Sarax indochinensis, Miranda & Giupponi & Prendini & Scharff, 2021

Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), European Journal of Taxonomy 772, pp. 1-409 : 186-188

publication ID 10.5852/ejt.2021.772.1505

publication LSID


persistent identifier

taxon LSID

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scientific name

Sarax indochinensis

sp. nov.

Sarax indochinensis View in CoL sp. nov.

Figs 103 View Fig , 112–114 View Fig View Fig View Fig ; Table 8

Sarax brachydactylus View in CoL – Fage 1946b: 79 (in part).

Sarax aff. brachydactylus View in CoL – Seiter et al. 2015: 545.


This species may be separated from other species of Sarax in Southeast Asia and Oceania by the following combination of characters: median and lateral eyes well developed ( Fig. 112A View Fig ); ventral sacs and ventral sac cover well developed; female gonopod finger-like with small invagination apically ( Fig. 113A View Fig ); cheliceral basal segment with small tooth adjacent to bifid tooth; secondary sexual dimorphism in males with longer pedipalp segments than females; pedipalp tarsus with two short dorsal spines; leg IV basitibia with four pseudo-articles, distitibia sc and sf series each with five trichobothria.

This species resembles S. brachydactylus , but the genital operculum is concave, with two symmetric bulges unlike that of S. brachydactylus , which is flat.


The species name is an adjective derived from Indochina, the region of Southeast Asia which included Cambodia, Laos, and Vietnam, under French control between ca 1890–1950.

Type material

Holotype THAILAND • ♀; Koh Kut [Ko Kut]; [11°40′05.07″ N, 102°33′51.21″ E]; B. Degerbøl leg.; 6 Apr. 1959; ZMUC 21613 View Materials . GoogleMaps

Paratypes THAILAND • 3 ♀♀, 7 ♂♂, 1 juv.; same locality as for holotype; B. Degerbøl le g.; 5 Apr. 1959; ZMUC 21612 View Materials GoogleMaps .

VIETNAM • 1 ♀; Annam, Ha Tien, Kien Giang Province; Hang Trong Nguc ( Da Dung mountain ); 10°25′44.8″ N, 104°28′36.6″ E; 7 Jan. 2010; H. Steiner leg. (Northern Lao-European Cave Project 2010); SMF 029/10 GoogleMaps .

Additional material

CAMBODIA • 1 ♀; Sré Umbel [ Srae Ambel ]; [11°07′01.82″ N, 103°44′43.14″ E]; Feb. 1939; Mission C. Dawydoff, Indochine, 1938–1939; MNHN GoogleMaps 1 ♂; Kampong Speu; Apr. 1939; Mission C. Dawydoff, Indochine, 1938–1939; MNHN 1 ♀, 2 juv.; Ream; [10°35′02.75″ N, 103°38′35.34″ E]; Mar. 1939; Mission C. Dawydoff, 1938–1939, Indochine; MNHN GoogleMaps 1 ♂; [11°34′37.44″ N, 103°07′41.04″ E]; 10 m a.s.l.; 3 Dec. 2014; J. Nigl leg.; NHMW 27601 View Materials GoogleMaps 1 juv.; Koh Chang; [12°04′28.60″ N, 102°18′35.95″ E]; under bark; 13 Jan. 1900; Th. Mortensen leg.; ZMUC 21614 View Materials GoogleMaps .

VIETNAM • 1 ♂; Phaurang , Tour Cham; May 1939; Mission C. Dawydoff, Indochine 1938–1939; MNHN 1 ♂; Banghoi [Ba Ngôi], Indochine; May 1939; Mission C. Dawydoff; 1938–1939; MNHN 2 juv.; Golf Du Siam [ Gulf of Thailand ]; Nov. 1938; P. Dama, Mission C. Dawydoff, Indochine, 1938– 1939; MNHN 1 ♀, 1 ♂; Hang Bong Lai; 10°25′42″ N, 104°28′39″ E; H. Steiner leg.; SMNS 86 View Materials GoogleMaps 1 ♀; Hang 8; 10°25′45″ N, 104°28′37.8″ E; H. Steiner leg.; SMNS 87 View Materials GoogleMaps .

NO DATA • 4 juv.; MNHN 1 ♀, 7 protonymphs; MNHN 1 ♀, 1 ♂, 1 juv.; MNHN 2 ♀; MNHN .


CARAPACE. Six anterior setae ( Fig. 112A View Fig ); frontal process triangular ( Fig. 112C View Fig ). Small granules densely scattered between ocular triads and among sulci. Median eyes and median ocular tubercle well developed ( Fig. 112C View Fig ); pair of setae on median ocular tubercle; lateral eyes well developed, pale, seta lateral to lateral ocular triad; lateral ocular triad near carapace margin; curved carina between ocular triads and carapace margin ( Fig. 112C View Fig ).

STERNUM. Tritosternum projected anteriorly with typical setation, long, surpassing base of pedipalp coxae ( Fig. 112B View Fig ); other sternal platelets narrow, projecting, with pair of setae anteriorly on plaque and some smaller setae posteriorly; pentasternum with four setae near membranous region.

OPISTHOSOMA. Ventral sacs cover well developed.

GENITALIA. Female genital operculum with short setae posteromedially; pair of white bulges medially and slender setae apically ( Fig. 113A–C View Fig ); gonopod finger-like, unsclerotized basally ( Fig. 113A, C, F View Fig ); denticulate margin between gonopod and posterior margin of genital operculum ( Fig. 113E View Fig ); slit sensilla laterally on genital operculum ( Fig. 113E View Fig ). Male gonopod as wide as long, wider in distal third ( Fig. 114C View Fig ); Lol2 fimbriate with small spines apically on projections ( Fig. 114A–B, D View Fig ); PI with smooth surface and acute apex ( Fig. 114F View Fig ); dorsal lobe short, with spines on inner margin and ventrally, and apex curved outwards ( Fig. 114C, G View Fig ); LaM short, not fused medially, with smooth surface ( Fig. 114A– B View Fig ); fistula with inner spines ( Fig. 114E View Fig ).

CHELICERAE. Retrolateral surface of basal segment with short projection, opposite to bifid tooth; retrolateral surface of claw with row of setae basally to medially; claw with five teeth; more than two rows of several setae on prolateral surface of basal segment; bifid tooth on basal segment with dorsal cusp larger than ventral cusp.

PEDIPALPS. Coxae without seta encircled by round carina and with three setae on margin. Femur with four dorsal spines and four ventral spines ( Fig. 112E–F View Fig ); two prominent setiferous tubercles between dorsal spine 1 and proximal margin; long spine between ventral spine 1 and proximal margin, two-thirds length of spine 1. Patella with four dorsal spines in primary series ( Fig. 112E View Fig ); prominent setiferous tubercle distal to spine I; three ventral spines ( Fig. 112F View Fig ); setiferous tubercle between spine I and distal margin. Tibia with ventral spine distally and two setae between spine and distal margin. Tarsus with two subequal dorsal spines ( Fig. 112D View Fig ); cleaning organ with 30–32 setae in ventral row.

LEGS. Tibia of leg I with 23 articles; tarsus I with 41 articles; first and second tarsal articles equal in length. Leg IV basitibia with four pseudo-articles, with sclerotized, denticulate margin at apex of articles; trichobothrium bt situated slightly closer to distal third of pseudo-article; distitibia trichobothrium bc situated equidistant between bf and s bf, and sc and sf series each with five trichobothria.


See Table 8.


Known from Cambodia, Thailand and Vietnam. The distribution of this species overlaps that of Weygoldtia davidovi .


Material from Cambodia (Sre Unebell, Réam) in the MNHN was misidentified as S. brachydactylus by Fage (1946).


Museum National d'Histoire Naturelle














Sarax indochinensis

Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj 2021

Sarax aff. brachydactylus

Seiter M. & Wolff J. & Horweg C. 2015: 545

Sarax brachydactylus

Fage L. 1946: 79
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