36; Werner, 1927: 358; Mello-Leitão, 1932: 16; Prado, 1940: 26, 34; Mello-Leitão, 1945: 267, 285– 288, figs. 118, 119; Bücherl, 1959: 269; 1971: 327; Lucas and Bücherl, 1972: 263; Stahnke and Calos, 1977: 119; Lourenço, 1979: 215, fig. 7; Araújo, 1981: 235; Lucas et al., 1981: 128, table 1 ; Fet and Lowe, 2000: 222; Brazil and Porto, 2010: 25, 50, 59, 66, figs. 5B, 7A; Porto et al., 2010: 292–294, 296, fig. 2B, table 1 ; Ubinski et al., 2016: 122.

Rhopalurus stenochirus: Mello-Campos, 1924 a: 252 , 275, 278, 279, pl. 11, fig. 39; 1924b: 318, 341, 343–345, pl. 11, fig. 39; Mello-Leitão, 1932: 15, 31; Meise, 1934: 42; Prado, 1940: 26, 30, 31; Mello-Leitão, 1945: 267, 288–290, figs. 120, 121; Bücherl, 1971: 327; Lucas and Bücherl, 1972: 263; Stahnke and Calos, 1977: 119; Araújo, 1981: 235; Lourenço, 1982 a: 122, 123, figs. 49, 50.

Centruroides stenochirus: Werner, 1927: 357 .

Ropalurus barythenar: Viquez, 1935: 111 , fig. 40.

TYPE MATERIAL: BRAZIL: Bahía: Centrurus barythenar : 12 ♂ syntypes (NMW 2248), Joazeiro. Centrurus stenochirus : 2 ♂, 1 juv.? ♀ syntypes (NMW 2251), Barra. Ceará: Rhopalurus rochae : Lectotype ♀, paralectotype ♂ [desig. Lourenço, 1982 a: 138] (MIZT Sc 421 ex 645), Fortaleza.

DIAGNOSIS: Jaguajir rochae can be differentiated from J. agamemnon and J. pintoi by its paler coloration and more slender metasoma. Whereas the base coloration is pale to dark yellow, and the metasoma of the male slightly wider posteriorly in J. rochae , the base coloration is brown to black, and the metasoma of the male markedly wider posteriorly in J. agamemnon and J. pintoi . Additionally, the fixed and movable fingers of the pedipalp chela of the male are slightly curved, creating a small proximal gap between them, and the proximal lobe on the fixed finger of the male exhibits a prominent proximal lobe in J. rochae , whereas the fixed and movable fingers of the pedipalp chela of the male are markedly curved, creating a large proximal gap between them, and the proximal lobe on the fixed finger of the male is reduced or absent in J. agamemnon and J. pintoi . Furthermore, the subaculear tubercle of the telson is much reduced, forming a small protuberance in J. rochae , as in J. pintoi , but well developed and spinoid in J. agamemnon .

DISTRIBUTION: Jaguajir rochae is endemic to northeastern Brazil and recorded from the states of Bahía, Ceará, Paraíba, Pernambuco, Piauí, Río Grande do Norte and Sergipe (fig. 8). A record from Costa Rica ( Viquez, 1935) is based on a misidentification.

ECOLOGY: This species inhabits open savannah (fig. 2D). Specimens collected by the authors were found under stones during the day and with UV light detection at night. The habitat and habitus are consistent with the lapidicolous ecomorphotype ( Prendini, 2001 b).

REMARKS: Borelli (1910) named the species after Francisco Diaz da Rocha, but the original spelling was feminine, rochae . Fet and Lowe (2000) noted that the masculine spelling is rochai and changed it accordingly. Although the corrected spelling has been adopted by others (e.g., Teruel, 2006: 52), we consider it an unjustified emendation and continue to use Borelli’s (1910) original spelling. It is noteworthy that, on the page preceding the description of R. rochae, Borelli (1910) described Tityus duckei Borelli, 1910 , after Mr. Ducke, implying he was aware of the proper ending for a masculine name and it was not a lapsus. If Rocha is considered a Latin name, the proper ending, even if dedicated to a man, is rochae ( ICZN, 1999: Art. 31.1.1. Examples), which may be what Borelli (1910) intended.

MATERIAL EXAMINED: BRAZIL: Bahía: Barra, Igarité, 5.vi.2008, 1 ♀ (MZSP 31122/ AMCC [LP 9963]); Catu, Ibiraba, 2.viii.2000 , 1 ♀ (MZSP 31123/AMCC [LP 9953]); Guanambí, Ceraíma, 14°13′S 42°46′W, 10–17. xii.2007, H.Y. Yamaguti et al. , 1 ♂ (MZSP 30881/AMCC [LP 9684]); Guanambí, 7 km S, 14°17′5.6″S 42°47′2.2″W, 533 m, 24.i.2007, C.I. Mattoni, R. Pinto-da-Rocha, and H.Y. Yamaguti, UV sampling, modified savannah, cloudy and raining , 1 juv. (AMCC [LP 7638]); Guanambí , 16 km SE, 14°17′19″S 42°41′31.1″W, 559 m, 25.i.2007, C.I. Mattoni, R. Pinto-da- Rocha, and H.Y. Yamaguti, UV sampling and under leaf litter, banana plantation and surrounds , 1 juv. (AMCC [LP 7655]); Fazenda do Fabiano , 8 km NE Guanambí, 14°10′17.6″S 42°43′56.4″W, 539 m, 24.i.2007, C.I. Mattoni, R. Pinto-da-Rocha, and H.Y. Yamaguti, under rocks, rocky hill and surrounds, open savannah modified , 1 ♀ (AMNH), 1 ♂, 2 juv. (AMCC [LP 7639]); Jeremoabo, Espadoado de Cima , 10°04′S 38°23′W, 27.v.2008, H.Y. Yamaguti, T.J. Porto, and M.B. da Silva , 1 subad. ♀ (MZSP 30882/AMCC [LP 9685]); Ibotirama, 12°10′34″S 43°11′33″W, 8.vi.2008, H.Y. Yamaguti, M.B. da Silva, and T.J. Porto , 1 ♂ (MZSP 31125/AMCC [LP 9956]), 1 ♀ (MZSP 31124/ AMCC [LP 9947]); Jeremoabo, 10°04′S 38°23′W, 25.v.2008, H.Y. Yamaguti, M.B. da Silva, and T.J. Porto , 1 ♂ (MZSP 31126/AMCC [LP 9945]); Várzea do Poço , 19.x.2007, 1 ♀ (MZSP 31127/AMCC [LP 9926]). Minas Gerais: Janaúba, 11.xii.2007, H.Y. Yamaguti, S. Outeda- Jorge, and C.A. Souza , 1 juv. ♂ (MZSP 31136/ AMCC [LP 9957]); Janaúba, UNIMONTES, Campus Janaúba, 11.xii.2007, H.Y. Yamaguti et al. , 1 subad. ♂ (MZSP 30879/AMCC [LP 9682]). Paraíba: Soledade, 07°02.118′S 36°27.311′W, 575 m, 16.iii.1999, A. Kury and A. Giupponi , 3 ♂ (AMCC [LP 1581, 1582, 1775]). Pernambuco: Escola Aquicola, Exu , 30.iii.1977, L.J. Vitt , caatinga, 1 ♂ (AMNH), 27.vi.1977, L.J. Vitt , 1 ♂ (AMNH); Exu , 07°26′44″S 39°44′21″W, 1.vi.2008, H.Y. Yamaguti, M.B. da Silva, and T.J. Porto , 1 ♀ (MZSP 31144/AMCC [LP 9955]), 1 juv. ♂ (MZSP 31143/AMCC [LP 9951]); Exu , 18 km E, 5. iii.1977, L.J. Vitt , under leaf of granite on boul- der, caatinga habitat , 2 ♀, 68 1 st instars (AMNH); Exu , 20 km E, 30.iii.1977, L.J. Vitt , 1 ♂, 1 ♀, 2 juv. (AMNH); Exu , 3 km NW, 10. iii.1977, L.J. Vitt , 2 ♂, 1 ♀, 3 juv. (AMNH); Exu , 3 km W, 30.v.1977, L.J. Vitt , 2 ♂, 4 ♀, 4 juv. (AMNH), 1.vi.1977, L.J. Vitt , 1 ♀ (AMNH); Exu , 5 km E, 8.v.1977, L.J. Vitt , 1 juv. (AMNH); Exu , 5 km N, 6.iv.1977, L.J. Vitt , caatinga, 1 ♂, 1 juv. (AMNH), 18.i.1978, L.J. Vitt and K.E. Streilein, 1 juv. (AMNH); Exu , 6 km N, 15. iii.1977, L.J. Vitt , open fields (cotton), under fallen logs, 1 ♀, 1 juv. ♂ (AMNH); Exu , 6 km NE, 16.iii.1977, L.J. Vitt , under rock on larger rock, caatinga habitat, 1 ♀, 49 1 st instars (AMNH); Fazenda Batente, 5 km NE Exu, 29. iii.1977, L.J. Vitt , 1 juv. (AMNH); Fazenda Caterino, 10 km NE Exu, 1.viii.1977, L.J. Vitt , 7 ♂, 3 ♀, 3 juv. (AMNH); Fazenda Chelonia, 8 km S Exu, 28.vii.1977, L.J. Vitt , 2 juv. (AMNH); Fazenda Guarani, 3 km N Exu, 14.vii.1977, L.J. Vitt , 1 ♂, 3 ♀, 1 subad., 3 juv. (AMNH); Fazenda Guarani, 5 km N Exu, 29.vii.1977, L.J. Vitt , 1 ♀, 3 juv. (AMNH), 19.ii.1978, L.J. Vitt , 1 ♀ (AMNH); Serra Talhada, 07°58′11″S 38°19′16″W, 30.v.2008, H.Y. Yamaguti, M.B. da Silva, and T.J. Porto , 2 ♀ (MZSP 31146/AMCC [LP 9943], MZSP 31147/AMCC [LP 9930]); Vitória do Santo Antão , 08°07′S 35°25′W, 28.v.2008, H.Y. Yamaguti, M.B. da Silva, and T.J. Porto , 1 ♀ (MZSP 31148/AMCC [LP 9946]). Piauí: Castelo do Piauí, 05°13′43″S 41°41′57″W, 13.viii.2008, R. Pinto-da-Rocha and L.S. Carvalho , 1 ♂ (MZSP 31150/AMCC [LP 9961]), 1 ♀ (MZSP 31151/AMCC [LP 9941]). Sergipe: near Genipapo, vii.1982, O.F. Francke , 1 ♂, 2 ♀ (AMNH); UHE Xingó, Canindé de São Francisco , 22.viii.2007, Arnaldo Jr., 1 ♀ (MZSP 30880/AMCC [LP 9683]) .

Figures

FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil.

FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil.

FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004.

FIG. 8. Map of northeastern South America, plotting known locality records of Jaguajir rochae (Borelli, 1910), comb. nov. (circles).

FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP).

FIG. 15. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♀ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (AMNH). Scale bars = 2 mm.

FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm.

FIG. 19. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♀ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). Scale bars = 2 mm.

FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm.

FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm.

FIG. 24. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (B, H, M, P), ectal aspect (F, I, N, S), ectal aspect, detail (D, J, R, T, U), ental aspect (A, G, K, O), and ental aspect, detail (C, L, Q). A–D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E–H. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). I–M. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP). N–R. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (MZSP). S–U. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). Scale bars = 1 mm.

FIG. 41. Jaguajir, gen. nov., pedipalp chela, retrolateral aspect. A. J. agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). B. J. pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). C. J. rochae (Borelli, 1910), comb. nov., ♂ (MZSP). Scale bars = 2 mm.

FIG. 42. Jaguajir, gen. nov., pedipalp chela, retrolateral aspect. A. J. agamemnon (C.L. Koch, 1839), comb. nov., ♀ (MZSP). B. J. pintoi (Mello-Leitão, 1932), comb. nov., ♀ (MZSP). C. J. rochae (Borelli, 1910), comb. nov., ♀ (AMNH). Scale bars = 2 mm.

FIG. 43. Jaguajir, gen. nov., metasoma and telson, dorsal (A–C) and ventral (D–F) aspects. A, D. J. agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). B, E. J. pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). C, F. J. rochae (Borelli, 1910), comb. nov., ♂ (AMNH). Scale bars = 5 mm.

FIG. 44. Jaguajir, gen. nov., metasoma and telson, lateral aspect. A. J. agamemnon (C.L. Koch, 1839), comb. nov., ♂ (MZSP). B. J. pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). C. J. rochae (Borelli, 1910), comb. nov., ♂ (AMNH). Scale bars = 5 mm.

FIG. 47. Jaguajir rochae (Borelli, 1910), comb. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. ♂ (AMNH). C, D. ♀ (AMNH). Scale bars = 10 mm.