Myrmicinae Lepeletier

Kiran, Kadrï & Karaman, Celal, 2012, First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae), Zootaxa 3548, pp. 1-38 : 16-29

Subfamily Myrmicinae Lepeletier

Anergates atratulus (Schenck, 1852) : Thrace (Çamlıtepe & Aktaç 1987; Radchenko 2007); [13] ( Heinze 1987); [4] (Aktaç et al. 1994); [45] ( Schulz & Sanetra 2002); [22], [39] (Lapeva-Gjonova et al. 2012).

Aphaenogaster aktaci Kiran & Tezcan, 2008 : [3] (Kiran et al. 2008).

Aphaenogaster balcanica ( Emery, 1898) : Thrace (Radchenko 2007).

as Stenamma testaceo-pilosa var. balcanica : [10], [12] ( Emery 1898).

as Aphaenogaster testaceopilosa var. balcanica : Asia Minor ( Ruzsky 1905).

as Aphaenogaster testaceo-pilosa subsp. balcanica : [3], [10] ( Forel 1911).

as Aphaenogaster testaceo-pilosa subsp. simonellii var. balcanica : [10] ( Emery 1915); [9] ( Emery 1921a). as Aphaenogaster testaceo-pilosa subsp. semipolita var. balcanica : [10] ( Schkaff 1924). as Aphaenogaster simonelli (sic!) var. balcanica : [3], [10] ( Santschi 1934).

as Aphaenogaster simonellii var. balcanica : [9] ( Aktaç 1976).

Aphaenogaster burri (Donisthorpe, 1950)

as Goniomma burri : [70] ( Donisthorpe 1950a; c).

Aphaenogaster depressa Bolton, 1995

as Ceratopheidole pergandei Donisthorpe, 1950 : [53] ( Donisthorpe 1950a). as Ceratopheidole perganderi (sic!): [53] ( Donisthorpe 1950d). as Aphaenogaster pergandei : [53] ( Brown 1981).

Aphaenogaster epirotes (Emery, 1895) : Thrace (Radchenko 2007); [4] (Lapeva-Gjonova & Kiran 2012). as Aphaenogaster obsidiana subsp. epirotes : [10] ( Schkaff 1924).

as Aphaenogaster obsidiana var. epirotes : [10], [12] ( Finzi 1930).

as Aphaenogaster obsidiana ( Mayr, 1861) : Thrace ( Aras & Aktaç 1990; Radchenko 2007); [2], [4], [5] (Aktaç et al. 1994) (misidentifications).

Comments: See the comments for Aphaenogaster obsidiana below.

Aphaenogaster festae Emery, 1915

as Aphaenogaster festai : Turkey (Arnol'di 1976).

Aphaenogaster gibbosa (Latreille, 1798) : [10] ( Forel 1911; Donisthorpe 1950a); Thrace (Radchenko 2007); [4] (Lapeva-Gjonova & Kiran 2012).

as Aphaenogaster striola ( Roger, 1859) : Asia Minor ( André 1883a).

as Aphaenogaster subterranaea subsp. gibbosa : Asia Minor ( Ruzsky 1905).

Aphaenogaster holtzi ( Emery, 1898) : [36] ( Emery 1908b); [45] ( Finzi 1930); [77] ( Aktaç 1976); [61], [74] ( Aktaç 1988).

as Stenamma subterraneum subsp. holtzi : [36] ( Emery 1898).

Aphaenogaster ionia Santschi, 1933 : [20] ( Lush 2009).

Aphaenogaster kervillei Forel, 1910 : [77] ( Aktaç 1976).

Aphaenogaster maculifrons Kiran & Aktaç, 2008 : [23], [28] (Kiran et al. 2008).

Aphaenogaster obsidiana ( Mayr, 1861) : [70] ( Karavaiev 1926a).

Comments: Although Aras & Aktaç (1990) and Aktaç et al. (1994) recorded this species from the European part of Turkey (Thrace), the material is misidentified and belongs to A. epirotes . Radchenko (2007) also recorded A.

obsidiana from Thrace but his record is based on the previously recorded material of Aras & Aktaç (1990) and Aktaç et al. (1994). Therefore, this species is known only from Anatolia. Its distribution in the European part of Turkey is removed.

Aphaenogaster radchenkoi Kiran, Aktaç & Tezcan, 2008 : [3] (Kiran et al. 2008).

Aphaenogaster semipolita (Nylander, 1856)

as Aphaenogaster testaceo-pilosa semipolita : Asia Minor ( Emery 1908b).

Aphaenogaster simonellii Emery, 1894 : [9] ( Aktaç 1978); [15] ( Baroni Urbani & Aktaç 1981); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [6] (Aktaç & Çamlıtepe 1987); [1] ( Aras & Aktaç 1987); [2] ( Aras & Aktaç 1992;1994).

Aphaenogaster splendida ( Roger, 1859) : [10] ( Forel 1895; Schkaff 1924); [9] ( Emery 1921a); [3], [20] ( Donisthorpe 1950a); Thrace (Radchenko 2007).

Aphaenogaster subcostata Viehmeyer, 1922 : [46] ( Viehmeyer 1922).

Aphaenogaster subterranea (Latreille, 1798) : [10] ( Forel 1895; Schkaff 1924); [12] ( Emery 1897); [10], [12] ( Finzi 1930); [10], [70] ( Donisthorpe 1950a); [59] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987; Kiran & Karaman 2012); [2] ( Aras & Aktaç 1992); [11], [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

as Formica subterranea : [10] ( Rigler 1852).

as Stenamma subterraneum : Asia Minor ( Emery 1898); [45] ( Kohl 1905).

Aphaenogaster subterraneoides Emery, 1881 : Thrace (Radchenko 2007). as Aphaenogaster pallida subsp. subterraneoides : [10] ( Schkaff 1924).

Aphaenogaster testaceopilosa (Lucas, 1849) : [12] ( Emery 1897); Asia Minor ( Ruzsky 1905); Thrace (Radchenko 2007).

as Aphaenogaster testaceo-pilosa: Asia Minor ( André 1883a); [10] ( Forel 1895; Donisthorpe 1950a). as Stenamma testaceo-pilosa: Sultan Mt., [12] ( Forel 1906).

Cardiocondyla brachyceps Seifert, 2003 : [70] ( Seifert 2003).

Cardiocondyla bulgarica Forel, 1892 : [3], [8], [15], [19], [20], [23], [76] ( Seifert 2003); [16] (Kiran & Aktaç 2006).

as Cardiocondyla elegans var. eleonorae Forel, 1911 : [3] ( Forel 1911; Santschi 1934).

Cardiocondyla elegans Emery, 1869 : [10], [27] ( Donisthorpe 1950a); [77] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Aktaç et al. 1994); [1] ( Aras & Aktaç 1987); [3], [20], [49] ( Seifert 2003); [12] (Kiran & Aktaç 2006).

Cardiocondyla mauritanica Forel, 1890 : [29], [36], [45] ( Seifert 2003).

Cardiocondyla nigra Forel, 1905 : Thrace ( Agosti & Collingwood 1987); [58] ( Seifert 2003).

Cardiocondyla sahlbergi Forel, 1913 : [3], [11], [28], [34], [45], [46], [49], [61], [72], [79] ( Seifert 2003).

Cardiocondyla semirubra Seifert, 2003 : [58] ( Seifert 2003).

Cardiocondyla stambuloffii Forel, 1892 : Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994); [1] ( Aras & Aktaç 1987); [28], [70], [74], [79] ( Seifert 2003).

as Cardiocondyla stambuloffi (sic!): Thrace (Radchenko 2007).

as Cardiocondyla bogdanovi Ruzky, 1905 : [79] ( Ruzsky 1905); Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Carebara oertzeni subsp. aeolia ( Forel, 1911)

as Oligomyrmex oertzeni var. aeolia : [3] ( Forel 1911).

as Oligomyrmex oertzeni var. aeolius : Turkey ( Ettershank 1966).

Chalepoxenus muellerianus (Finzi, 1922) : [21] ( Heinze 1987); [36] ( Schulz & Sanetra 2002).

Crematogaster auberti Emery, 1869 : Thrace (Radchenko 2007).

as Cremastogaster auberti : [16] ( Forel 1906); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994).

Crematogaster ionia Forel, 1911 : [12], [21] (Kiran & Aktaç 2006); Thrace (Radchenko 2007). as Cremastogaster scutellaris var. ionia : [3], [6], [16] ( Forel 1911); Bolkar Mt. ( Forel 1913). as Crematogaster scutellaris subsp. schmitti (sic!) var. ionia : [10] ( Schkaff 1924). as Crematogaster scutellaris st. schmidti var. ionia : [10] ( Santschi 1934).

as Cremastogaster ionia: Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994); [2] ( Aras & Aktaç 1992;1994); [19] ( Ülgentürk 2001).

Crematogaster laestrygon Emery, 1869

as Cremastogaster laestrygon : [77] ( Aktaç 1976); [1] ( Aras & Aktaç 1987); Thrace (Aktaç et al. 1994).

Crematogaster lorteti Forel, 1910 : [36] ( Santschi 1934).

Crematogaster schmidti (Mayr, 1853) : Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [4] (Lapeva- Gjonova & Kiran 2012).

as Cremastogaster scutellaris var. schmidti : [10] ( Forel 1895; 1911; Schkaff 1924); [70] ( Forel 1904). as Crematogaster scutellaris subsp. schmitti (sic!): [10] ( Schkaff 1924).

Crematogaster scutellaris (Olivier, 1792) : [10] ( Donisthorpe 1950a); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006); Thrace (Radchenko 2007).

as Cremastogaster scutellaris : [10] ( Emery 1870); Asia Minor ( André 1883a); [1] ( Forel 1895; Aras & Aktaç 1987); [3] ( Baroni Urbani 1964); [9] ( Aktaç 1976); [6] (Aktaç & Çamlıtepe 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994); [2] ( Aras & Aktaç 1992).

Crematogaster sordidula (Nylander, 1849) : [27] ( Emery 1897); [11] ( Donisthorpe 1950a); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [11], [12], [21] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008); [4] (Lapeva-Gjonova & Kiran 2012).

as Crematogaster sordidula var. flachi Forel, 1895 : [22] ( Emery 1897).

as Cremastogaster sordidula: Sultan Mt. ( Forel 1906) ; [81] ( Aktaç 1988); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994); [2] ( Aras & Aktaç 1992;1994).

as Cremastogaster sordidula var. flachi : [10], [16] ( Forel 1911); [27] ( Santschi 1921). as Crematogaster sordidula var. mayri (Mayr, 1853) : [10] ( Schkaff 1924).

as Cremastogaster sordidula subsp. mayri: [9] ( Aktaç 1976; 1978)

Crematogaster sordidula subsp. aeolia Forel, 1911: [3] ( Emery 1912). as Cremastogaster sordidula var. aeolia : [3] ( Forel 1911; 1913).

Crematogaster sordidula subsp. osmanica Santschi, 1921

as Cremastogaster sordidula var. osmanica : [27] ( Santschi 1921).

Formicoxenus nitidulus (Nylander, 1846) : [12] ( Schulz & Sanetra 2002).

Harpagoxenus sublaevis (Nylander, 1849) : [32] ( Heinze & Kauffmann 1993).

Leptothorax acervorum (Fabricius, 1793) : Thrace ( Agosti & Collingwood 1987); [73], [74] ( Aktaç 1988); [32] (Heinze et al. 1993).

Leptothorax muscorum (Nylander, 1846) : [59] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987); [74] ( Aktaç 1988); [47] ( Heinze 1988); [32] (Heinze et al. 1993); [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

Leptothorax scamni Ruzsky, 1905 : [25], [50], [70] (Heinze et al. 1993); [4] (Lapeva-Gjonova & Kiran 2012).

Manica rubida (Latreille, 1802) : [59] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987); [61], [73], [74] ( Aktaç 1988).

as Myrmica rubida : Turkey (in Asia) ( André 1883b); [70] ( Forel 1904); [74] ( Karavaiev 1926a).

Messor barbarus subsp. mediosanguineus Donisthorpe, 1946: Turkey ( Donisthorpe 1946); [53] ( Donisthorpe 1950a).

Comments: Messor barbarus subsp. mediosanguineus was collected by Donisthorpe from an area very near to the Turkish-Syrian border and is probably M. syriacus (see Tohmé & Tohmé 1981).

Messor caducus (Victor, 1839) : Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [11], [12] (Kiran & Aktaç 2006); [20] ( Lush 2009); [4] (Lapeva- Gjonova & Kiran 2012).

Messor dentatus Santschi, 1927

as Messor semirufus var. dentata : [77] ( Aktaç 1976).

Messor denticulatus Santschi, 1927 : [20] ( Lush 2009).

Messor ebeninus Santschi, 1927 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007); Turkey ( Kugler 1988).

Messor himalayanus (Forel, 1902) : [61] ( Aktaç 1988).

Comments: Arnol’di (1977) gives the distribution of this species as Himalayas and Afghanistan. This record appears to be doubtful for Turkey.

Messor intermedius Santschi, 1927

as Messor barbarus st. semirufus var. intermedia : [27] ( Santschi 1921).

Messor lobicornis subsp. rugosus (André, 1881)

as Stenamma barbarum r. capitatum v. rugosa : [16] ( Forel 1906).

Messor luridus Santschi, 1927

as Stenamma barbarum subsp. caducum var. lurida : [17] ( Emery 1898). as Messor barbarus subsp. semirufus var. lurida : [10] ( Schkaff 1924).

Messor meridionalis (André, 1883) : [3] ( Baroni Urbani 1964); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994); [2] ( Aras & Aktaç 1992;1994); [46] ( Arndt et al. 2005).

as Aphaenogaster barbara var. meridionalis : [10] ( André 1883a).

as Messor barbarus var. meridionalis : [17] ( Emery 1897); [10], [27], [52] ( Donisthorpe 1950a). as Stenamma barbarum r. capitatum v. meridionalis: Sultan Mt. , [16] ( Forel 1906). as Messor barbarus subsp. meridionalis : [3], [10] ( Forel 1911); [9] ( Emery 1921a). as Messor barbarus st. meridionalis : [3], [27] ( Santschi 1921).

as Messor barbarus subsp. instabilis var. meridionalis : [10] ( Schkaff 1924).

Messor oertzeni Forel, 1910 : [3] ( Forel 1910); [9] ( Emery 1921a); [15] ( Baroni Urbani & Aktaç 1981); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987; Aktaç et al. 1990); [2] ( Aras & Aktaç 1992); [12], [16], [21] (Kiran & Aktaç 2006). as Messor oertzeni var. amphigea Forel, 1911 : [3] ( Forel 1911); [10], [15], [20], [27] ( Donisthorpe 1950a). as Messor oerzenitor (sic!) var. amphigera (sic!): [27] ( Santschi 1921).

as Messor oerzeni (sic!): [2] (Aras & Aktaç 1994).

Messor oertzeni subsp. amasiensis Emery, 1921

as Messor oertzeni var. amasiensis : [44] ( Emery 1921a).

Messor orientalis ( Emery, 1898) : Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [61], [67], [71], [75], [76], [80], [81] ( Aktaç 1988).

as Stenamma structor var. orientalis : [36] ( Emery 1898).

as Messor barbarus subsp. structor var. orientalis : [3] ( Forel 1911).

as Messor structor st. rufitarsis var. orientalis : [36] ( Santschi 1926).

as Messor structor var. orientalis : [9] ( Aktaç 1976).

Messor semirufus (André, 1883) : [3], [10], [23], [27] ( Baroni Urbani 1974); [9], [77] ( Aktaç 1976; 1978); [15] ( Baroni Urbani & Aktaç 1981); [1], [4] ( Baroni Urbani et al. 1989); [1] (Aktaç et al. 1990). as Stenamma barbarum r. capitatum var. semirufa: Taurus Mts. ( Forel 1906) .

as Messor barbarus subsp. semirufa: [20] ( Donisthorpe 1950a).

Messor structor (Latreille, 1798) : [12], [17], [27] ( Emery 1897); [1] ( Aras & Aktaç 1987); [61], [67], [73], [74], [81] ( Aktaç 1988); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992;1994); [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012). as Aphaenogaster structor : [10] ( Forel 1895).

as Stenamma structor : [12] ( Forel 1906).

as Messor barbarus subsp. clivorum Ruzsky, 1905 : [3] ( Forel 1911).

as Messor barbarus subsp. structor var. mutica Nylander, 1849 : [10], [16] ( Forel 1911). as Messor barbarus subsp. structor var. tyrrhena ( Emery, 1898) : [3], [10] ( Forel 1911). as Messor barbarus subsp. varrialei Emery, 1921: [9] ( Emery 1921a).

as Messor barbarus subsp. rufitarsis Fabricius, 1804 : [10] ( Schkaff 1924). as Messor clivorum : [10] ( Schkaff 1924).

as Messor barbarus subsp. structor : [27], [76] ( Donisthorpe 1950a).

as Messor rufitarsis : [10] ( Collingwood 1963).

as Messor muticus: Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Messor structor subsp. aegaeus Santschi, 1926 : Thrace (Radchenko 2007). as Messor barbarus subsp. structor var. aegaea : [9] ( Emery 1921a).

as Messor structor var. aegaea : [10] ( Baroni Urbani 1964); [9] ( Aktaç 1976; 1978).

Messor sultanus Santschi, 1917 : Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992;1994).

Messor syriacus Tohmé, G. 1969

as Messor minor subsp. syriacus : Turkey (Arnol'di 1977).

New localities: [72], 05. VII.2010, 11 workers; Hasankeyf, 26. VI.2010, 3 workers, leg. F.Toprak, det. K.Kiran.

Messor testaceus Donisthorpe, 1950 : [63] ( Donisthorpe 1950b).

Messor wasmanni Krausse, 1910

as Messor barbarus st. semirufus var. concolor Forel ?: [27] ( Santschi 1921). as Messor barbarus subsp. semirufus var. concolor Emery, 1908 : [10] ( Schkaff 1924). as Messor semirufus var. concolor : Asia Minor ( Santschi 1927).

as Messor semirufus subsp. concolor : [3], [10] ( Baroni Urbani 1964). as Messor concolor: Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Monomorium dentigerum (Roger, 1862) : [3] ( Forel 1911); [9], [77] ( Aktaç 1976; 1978); [15] ( Baroni Urbani & Aktaç 1981); Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

as Holcomyrmex dentiger: Taurus Mts. ( Forel 1906) .

Comments: Before the revision of Radchenko (1997d) all similar species from Turkey and neighbouring countries were determined as M. dentigerum , but most probably these records are M. perplexum (Radchenko, pers. comm.).

Monomorium kusnezowi Santschi, 1928 : [77] ( Aktaç 1976; 1978).

Comments: Based on the data of Dlussky et al. (1990), we suggest that these records belong to M. ruzskyi Dlussky & Zabelin, 1985 . Monomorium ruzskyi is distributed in Transcaucasus and the mountains of the Middle Asia, while M. kusnezowi occur in Middle Asian deserts.

Monomorium monomorium Bolton, 1987 : [16] (Kiran & Aktaç 2006); Thrace (Radchenko 2007). as Monomorium minutum Mayr, 1855 : [3] ( Forel 1911); [10] ( Schkaff 1924); [9] ( Aktaç 1976); [15] ( Baroni Urbani & Aktaç 1981); [2] ( Aras & Aktaç 1992).

Monomorium perplexum Radchenko, 1997 : Turkey ( Radchenko 1997d).

as Monomorium glabrum (André, 1883) : [19] ( Donisthorpe 1950a – misidentification).

New localities: [46] (154m.), N 37° 03', E 35° 21', 22.VI.2004, 1 worker, leg and det. K.Kiran; [7] Gölmarmara- Musalar Village (653m.), N 38° 41', E 28° 08', 04.VII.2007, 29 workers, leg. C.Karaman, det. K.Kiran; Köprübaşı- Kozaklı Village (600m.), N 38° 43', E 28° 17', 05.VII.2007, 10 workers, leg. C.Karaman, det. K.Kiran; [15] Irlıganlı (465m.), N 37° 53', E 29° 11', 10.VII.2007, 3 workers, leg. C.Karaman, det. K.Kiran.

Comments: See the comments for M. glabrum in species excluded from the list of Turkish ants.

Monomorium pharaonis (Linnaeus, 1758) : [3] ( Forel 1911); [1] (Aktaç & Kiran 2006; Kiran & Karaman 2012); Thrace (Radchenko 2007).

Monomorium sommieri Emery, 1908

as Monomorium salomonis subsp. sommieri : Asia Minor ( Baroni Urbani 1971).

Comments: According to Baroni Urbani (1971), this species is distributed in nearly all of the Mediterranean region, in equatorial and austral Africa, in Asia Minor , the Caucasus, Central Asia, and in India and Burma. The type locality is Tunisia. Baroni Urbani's locality “ Asia Minor ” is not based on any Turkish material and no other record has been reported from Turkey. Therefore, the existence of this species in Turkey is doubtful.

Monomorium subopacum (Smith, F., 1858) : Asia Minor ( Baroni Urbani 1971).

Myrmecina graminicola (Latreille, 1802) : Asia Minor ( Stitz 1939; Kutter 1977); Anatolia ( Baroni Urbani 1971); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [11], [12] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

Myrmica anatolica Elmes, Radchenko & Aktaç, 2002 : [32], [59], [61], [66], [67] ( Elmes et al. 2002; Radchenko & Elmes 2010).

Myrmica bergi Ruzsky, 1902 : [59] ( Aktaç 1976).

Comments: This species is distributed in the Steppe Zone from the delta of the Danube River to Altai Mts., Transcaucasus, NE Iran, and the plains and mountains of Middle Asia ( Radchenko & Elmes 2010). Therefore this record is doubtful for Turkey.

Myrmica deplanata Emery, 1921 : Thrace ( Agosti & Collingwood 1987; Seifert 1988; Radchenko & Elmes 2010). as Myrmica lacustris Ruzsky, 1905 : Thrace (Radchenko 2007).

Myrmica gallienii Bondroit, 1920 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [4] (Lapeva-Gjonova & Kiran 2012).

Myrmica hellenica Finzi, 1926 : [26], [27], [30], [31], [50], [59], [66], [70], [73] (Seifert et al. 2009); Turkey ( Radchenko & Elmes 2010); [4] (Lapeva-Gjonova & Kiran 2012).

as Myrmica rugulososcabrinodis Karavaiev, 1929 : Northern Turkey ( Radchenko & Antanova 2004); [12], [16], [21] (Kiran & Aktaç 2006).

as Myrmica caucasica Arnol’di, 1934: Turkey ( Seifert 1988).

Myrmica jennyae Elmes, Radchenko & Aktaç, 2002 : [59], [66], [70] ( Elmes et al. 2002; Radchenko & Elmes 2010).

Myrmica lobicornis Nylander, 1846 : [74], [76] ( Donisthorpe 1950a); [59] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [61], [73], [74] ( Aktaç 1988).

Myrmica lonae Finzi, 1926 : Turkey (Seifert 2005); [4] (Lapeva-Gjonova & Kiran 2012).

Myrmica ravasinii Finzi, 1923 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [31] ( Seifert 1988); Turkey ( Radchenko & Elmes 2010).

Myrmica rubra (Linnaeus, 1758) : Thrace ( Agosti & Collingwood 1987; Radchenko 2007). as Formica rubra : [10] ( Rigler 1852).

as Myrmica laevinodis Nylander, 1846 : [74] ( Ruzsky 1905).

Myrmica ruginodis Nylander, 1846 : [59] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aktaç et al. 1994; Radchenko 2007); [61], [67], [73], [74], [81] ( Aktaç 1988); [12], [21] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008); Turkey ( Radchenko & Elmes 2010).

as Myrmica rubra subsp. ruginodis : [70], [73] ( Karavaiev 1926a).

Myrmica rugulosa Nylander, 1849 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007; Radchenko & Elmes 2010).

as Myrmica scabrinodis subsp. rugulosa : [73] ( Karavaiev 1926a).

Myrmica sabuleti Meinert, 1861 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [67], [71], [75] ( Aktaç 1988); Anatolia ( Seifert 1988); Turkey (Seifert 2005; Radchenko & Elmes 2010); [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

as Myrmica scabrinodis var. sabuleti : Turkey ( Finzi 1926).

Myrmica salina Ruzsky, 1905 : [19], [28], [42], [45], [67], [68], [74] ( Seifert 2011).

Myrmica scabrinodis Nylander, 1846 : Asia Minor ( André 1883b); [27] ( Santschi 1921); [73] ( Karavaiev 1926a); [3], [18] ( Donisthorpe 1950a); [59] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [1] ( Aras & Aktaç 1987); [61], [67], [73], [75], [81] ( Aktaç 1988); Turkey ( Radchenko & Elmes 2010); [4] (Lapeva-Gjonova & Kiran 2012).

Myrmica schencki Viereck, 1903 : [73] ( Karavaiev 1926a); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); North-eastern Turkey ( Radchenko & Elmes 2010).

Myrmica specioides Bondroit, 1918 : Thrace ( Agosti & Collingwood 1987); Turkey ( Radchenko & Elmes 2010); [1], [20], [28], [31], [51], [67], [79] ( Seifert 2011); [4] (Lapeva-Gjonova & Kiran 2012). as Myrmica scabrinodis var. turcica Santschi, 1931 : [27] ( Santschi 1931).

as Myrmica vandeli Bondroit, 1920 : Thrace ( Agosti & Collingwood 1987).

as Myrmica sancta Karavaiev, 1926 : [27] ( Seifert 1988).

as Myrmica turcica : [27] ( Seifert 1988); Thrace (Radchenko 2007).

as Myrmica bessarabica Nasonov, 1888 : Thrace (Radchenko 2007); Eastern Black Sea ( Ocak & Aktaç 2012). Comments: See the comments for M. vandeli in species excluded from the list of Turkish ants.

Myrmica spinosior Santschi, 1931 : [12], [43] (Seifert 2005).

Comments: According to Radchenko & Elmes (2010), records of Seifert (2005) for Turkey are doubtful.

Myrmica sulcinodis Nylander, 1846 : [12] ( Donisthorpe 1950a); [59] ( Aktaç 1976); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [67], [73], [74] ( Aktaç 1988); Anatolia ( Radchenko & Elmes 2010).

Myrmica tamarae Elmes, Radchenko & Aktaç, 2002 : [67] ( Elmes et al. 2002; Radchenko & Elmes 2010).

Myrmica tulinae Elmes, Radchenko & Aktaç, 2002 : [4], [70] ( Elmes et al. 2002; Radchenko & Elmes 2010); [4] (Lapeva-Gjonova & Kiran 2012).

as Myrmica tulini: Thrace (Radchenko 2007) .

Myrmoxenus gordiagini Ruzsky, 1902 : [3], [20], [28], [41] ( Schulz & Sanetra 2002).

Myrmoxenus kraussei ( Emery, 1915)

as Epimyrma kraussei : [20] ( Heinze 1987); [21] (Kiran & Aktaç 2006).

Myrmoxenus ravouxi (André, 1896) : [70], [73] ( Schulz & Sanetra 2002).

Myrmoxenus stumperi (Kutter, 1950) : [31] ( Schulz & Sanetra 2002).

Oxyopomyrmex krueperi Forel, 1911 : [1], [2] (Aktaç & Kiran 2004).

Oxyopomyrmex oculatus André, 1881 : [3] ( Forel 1911).

Pheidole megacephala (Fabricius, 1793) : [3], [15], [20] ( Donisthorpe 1950a).

Pheidole pallidula (Nylander, 1849) : [1] ( Forel 1895; Aras & Aktaç 1987); [12], [17], [27] ( Emery 1897); Sultan Mt., [16] ( Forel 1906); [3], [10], [16] ( Forel 1911); [3], [10], [12], [20], [27], [28] ( Donisthorpe 1950a); [3] ( Baroni Urbani 1964); [9], [77] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [6] (Aktaç & Çamlıtepe 1987); [74], [76], [80], [81] ( Aktaç 1988); [2] ( Aras & Aktaç 1992;1994); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006); [20] ( Lush 2009); [1] (Kiran & Karaman 2012); [4] (Lapeva-Gjonova & Kiran 2012).

as Pheidole pallidula st. arenarium (sic!) var. orientalis Emery, 1915 : [3], [27] ( Santschi 1921); [10] ( Santschi 1934).

as Pheidole pallidula subsp. arenarum var. orientalis : [10] ( Schkaff 1924).

Pheidole teneriffana Forel, 1893 : [3] ( Forel 1911).

Solenopsis fugax (Latreille, 1798) : [22] ( Emery 1897); Sultan Mt. ( Forel 1906); [10] ( Forel 1911; Schkaff 1924); [10], [18] ( Donisthorpe 1950a); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [11], [12], [16], [21] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

as Diplorhoptrum fugax : [59] ( Aktaç 1976); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994); [2] ( Aras & Aktaç 1992).

Solenopsis latro Forel, 1894 : Thrace ( Agosti & Collingwood 1987).

Solenopsis orbula Emery, 1875

as Diplorhoptrum orbulum : [67] ( Aktaç 1988).

Stenamma debile (Förster, 1850) : [11] ( DuBois 1998); Thrace (Radchenko 2007); [20] ( Rigato 2011). as Stenamma westwoodii Westwood, 1839 : [12] (Kiran & Aktaç 2006 – misidentification).

Comments: See the comments for S. westwoodii in species excluded from the list of Turkish ants.

Stenamma striatulum Emery, 1895 : [59] ( DuBois 1998); [55], [70] ( Rigato 2011); [4] (Lapeva-Gjonova & Kiran 2012).

as Stenamma westwoodii Westwood, 1839 : [11] (Kiran & Aktaç 2006 – misidentification). Comments: See the comments for S. westwoodii in species excluded from the list of Turkish ants.

Strongylognathus cf. alpinus Wheeler, 1909 : [45] ( Schulz & Sanetra 2002).

Strongylognathus huberi Forel, 1874 : [21] (Kiran & Aktaç 2006).

Strongylognathus kervillei Santschi, 1921 : [27] ( Santschi 1921); [46], [49], [53], [63] ( Schulz & Sanetra 2002).

Strongylognathus silvestrii Menozzi, 1936 : [36], [45] ( Schulz & Sanetra 2002).

Strongylognathus testaceus (Schenck, 1852) : [2] ( Aras & Aktaç 1992); [9], [28], [31], [45] ( Schulz & Sanetra 2002).

Strumigenys argiola (Emery, 1869)

as Epitritus argiolus : [10] ( Schkaff 1924).

as Pyramica agriola (sic!): Thrace (Radchenko 2007).

Strumigenys baudueri (Emery, 1875) : [10] ( Bondroit 1918; Emery 1916; 1922). as Smithistruma baudueri : Turkey ( Baroni Urbani 1971)

as Pyramica baudueri: Thrace (Radchenko 2007) .

Strumigenys membranifera Emery, 1869

as Trichoscapa membranifera : Turkey ( Kugler 1988).

Strumigenys tenuipilis Emery, 1915

as Smithistruma baudueri var. tenuipilis : [10] ( Bernard 1968).

Temnothorax aeolius ( Forel, 1911)

as Leptothorax bulgaricus subsp. aeolius : [3] ( Forel 1911). as Leptothorax bulgaricus var. aeolia: Menderes river ( Forel 1913).

Temnothorax affinis (Mayr, 1855) : [4] (Lapeva-Gjonova & Kiran 2012). as Leptothorax affinis : [12] ( Heinze 1988; Kiran & Aktaç 2006).

Temnothorax antigoni ( Forel, 1911)

as Leptothorax antigoni : [3] ( Forel 1911).

Temnothorax artvinensis Seifert, 2006

as Temnothorax artvinense : [70] (Seifert 2006).

Temnothorax bulgaricus ( Forel, 1892)

as Leptothorax bulgaricus: Bolkar Mt. ( Forel 1913) ; Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [3], [8], [11], [12], [13], [20] ( Heinze 1988); [11], [12], [16], [21] (Kiran & Aktaç 2006).

Temnothorax bulgaricus subsp. smyrnensis ( Forel, 1911)

as Leptothorax bulgaricus subsp. smyrnensis: [3] ( Forel 1911).

Temnothorax corticalis (Schenck, 1852)

as Leptothorax corticalis: Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Temnothorax crassispinus (Karavaiev, 1926) : [4] (Lapeva-Gjonova & Kiran 2012).

as Leptothorax crassispinus: Thrace (Radchenko 2007) .

Comments: The name T. crassispinus was considered a junior synonym of T. nylanderi for many years until Radchenko (2000) separated these two species. T. crassispinus has mainly an East European and Caucasian distribution, while T. nylanderi has a Central and West European distribution. Considering the geographical position of Turkey, old records referring to T. nylanderi may also be relevant to T. crassispinus (or even some other species) and require further investigation.

Temnothorax dessyi ( Menozzi, 1936) : Sultan Mts. ( Kiran & Aksoy 2011); Kaz Mt. ( Aktaç & Karaman 2012).

Temnothorax exilis (Emery, 1869)

as Leptothorax exilis : Asia Minor ( Baroni Urbani 1971); Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [3] ( Heinze 1988); [12], [21] (Kiran & Aktaç 2006).

as Leptothorax exilis var. darii Forel, 1911 : [3] ( Forel 1911).

Temnothorax graecus ( Forel, 1911)

as Leptothorax bulgaricus subsp. graeca: [11] ( Donisthorpe 1950a). as Leptothorax graecus : [2] ( Aras & Aktaç 1992); Thrace (Radchenko 2007).

Temnothorax interruptus (Schenck, 1852) : [4] (Lapeva-Gjonova & Kiran 2012).

Temnothorax kemali ( Santschi, 1934)

as Leptothorax kemali : [3] ( Santschi 1934).

Temnothorax korbi (Emery, 1924) : Kaz Mt. ( Aktaç & Karaman 2012).

Temnothorax laestrygon ( Santschi, 1931) : Kaz Mt. ( Aktaç & Karaman 2012).

Temnothorax luteus (Forel, 1874) : [1] (Kiran & Karaman 2012). as Leptothorax tuberum r. luteus : [1] ( Forel 1895); Bolkar Mt. ( Forel 1913). as Leptothorax tuberum subsp. luteus : [3] ( Forel 1911).

as Leptothorax luteus : [11], [12], [16], [21] (Kiran & Aktaç 2006).

Temnothorax nadigi (Kutter, 1925)

as Leptothorax nadigi : [25], [31], [60], [61], [67], [70] ( Czechowska et al. 1998); [21] (Kiran & Aktaç 2006).

Temnothorax nigriceps (Mayr, 1855)

as Leptothorax nigriceps: Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [37] ( Heinze 1988).

Temnothorax nylanderi (Förster, 1850) : [4] (Lapeva-Gjonova & Kiran 2012).

as Leptothorax nylanderi : [10] ( Forel 1895); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aktaç et al. 1994); [13], [25], [47] ( Heinze 1988); [11], [12], [16], [17], [21] (Kiran & Aktaç 2006). Comments: See the comments for T. crassispinus .

Temnothorax parvulus (Schenck, 1852) : [4] (Lapeva-Gjonova & Kiran 2012). as Leptothorax parvulus: Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [12], [21], [25] ( Heinze 1988); [11], [16], [21] (Kiran & Aktaç 2006).

Temnothorax recedens (Nylander, 1856) : [4] (Lapeva-Gjonova & Kiran 2012). as Leptothorax recedens: Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [12], [16] (Kiran & Aktaç 2006).

Temnothorax rottenbergii subsp. jesus ( Forel, 1913)

as Leptothorax rottenbergi (sic!) var. jesus : [3] ( Forel 1913).

Temnothorax schoedli Seifert, 2006 : [49] (Seifert 2006).

Temnothorax semiruber (André, 1881) : [4] (Lapeva-Gjonova & Kiran 2012). as Leptothorax semiruber: Thrace ( Agosti & Collingwood 1987; Radchenko 2007); [27], [31] ( Heinze 1988); [12], [16], [21] (Kiran & Aktaç 2006).

Temnothorax sordidulus (Müller, 1923)

as Leptothorax sordidulus: Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Temnothorax tuberum (Fabricius, 1775) : [4] (Lapeva-Gjonova & Kiran 2012).

as Leptothorax tuberum : Asia Minor ( André 1883b); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aktaç et al. 1994; Radchenko 2007); [61], [67], [73], [74] ( Aktaç 1988); [11], [12], [16], [21] (Kiran & Aktaç 2006).

Temnothorax turcicus ( Santschi, 1934)

as Leptothorax turcicus : [3] ( Santschi 1934).

Temnothorax unifasciatus (Latreille, 1798) : [4] (Lapeva-Gjonova & Kiran 2012). as Leptothorax unifasciatus : [12] ( Emery 1897); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [10], [21], [25], [59] ( Heinze 1988); [11], [12], [16] (Kiran & Aktaç 2006).

as Leptothorax tuberum st. unifasciatus : [3] ( Santschi 1921).

as Leptothorax tuberum subsp. unifasciatus var. unifasciato-interrupta Forel, 1874: [10] ( Schkaff 1924).

Tetramorium anatolicum Csösz & Schulz, 2007 : [67], [80] ( Csösz et al. 2007).

Tetramorium brevicorne Bondroit, 1918

as Tetramorium caespitum var. brevicornis : [26] ( Donisthorpe 1950a).

Comments: This species has a very limited distribution in islands of Corsica, Sardinia and Sicily ( Sanetra et al. 1999). Its existence in Turkey is doubtful, with older material probably misidentified.

Tetramorium caespitum (Linnaeus, 1758) : [1] ( Forel 1895; Aras & Aktaç 1987; Kiran & Karaman 2012); [2], [12], [17], [27] ( Emery 1897); [12], Sultan Mt. ( Forel 1906); [3], [10] ( Forel 1911; Baroni Urbani 1964); [27] ( Santschi 1921); [10] ( Schkaff 1924); [6], [12], [15], [70], [74], [76] ( Donisthorpe 1950a); [59] ( Aktaç 1976; 1978); Thrace ( Agosti & Collingwood 1987; Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [6] (Aktaç & Çamlıtepe 1987); [61], [67], [71], [73], [74], [75], [76], [80], [81] ( Aktaç 1988); [2] ( Aras & Aktaç 1992;1994); [11], [12], [16] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012).

as Formica cespitum (sic!): [10] ( Rigler 1852).

Comments: Schlick-Steiner et al. (2006) indicated in their study on cryptic species of Tetramorium in the Western Palaearctic that the Tetramorium caespitum / impurum complex comprises seven Palaearctic species. The taxonomy of this complex in the Palaearctic region has not been clarified yet and needs further investigation in Turkey.

Tetramorium caespitum subsp. flavidulum Emery, 1924

as Tetramorium caespitum caespitum var. flavidula : Asia Minor ( Emery 1909a); [36] ( Emery 1925d).

Tetramorium chefketi Forel, 1911 : Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992;1994); [31] ( Schulz 1996); [28], [31], [37], [42], [70] ( Güsten et al. 2006); [11], [12], [16], [21] (Kiran & Aktaç 2006); [3], [10], [13], [15], [17], [20], [27], [28], [31], [34], [35], [36], [37], [39], [45], [51], [55], [64], [70], [74] ( Csösz et al. 2007); [4] (Lapeva-Gjonova & Kiran 2012).

as Tetramonium (sic!) caespitum var. forte Forel, 1904 : [70] ( Forel 1904).

as Tetramorium fortis : [10] ( Emery 1909a); [6] (Aktaç & Çamlıtepe 1987); [1] ( Aras & Aktaç 1987); Thrace (Çamlıtepe & Aktaç 1987; Aras & Aktaç 1990); [2] ( Aras & Aktaç 1992).

as Tetramorium caespitum var. chefketi : [10] ( Forel 1911).

as Tetramorium caespitum var. sarkissiani Forel, 1911 : [16] ( Forel 1911).

as Tetramorium caespitum st. turcomana Emery, 1909: [27] ( Santschi 1921).

as Tetramorium caespitum subsp. ferox var. chefketi : [10] ( Schkaff 1924).

as Tetramorium caespitum var. fortis : Turkey ( Baroni Urbani 1971).

as Tetramorium chefteki (sic!): Thrace ( Agosti & Collingwood 1987); [27] ( Özdemir et al. 2008). as Tetramorium forte: Thrace ( Agosti & Collingwood 1987; Aktaç et al. 1994; Radchenko 2007); [11], [12], [16], [21] (Kiran & Aktaç 2006); [27] ( Özdemir et al. 2008).

as Tetramorium sarkissiani : Turkey ( Collingwood & Heatwole 2000).

Comments: See the comments for Tetramorium forte in species excluded from the list of Turkish ants.

Tetramorium davidi Forel, 1911 : Thrace ( Agosti & Collingwood 1987; Radchenko 2007).

Tetramorium densopilosum Radchenko & Arakelian, 1990 : [75] ( Csösz & Schulz 2010).

Tetramorium diomedeum Emery, 1908 : Western Turkey ( Sanetra et al. 1999); [9], [20], [36], [45], [49], [53] ( Csösz & Schulz 2010).

as Tetramorium caespitum subsp. ferox var. laevior Forel, 1911 : [10], [16] ( Forel 1911). as Tetramorium ferox var. laevior : [10] ( Finzi 1939); Thrace ( Sanetra et al. 1999).

Tetramorium ferox Ruzsky, 1903 : [3] ( Santschi 1934); Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992); [27], [51] ( Ülgentürk 2001); [11], [12], [16], [21] (Kiran & Aktaç 2006); [13], [28], [31], [34], [36], [42], [45], [51], [74] ( Csösz & Schulz 2010); [4] (Lapeva- Gjonova & Kiran 2012).

as Tetramorium caespitum st. ferox var. perspicax Santschi, 1921 : [27] ( Santschi 1921). as Tetramorium caespitum ferox : Asia Minor ( Emery 1925d).

as Tetramorium perspicax : Asia Minor ( Radchenko 1992).

Tetramorium feroxoide Dlussky & Zabelin, 1985 : Thrace (Radchenko 2007); [45], [53], [74] ( Csösz & Schulz 2010).

Tetramorium goniommoide Poldi, 1979 : [15] ( Poldi 1979; Baroni Urbani & Aktaç 1981); [21] (Kiran & Aktaç 2006).

Tetramorium hippocrate Agosti & Collingwood, 1987

as Tetramorium caespitum semilaeve var. hippocratis : [9] ( Emery 1921a); [10] ( Schkaff 1924). as Tetramorium semilaeve var. hippocratis : [9] ( Aktaç 1976; 1978).

as Tetramorium hippocratis: Thrace ( Agosti & Collingwood 1987; Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] ( Aras & Aktaç 1992).

Tetramorium hungaricum Röszler, 1935 : [4] (Lapeva-Gjonova & Kiran 2012).

Tetramorium impurum (Förster, 1850) : Thrace ( Agosti & Collingwood 1987; Radchenko 2007). as Tetramorium caespitum var. impura : [10] ( Schkaff 1924).

Comments: According to Schlick-Steiner et al. (2006), T. impurum is a Northwest and Central European species. The occurrence of the species in Turkey does not fit with its distribution and its existance in Turkey is doubtful.

Tetramorium lucidulum Menozzi, 1933 : Thrace ( Agosti & Collingwood 1987; Aktaç et al. 1994; Radchenko 2007); [11], [12], [16] (Kiran & Aktaç 2006); [4] (Lapeva-Gjonova & Kiran 2012). as Tetramorium caespitum punicum var. lucidula : Asia Minor ( Emery 1909a).

as Tetramorium caespitum st. punicum var. lucidula : [27] ( Santschi 1921).

as Tetramorium caespitum subsp. punica var. lucidula : Asia Minor ( Emery 1922). as Tetramorium lucidula : [77] ( Aktaç 1976; 1978); Thrace ( Aras & Aktaç 1990); [28], [77] ( López 1991); [2] (Aras & Aktaç 1994).

Tetramorium moravicum Kratochvil, 1941 : [73], [80] ( Güsten et al. 2006); [64], [66], [67], [74], [75], [76], [80] ( Csösz et al. 2007); [4] (Lapeva-Gjonova & Kiran 2012).

Tetramorium punicum (Smith, F., 1861) : Thrace ( Agosti & Collingwood 1987; Radchenko 2007). as Tetramorium caespitum subsp. punicum : [12] ( Emery 1897).

as Tetramorium caespitum var. punica : [3], [16] ( Forel 1911).

as Tetramorium caespitum st. punicum : [27] ( Santschi 1921).

Tetramorium rhodium Emery, 1924 : [3], [15] ( Csösz et al. 2007).

Tetramorium schmidti Forel, 1904 : [77] ( Aktaç 1976).

Tetramorium semilaeve André, 1883 : [3] ( Baroni Urbani 1964); [9] ( Aktaç 1978); [6] (Aktaç & Çamlıtepe 1987); [1] ( Aras & Aktaç 1987); [61], [67], [71], [74], [75], [76], [80], [81] ( Aktaç 1988); Thrace ( Aras & Aktaç 1990; Aktaç et al. 1994; Radchenko 2007); [2] (Aras & Aktaç 1994); [20] ( Csösz & Markó 2004); [11], [12], [16], [17] (Kiran & Aktaç 2006).

as Tetramorium caespitum subsp. semilaeve : [12] ( Emery 1897); [45] ( Kohl 1905). as Tetramorium caespitum v. semilevis (sic!): [3], [10], [16] ( Forel 1911).

as Tetramorium caespitum subsp. semilaevis: [12], [15], [53] ( Donisthorpe 1950a). as Tetramorium semilaeve semilaeve var. siciliensis Santschi, 1927 : [9] ( Aktaç 1976). as Tetramorium semileave (sic!): Thrace (Çamlıtepe & Aktaç 1987).

as Tetramorium semilaeve siciliensis : [9] ( López 1991).

Tetramorium semilaeve subsp. galaticum Menozzi, 1936: [27] ( Menozzi 1936). as Tetramorium caespitum st. biskrensis var. galatica : [27] ( Santschi 1921).

Tetramorium syriacum Emery, 1924 : [53], [58] ( Csösz et al. 2007).