Corticotomus Sharp, 1891

Kolibac, Jiri, 2013, Trogossitidae: A review of the beetle family, with a catalogue and keys, ZooKeys 366, pp. 1-194 : 44-45

Genus Corticotomus Sharp, 1891 Figs 1 , 6 ; Map 5

Corticotomus Sharp, D. 1891: 390.

Type species.

Corticotomus basalis Sharp, 1891 [designated by Barron 1971]

Léveillé, A. 1910: 7. Downie, N. M. & Arnett, R. H. Jr. 1996: 936 (key). Kolibáč, J. 2005: 50 (redescription). Kolibáč, J. 2006: 111 (phylogeny). Lepesme, P. & Paulian, R. 1944: 138 ( Colydobius Sharp, 1891) (key). Léveillé, A. 1910: 20 ( Colydobius Sharp, 1891)

Colydobius Sharp, 1891 [type species: Colydobius divisus Sharp, 1891; designated by Kolibáč 2005]

Léveillé, A. 1910: 20. Kolibáč, J. 2005: 50 (synonymized)

Leveillesoma Lepesme & Paulian, 1944 [type species: Nemosomia fulva Léveillé, 1905; by original designation]

Lepesme, P. & Paulian, R. 1944: 139. Kolibáč, J. 2005: 50 (synonymized)

Parafilumis Casey, 1916 [type species: Parafilumis estriata Casey, 1916; by original designation and monotypy]

Barron, J. R. 1971: 53 (synonymized)

Description.

Body size: about 3.0-5.0 mm. Body shape elongate. Gular sutures narrow, subparallel at apex. Frontoclypeal suture absent. Frons: longitudinal groove or depression absent. Cranium ventrally: tufts of long setae at sides absent. Submentum: ctenidium absent. Antennal groove present. Eyes: size moderate. Eyes number: two. Epicranial acumination deep. Lacinial hooks absent. Galea: shape elongate. Galea: ciliate setae present. Mediostipes-Lacinia not fused. Palpifer: outer edge denticulate. Mandibular apical teeth number: two, horizontally situated. Mola reduced but present. Penicillus (at base) present (fine, often membranous). Pubescence above mola or cutting edge absent. Ventral furrow present. Basal notch shallow or absent. Labrum-Cranium not fused. Epipharyngial sclerite present. Lateral tormal process: projection curved downwards, processes not connected ( Airora ). Ligula: ciliate setae present. Ligula rigid, not retroflexed, weakly emarginate. Hypopharyngeal sclerite sickle shaped. Antenna 11-segmented. Antennal club asymmetrical, sensorial fields present. Front coxal cavities externally closed, internally open. Pronotum elongate. Prepectus absent. Middle coxal cavities open. Elytra: long hairs absent. Epipleuron moderate. Elytral interlocking mechanism present, carinae reduced. Elytral punctation regular, scales absent. Wing: radial cell open (outer vein present), wedge cell absent, cross vein MP3-4 absent, cross vein AA1+2-3+4 absent. Front tibiae: spines along side large. Hooked spur present. Claws: denticle absent. Spiculum gastrale absent. Tegmen composed of two parts. Coxitae undivided.

Biology.

Predatory. North American species have been both reared and collected from various species of pine and willow, also from Rhus , Pseudotsuga , and other trees ( Barron 1971, Dajoz 1997). The Corticotomus species were mostly found under bark but also in the burrows of Cryphalus ( Scolytinae ) ( Barron 1971). Dajoz (1997) noted Corticotomus apicalis associated with the bark beetle Chaetophloeus parkinsoniae . The way of life of Corticotomus is probably similar to that of Nemozoma .

Distribution.

Distributed from Brazil to Canada. One species, unknown to me, is described from Chile.

Species:

Corticotomus apicalis Van Dyke, 1944; Western USA (JRB)

Barron, J. R. 1971: 60. Dajoz, R. 1997: 42 (biology). Van Dyke, E. C. 1944: 151

Corticotomus basalis Sharp, 1891; Guatemala (AL)

Léveillé, A. 1910: 7. Kolibáč, J. 2005: 50

Corticotomus bicolor Léveillé, 1895; Chile (AL)

Léveillé, A. 1910: 7

Corticotomus californicus Van Dyke, 1915; Western USA (JRB)

Barron, J. R. 1971: 60 (syn. Parafilumis estriata Casey, T. L. 1916: 207, 283; synonymized by whom?). Van Dyke, E. C. 1915: 28. Schaeffer, C. F. A. 1920: 193

Corticotomus caviceps Fall, 1910; Western Canada and USA (JRB, varA)

Barron, J. R. 1971: 58. Dajoz, R. 1997: 41 (diagnosis). Barron, J. R. 1971: 58 (syn. Corticotomus laeviventris Casey, 1916; synonymized by whom?). Fall, H. C. 1910: Lepesme, P. & Paulian, R. 1944: 140 ( Nemosoma s.str. caviceps Fall, 1910)

Corticotomus cylidricus LeConte, 1863; Eastern USA: Iowa (JRB)

Léveillé, A. 1910: 5 ( Nemosoma ). Barron, J. R. 1971: 57 (syn. Corticotomus cylindricus var. texanus Schaeffer, 1918). Schaeffer, C. F. A. 1920: 193 ( Corticotomus cylindricus subsp. texanus Schaeffer, C. F. A. 1918: 192). Barron, J. R. 1971: 56. Van Dyke, E. C. 1915: 26. Kolibáč, J. 2005: 51 (redescription). Böving, A. G. & F. C. Craighead, 1931: 273 (larva). Reitter, E. 1876: 14 ( Nemosoma ). Lepesme, P. & Paulian, R. 1944: 140 ( Nemosoma s.str.)

Corticotomus depressus Schaeffer, 1918; Eastern USA (JRB)

Barron, J. R. 1971: 54. Schaeffer, C. F. A. 1918: 192

Corticotomus divisus Sharp, 1891; Panama (AL)

Léveillé, A. 1910: 20 ( Colydobius ). Kolibáč, J. 2005: 51 (combination)

Corticotomus dufaui Léveillé, 1907; Guadeloupe (AL)

Léveillé, A. 1910: 20. Kolibáč, J. 2005: 52 (redescription)

Corticotomus fulva Léveillé, 1905; Brazil (AL)

Léveillé, A. 1910: 4 ( Nemosomia ). Kolibáč, J. 2005: 50 (combination). Lepesme, P. & Paulian, R. 1944: 139 ( Nemosomia ) (combined with Leveillesoma )

Corticotomus parallelum Melsheimer, 1844; Eastern USA (JRB)

Léveillé, A. 1910: 6 ( Nemosoma ). Barron, J. R. 1971: 55. Lepesme, P. & Paulian, R. 1944: 140 ( Nemosoma ). Reitter, E. 1876: 14 ( Nemosoma )

Corticotomus quadrimaculatus Léveillé, 1894; Brazil (AL)

Léveillé, A. 1910: 7

Corticotomus sharpi Léveillé, 1905; Mexico (AL)

Léveillé, A. 1910: 7

Corticotomus signatus Sharp, 1891; Guatemala (AL)

Léveillé, A. 1910: 20

Corticotomus testaceus Dajoz, 1997; USA: Arizona (AD)

Dajoz, R. 1997: 40

Figures

Figure 1. Adult morphology: A Alindria sp. from Laos, ventral surface B Leipaspis lauricola, labium C Airora cylindrica, antennal club D Corticotomus cylindricus, mandible dorsally E Airora cylindrica, mandible ventrally F Acalanthis quadrisignata, labium G Acalanthis quadrisignata, maxilla H Acalanthis quadrisignata, wing I Peltonyxa deyrollei, tegmen composed of 3 parts J Airora cylindrica, tegmen composed of 2 parts.

Map 5. A distribution of the tribe Trogossitini.

Figure 6. A Alindria spectabilis B Alindria elongata C Alindria sp., Thailand D Airora cylindrica E Airora (syn. Temnochilodes) dugesi F Corticotomus (syn. Colydobius) divisus G Elestora fulgurata.