Tanindrazanus anjozorobeus, Forthman & Chłond & Weirauch, 2016

Forthman, Michael, Chłond, Dominik & Weirauch, Christiane, 2016, Taxonomic Monograph Of The Endemic Millipede Assassin Bug Fauna Of Madagascar (Hemiptera: Reduviidae: Ectrichodiinae), Bulletin of the American Museum of Natural History 2016 (400), pp. 1-1 : 1-

Tanindrazanus anjozorobeus , new species

Plates 2, 6; map 9

DIAGNOSIS: Males are recognized among other Tanindrazanus species by the tricolor body (red, orange, and brown coloration), elevated clypeal apex relative to the labrum, synthlipsis smaller than the width of the eye, and broad postocular in dorsal view. This species is very similar to T. kathrynae , but it can be easily distinguished by the larger body size, small width of the synthlipsis, larger eyes that surpass the dorsal and ventral margins of the head in lateral view, very large ocelli separated by less than one-third of the diameter of an ocellus, and lack of anterolateral projections on the pronotum.

DESCRIPTION: Large body size (length: 17.32 mm, holotype; 16.20–17.32 mm). COLOR-

ATION: Red with pale orange-brown mandibular and maxillary plates, clypeus, dorsal postocular near ocellar tubercle, labium, anterior pronotal lobe except lateral margins, tibiae except apically, and corium except distally. Remainder of head, lateral margins of anterior pronotal lobe, anterior part of propleura, meso- and metapleura, scutellum, corium distally, forewing membrane, femora and apical part of tibiae, ventrolateral spot on abdominal sternites

MAP 9. Localities of Tanindrazanus amboasaricus, II –V, and pygophore dark brown to black. VES-

Ta. anjozorobeus , and Toliarus (To.) species. TITURE: As in generic description. STRUC-

TURE: HEAD: Ventrally with shallow

HOLOTYPE: Male: Madagascar: Toliara: Parc anteromedial depression; clypeal apex dorsally

National d’Andohahela, Ihazofotsy Parcelle III, elevated relative to labrum (pl. 13G, H);

24.83083 ° S 46.53616 ° E, 80 m, 16 Dec 2002 –26 postclypeus with relatively deep, broad medial

Dec 2002, M. Irwin, F. Parker, R. Harin’Hala longitudinal depression to middle of interocular

(00044865) ( CAS). area; synthlipsis smaller than the width of eye;

PARATYPES: Madagascar: Antananarivo: 46 km interocular sulcus posterior to hind margin of

NE of Ankazobe: Ambohitantely, 18.198 ° S eye; postocular broad in dorsal view (pl. 13A);

47.2815 ° E, 700 m, 17 Oct 2003 – 24 Oct 2003, M.

Irwin, R. Harin’Hala, 13 (00006126) ( BMNH), 13 ocelli very large, separated by less than one-

(00045559) ( USNM); 28 Dec 2003 – 10 Jan 2004, M. third diameter of ocellus, located on distinct

Irwin, R. Harin’Hala, 13 (00006093) ( UCR); 10 Jan median tubercle (pl. 13H); eye about one-third

2004– 20 Jan 2004, M. Irwin, R. Harin’Hala, 13 of head length, distinctly surpassing dorsal and

(00007177) ( CAS), 13 (00045466) (SU). Maha- ventral head surfaces; pedicel about one-fourth

janga: 160 km N of Maevatanana on RN 04, Ampi- longer than scape; labial segment III subequal

joroa National Park, 16.31933 ° S 46.81333 ° E, 43 m,

07 Feb 2005 – 19 Feb 2005, M. Irwin, R. Harin’Hala, to II. THORAX: Anterior pronotal lobe without

13 (00045044) ( CAS). Unknown: 23 (00044823, anterolateral projections; posterior pronotal

00044877) ( CAS). lobe smooth; pronotal transverse furrow con-

2016 FORTHMAN ET AL.: MILLIPEDE ASSASSIN BUG FAUNA 73

tinuous; legs slender; tarsomeres I and II com- apically, and tarsi. Meso- and metasterna medially bined subequal to III; hemelytron reaching or and abdomen, except pygophore, pale red. Distal surpassing abdominal apex. ABDOMEN: Apex half of antennal segment V and segments VI and rounded ; dorsal laterotergite II not expanded; VII white. VESTITURE: As in generic description. intersegmental sutures carinulate between ster- STRUCTURE: HEAD: Ventrally with shallow nites II – V and laterally between V and VI ; anteromedial depression; clypeal apex dorsally elepygophore process subtriangular in lateral view vated relative to labrum (pl. 13G, H); postclypeus (pl. 20H, I); BPE longer than basal plate; area of with relatively deep, broad medial longitudinal endosomal struts–DPS fusion ovate. depression to middle of interocular area; postocular ETYMOLOGY: The species epithet is a noun in narrow in dorsal view (pl. 13A) ; synthlipsis about apposition and is named after the type locality width of eye; interocular sulcus near hind margin Anjozorobe , Madagascar. of eye (pl. 13B) ; eyes about one-third of head DISTRIBUTION: Anjozorobe in the Antanana- length, not reaching dorsal and ventral head surrivo province (map 9). faces ; ocelli large, separated by less than diameter DISCUSSION: Coloration varies with the of ocellus, located on distinct median tubercle (pl. pronotal longitudinal furrow either light 13H) ; pedicel about one-fourth longer than scape; brown or entirely black and the abdominal labial segment III subequal to II. THORAX: Anterior sternites dark brown. pronotal lobe with very small anterolateral projec- HOLOTYPE: Male: Madagascar: Antanana- tions (pl. 15B) ; posterior pronotal lobe transversely rivo: Anjozorobe District , Anjozorobe, 18.2619 ° S striated ; pronotal transverse furrow continuous; 47.6849 ° E, 1091 m, Dec 1938 ( MNHN). legs slender ; tarsomeres I and II combined sub- PARATYPES: Madagascar: Antananarivo: Anjozo- equal to III on fore- and mid legs, longer on hind robe District, Anjozorobe, 18.2619 ° S 47.6849 ° E, 1091 legs ; hemelytron reaching or surpassing abdominal m, Dec 1938, 33 ( MNHN) .

apex. ABDOMEN: Apex rounded; dorsal laterotergite

II not expanded; intersegmental sutures carinulate Tanindrazanus antananarivo , new species between sternites II – V and laterally between V and

Plates 2, 6, 9; map 6 VI; pygophore process subtriangular in lateral view DIAGNOSIS: Males recognized among other (pl. 20H, I); BPE as long as basal plate; area of Tanindrazanus species by a combination of the endosomal struts–DPS fusion ovate (pl. 21H). following characters: the dark brown color with ETYMOLOGY: The species epithet is a noun in pale red abdomen, synthlipsis about the width of apposition and is named after the type locality the eye, eye not reaching the dorsal and ventral province Antananarivo, Madagascar. head margins, and continuous pronotal trans- DISTRIBUTION: Northern region of Antananaverse furrow. This species is similar to T. margin- rivo province (map 6). atus and T. tenebricus , but is easily distinguished DISCUSSION: The dorsal surface of the antenby the dark brown and pale red color pattern and nifers and postclypeus are light brown in some intersegmental sutures carinulate between ster- specimens. nites II – V and laterally between V and VI. HOLOTYPE: Male: Madagascar: Antanana- DESCRIPTION: Medium body size (length: 12.52 rivo: 46 km NE of Ankazobe: Ambohitantely, mm, holotype; 12.41–14.00 mm). COLORATION: 18.198 ° S 47.2815 ° E, 700 m, 16 Jan 2005 –27 Jan Dark brown with light brown clypeus, antennifer 2005, M. Irwin, R. Harin’Hala (00006330) ( CAS). except apically, postocular and neck dorsomedially, PARATYPES: Madagascar: Antananarivo: 46 km NE head posteroventrally, anteocular laterally and ven- of Ankazobe: Ambohitantely, 18.198 ° S 47.2815 ° E, 700

m, 24 Oct 2003 – 31 Oct 2003, M. Irwin, R. Harin’Hala, trally, labium, basal half of antennal segment V, 13 (00005367) ( CAS); 07 Nov 2003 – 15 Nov 2003, M. anterior margin of pronotum, prosternum, corium Irwin, R. Harin’Hala, 13 (00006334) ( CAS); 22 Nov basally, coxae, trochanters, femora basally, tibiae 2003– 29 Nov 2003, M. Irwin, R. Harin’Hala, 13

74 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 400

(00006333) ( CAS); 01 Nov 2004 – 14 Nov 2004, M. anterolateral projections (pl. 15B), slightly cari- Irwin, R. Harin’Hala, 13 (00006336) ( CAS); 07 Dec nate laterally; posterior pronotal lobe trans- 2004– 22 Dec 2004, M. Irwin, R. Harin’Hala, 13

versely striated; pronotal transverse furrow

(00006335) ( CAS); 22 Dec 2004 – 06 Jan 2005, M. Irwin, R. Harin’Hala, 13 (00006083) ( BMNH) ; 16 Jan 2005 – continuous; legs slender; tarsomeres I and II 27 Jan 2005, M. Irwin, R. Harin’Hala, 23 (00006331, combined subequal to III on mid and hind tarsi, 00006332) ( USNM) ; 23 Mar 2005 – 03 Apr 2005, M. foretarsi missing; wings spread out in specimens. Irwin, R. Harin’Hala, 23 (00006958, 00006959) ( UCR) ; ABDOMEN: Apex rounded ; dorsal laterotergite II 03 Apr 2005 – 17 Apr 2005, M. Irwin, R. Harin’Hala , 13 not expanded ; intersegmental sutures carinulate (00006127) (SU).

between sternites II – VI and laterally between VI

and VII; pygophore process subtriangular in lat-

Tanindrazanus bemaraha , new species eral view (pl. 20H, I); BPE as long as basal plate;

Plates 2, 6, 9; map 7 area of endosomal struts–DPS fusion elongate

DIAGNOSIS: Males are recognized among other subquadrate (pl. 21G). species in Tanindrazanus by the longer labial seg- ETYMOLOGY: The species epithet is a noun in ment II relative to III, anterior pronotal lobe with apposition and is named after the type locality, very small anterolateral protuberances, and ante- Tsingy de Bemaraha National Park, rior pronotal lobe slightly carinate laterally. This Madagascar. species is similar to T. andohahela and T. notatus , DISTRIBUTION: Tsingy de Bemaraha National but differs from both species by the presence of Park in the Mahajanga province (map 7). the anterolateral protuberances and laterally cari- HOLOTYPE: Male: Madagascar: Mahajanga: nate anterior pronotal lobe. Tanindrazanus bema- Parc National Tsingy de Bemaraha, 10.6 km ESE raha is further distinguished from T. notatus by 123° Antsalova, 19.70944 ° S 44.71806 ° E, 150 m, the lack of black markings on the hind femur and 16 Nov 2001 – 20 Nov 2001, Fisher et al. tibia and longer labial segment II. (00006473) (CAS).

DESCRIPTION: Medium body size (length: PARATYPE: Madagascar: Mahajanga: Parc National 13.93 mm, holotype; 13.07–13.93 mm). COLOR- Tsingy de Bemaraha , 10.6 km ESE 123° Antsalova ,

19.70944 ° S 44.71806 ° E, 150 m, 16 Nov 2001 –20 Nov

ATION: Orange-brown with dark brown scape

2001, Fisher et al., 13 (00005362) (CAS).

except basally, pedicel, and antennal segments III and IV, as well as V basally. Antennal segment V apically and segments VI and VII pale. Wing Tanindrazanus brunneus , new species membrane and distal wing veins brown. VESTI- Plates 2, 6, 9; map 8

TURE: As in generic description. STRUCTURE: DIAGNOSIS: Males recognized among other

HEAD: Ventrally with shallow anteromedial species in this genus by the very elongate subtridepression; clypeal apex dorsally elevated relative angular (almost cylindrical) head in lateral view, to labrum (pl. 13G, H); postclypeus with rela- postclypeal depression extending to interocular tively deep, broad medial longitudinal depression sulcus, broad postocular in dorsal view, synthlipto middle of interocular area; synthlipsis about sis about 2.5 width of an eye, eyes about one-fifth 1.5 times width of eye; interocular sulcus near of the head length, and small ocelli on a shalhind margin of eye (pl. 13B); postocular narrow lower ocellar tubercle. This species is similar to in dorsal view (pl. 13A); ocelli large, separated by T. joffrevillus , but is distinguished by the precedless than diameter of ocellus, located on distinct ing characters, labial segment II longer than III, median tubercle (pl. 13H); eye about one-fourth striated posterior pronotal lobe, and slender legs. of head length, not reaching dorsal and ventral DESCRIPTION: Medium body size (length: head surfaces; pedicel about one-third longer 10.31 mm, holotype). COLORATION: Orange than scape; labial segment III shorter than II. with pale thoracic pleura and sterna and brown

THORAX: Anterior pronotal lobe with very small pronotum and wing membrane. Scape, pedicel,

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antennal segments III and IV, as well as segment size, red and black coloration, flat ventral head V basally and segment VII apically, dark brown. surface, labial segment II shorter than III, later- Antennal segment V apically, segment VI, and ally carinate anterior pronotal lobe, slightly base of segment VII white. VESTITURE: As in incrassate forefemur, and intersegmental sutures generic description. STRUCTURE: HEAD: Elon- carinulate between sternites II and III and latergate; ventrally flat; clypeal apex not elevated rela- ally between III–V. This species is similar to T. tive to labrum (pl. 13C–F, I–K); postclypeus with marojejy , T. mahafaly , and T. irwini , but can be very shallow, narrow medial longitudinal depres- separated from the features mentioned, in parsion to interocular sulcus; synthlipsis about 2.5 ticular the coloration and sternal intersegmental times width of eye; interocular sulcus near hind carinulation pattern. margin of eye (pl. 13B); postocular broad in dor- DESCRIPTION: Large body size (length: 23.39 sal view (pl. 13B); ocelli relatively small, sepa- mm, holotype; 22.47–23.69 mm). COLORrated by less than diameter of ocellus, located on ATION: Red with black antenna (except pedicel shallow medial tubercle (pl. 13G); eyes about basally, apical half of segment V, and segment one-fifth of head length, not reaching dorsal and VI), posterior pronotal margin, scutellar proventral head surfaces; pedicel about one-fourth cesses apically, hemelytron except base of corium, longer than scape; labial segment III shorter than trochanters, femora basally, and posterolateral II. THORAX: Anterior pronotal lobe without dis- stripes on abdominal sternites III–VI. Hind tibia tinct anterolateral projections; posterior pronotal dark red and distal half of antennal segment V lobe transversely striated; pronotal transverse and segment VI white. VESTITURE: As in furrow divided by paramedian longitudinal generic description. STRUCTURE: HEAD: Venridges (pl. 15B); legs slender; tarsomeres I and II trally flat; clypeal apex dorsally elevated relative combined subequal to III; hemelytron surpassing to labrum (pl. 13G, H); postclypeus with relaabdominal apex. ABDOMEN: Apex rounded; dor- tively deep, broad medial longitudinal depression sal laterotergite II not expanded; intersegmental to middle of interocular area; synthlipsis about sutures carinulate between sternites II–VI and 1.5 times width of eye; interocular sulcus near laterally between VI and VII; pygophore process hind margin of eye (pl. 13B); postocular narrow very flattened in lateral view (pl. 20F); BPE as in dorsal view (pl. 13A); ocelli large, separated by long as basal plate; area of endosomal struts– less than diameter of ocellus, located on distinct DPS fusion ovate (pl. 21H). median tubercle (pl. 13H); eye about one-fourth ETYMOLOGY: The species epithet is an adjec- of head length, not reaching dorsal and ventral tive in the nominative case and is named for the head surfaces; pedicel about one-fourth longer color of the body. than scape; labial segment III longer than II. DISTRIBUTION: The only known specimen THORAX: Anterior pronotal lobe without distinct was collected from Marojejy Nature Reserve in anterolateral projections, laterally carinate; posthe Antsiranana province (map 8). terior pronotal lobe transversely striated; prono- HOLOTYPE: Male: Madagascar: Antsira- tal transverse furrow divided by paramedian nana: R.N. I. de Marojejy, 10.0 km NW Manan- longitudinal ridges (pl. 15B); forefemur slightly tenina, 14.43333 ° S 49.76167 ° E, 750 m, 15 Oct incrassate; tarsomeres I and II combined shorter 1996– 22 Oct 1996, E. Quinter and T. Nguyen than III on fore- and mid legs, subequal on hind (00078368) (AMNH). legs; hemelytron reaching or surpassing abdomi-

nal apex. ABDOMEN: Apex rounded or notched Tanindrazanus hannajagodae , new species medially; dorsal laterotergite II not expanded;

Plates 2, 6, 9; map 6 intersegmental sutures carinulate between ster- DIAGNOSIS: Males are recognized among nites II and III and laterally between III–V; pygoother Tanindrazanus species by the large body phore process subtriangular in lateral view (pl.

76 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 400

20H, I); BPE as long as basal plate; area of endo- erotergites except posterior half of VII, ventrolatsomal struts–DPS fusion circular. eral stripes on abdominal sternites III and IV, ETYMOLOGY: The species epithet is a noun in lateral margins of sternites II – VI, and anterolateral the genitive case and is named after the second area of VII, labial segment II basally, labial segauthor’s daughter, Hanna Jagoda Chłond. ment IV, prothorax, and dorsal half of mesopleu- DISTRIBUTION: Northern region of the ron orange. VESTITURE: As in generic description. Antsiranana province (map 6). STRUCTURE: HEAD: Ventrally with shallow DISCUSSION: The coloration is variable with anteromedial depression ; clypeal apex dorsally the hind femur dark red, the ventral interseg- elevated relative to labrum (pl. 13G, H); postclypmental sutures black, and the hemelytron light eus with shallow, broad medial longitudinal brown in some specimens. depression to middle of interocular area (pl. 13B); HOLOTYPE: Male : Madagascar: Antsiranana: synthlipsis about 1.5 times width of eye ; interocu- 7 km N of Joffreville , 12.45 ° S 49.23333 ° E, 426 m, lar sulcus near hind margin of eye (pl. 13B) ; post- 11 Feb 2007 – 18 Feb 2007, M. Irwin, R. Harin’Hala ocular narrow in dorsal view (pl. 13A) ; ocelli large, (00006261) ( CAS). separated by diameter of ocellus, located on dis- PARATYPES: Madagascar: Antsiranana: 7 km N of Jof- tinct median tubercle (pl. 13H) ; eye about onefreville, 12.45 ° S 49.23333 ° E, 426 m, 20 Jan 2007 –27 Jan third of head length, not reaching dorsal and 2007, M. Irwin, R. Harin’Hala, 13 (00006263) ( SU) ; 11

ventral head surfaces; pedicel about one-fourth Feb 2007– 18 Feb 2007, M. Irwin, R. Harin’Hala, 13 (00006262) ( UCR). Réserve Spéciale d’Ambre, 3.5 km 235° longer than scape ; labial segment III subequal to SW Sakaramy, 12.46888 ° S 49.24222 ° E, 325 m, 26 Jan II. THORAX: Anterior pronotal lobe without dis- 2001– 31 Jan 2001, Fisher et al., 13 (00006117) ( CAS). tinct anterolateral projections ; posterior pronotal

lobe transversely striated; pronotal transverse fur-

Tanindrazanus harinhali , new species row divided by paramedian longitudinal ridges (pl.

Plates 2, 6, 9 ; map 6 15B); forefemur slightly incrassate; tarsomeres I DIAGNOSIS: Males are recognized among and II combined subequal to III ; hemelytron surother species in this genus by a combination of passing abdominal apex. ABDOMEN: Apex the following characters: the distinct tricolor rounded ; dorsal laterotergite II not expanded; pattern (dark brown, orange, and pale yellow), intersegmental sutures carinulate between sternites synthlipsis about 1.5 times the width of the eye, II–VI and laterally between VI and VII; pygophore eye not reaching the dorsal and ventral head process subtriangular in lateral view (pl. 20H, I); margins, ocelli separated by diameter of ocellus, BPE shorter than basal plate; area of endosomal and the pronotal transverse furrow not continu- struts–DPS fusion elongate ovoid. ous. This species is similar to T. amboasaricus , ETYMOLOGY: The species epithet is a noun in but differs by the smaller body size ; orangish the genitive case and is named after Rasolondalao head, pronotum, and scutellum; greater syn- Harin’Hala Hasinjaka , who helped collect a majorthlipsis width ; and intersegmental sutures cari- ity of the specimens examined for this species. nulate between sternites II – VI and laterally DISTRIBUTION: Fianarantsoa and Toliara between VI and VII. provinces (map 6). DESCRIPTION: Medium body size (length: 10.99 DISCUSSION: The yellow coloration on the mm, holotype; 10.39–11.89 mm). COLORATION: head, legs, and stripes on the abdominal sternites Dark brown with pale yellow mandibular and may be reduced in size or absent, and the clypmaxillary plates, postclypeus, antennifer dorsally, eus, posteroventral head surface, and dorsoscape basally, scutellum medially, corium basally, medial neck surface may be yellow. Approximately coxae, trochanters, fore- and mid femora except half of the specimens examined have the DFLA dorsally, hind femur except apically, foretibia, mid subdivided into two pseudosegments instead of tibia except apically, hind tibia medially, tarsi, lat- three among and within individuals .

2016 FORTHMAN ET AL.: MILLIPEDE ASSASSIN BUG FAUNA 77

HOLOTYPE: Male: Madagascar: Toliara: Beza 2004, M. Irwin, F. Parker, R. Harin’Hala , 23 (00006544, Mahafaly Reserve , Parcelle I near research sta- 00006545) ( CAS); 06 Jun 2004 – 20 Jun 2004, M. Irwin, tion, 23.6865 ° S 44.591 ° E, 165 m, 28 Nov 2001 –04 F. Parker, R. Harin’Hala, 23 (00006533, 00006553)

( USNM); 22 Aug 2004 – 01 Sep 2004, M. Irwin, F. Dec 2001, R. Harin’Hala (00006418) ( CAS). Parker, R. Harin’Hala , 13 (00006530) ( UCR). Berenty PARATYPES: Madagascar: Fianarantsoa: 50 km S of Special Reserve, 8 km NW Amboasary, 25.021 ° S Farafangana , Mahabo Mananivo , Ampitavananima For- 46.3055 ° E, 35 m, 21 Dec 2003 – 01 Jan 2004, M. Irwin, est, 23.12983 ° S 47.717 ° E, 34 m, 03 Mar 2007 –10 Mar F. Parker, R. Harin’Hala, 13 (00006346) ( CAS) ; 02 Jan 2007, M. Irwin, F. Parker, R. Harin’Hala, 13 (00044892) 2004– 13 Jan 2004, M. Irwin, F. Parker, R. Harin’Hala, ( CAS). Parc National Ranomafana , radio tower at forest 13 (00006359) ( UCR) ; 24 Mar 2004 – 04 Apr 2004, M. edge, 21.251 ° S 47.40716 ° E, 1130 m, 16 Oct 2001 –08 Irwin, F. Parker, R. Harin’Hala , 23 (00006549, Nov 2001, M. Irwin, R. Harin’Hala, 13 (00006425) 00006550) ( CAS) ; 04 Apr 2004 – 15 Apr 2004, M. Irwin, ( CAS) ; 09 Aug 2005 – 25 Aug 2005, M. Irwin, R. F. Parker, R. Harin’Hala , 13 (00006531) ( SU) ; 23 Jun Harin’Hala, 13 (00006537) ( UCR). Radio tower 22 km 2004– 04 Jul 2004, M. Irwin, F. Parker, R. Harin’Hala , SW of Ilakaka, near Fianarantsoa / Toliara border, 23 (00006350, 00006351) ( AMNH) ; 04 Jul 2004 –18 Jul 22.77917 ° S 45.025 ° E, 1100 m, 06 Mar 2002 – 11 Mar 2004, M. Irwin, F. Parker, R. Harin’Hala, 13 (00006356) 2002, M. Irwin, R. Harin’Hala, 13 (00006340) ( CAS). ( CAS). Beza Mahafaly Reserve , Parcelle II near Bel- Toliara: Andohahela Natl Park, Tsimelahy, Parcelle II, levue, 23.68983 ° S 44.5755 ° E, 180 m, 10 Nov 2001 –21 24.93683 ° S 46.62666 ° E, 180 m, 27 Dec 2002 –06 Jan Nov 2001, M. Irwin, R. Harin’Hala, 13 (00006424) 2003, M. Irwin, F. Parker, R. Harin’Hala, 13 (00006391) ( MNHN) ; 18 Dec 2001 – 25 Dec 2001, M. Irwin, R. ( CAS) ; 06 Jan 2003 – 16 Jan 2003, R. Harin’Hala, 13 Harin’Hala, 13 (00006426) ( SU) ; 16 Jan 2002 –18 Jan (00006376) ( CAS), 13 (00006377) ( USNM) ; 08 Mar 2002, M. Irwin, R. Harin’Hala, 13 (00006385) ( CAS) ; 2003– 18 Mar 2003, M. Irwin, F. Parker, R. Harin’Hala , 08 Feb 2002 – 15 Feb 2002, M. Irwin, R. Harin’Hala, 13 23 (00006337, 00006344) ( SU) ; 28 Mar 2003 –08 Apr (00006540) ( UCR). Beza Mahafaly Reserve, Parcelle I 2003, M. Irwin, F. Parker, R. Harin’Hala , 13 (00006552) near research station, 23.6865 ° S 44.591 ° E, 165 m, 15 ( CAS) ; 22 Jun 2003 – 29 Jun 2003, M. Irwin, F. Parker, Oct 2001– 10 Nov 2001, M.E. Irwin, F.D. Parker, R. R. Harin’Hala , 13 (00006375) ( AMNH) ; 17 Aug 2003 – Harin’Hala, 13 (00006387) ( CAS) ; 10 Nov 2001 –21 24 Aug 2003, M. Irwin, F. Parker, R. Harin’Hala , 13 Nov 2001, R. Harin’Hala, 13 (00006557) ( UCR) ; 21 (00006554) ( UCR); 09 Nov 2003 – 20 Nov 2003, M. Nov 2001– 28 Nov 2001, R. Harin’Hala, 23 (00006341, Irwin, F. Parker, R. Harin’Hala, 13 (00006551) ( CAS). 00006392) ( CAS) ; 28 Nov 2001 – 04 Dec 2001, R. Berenty Special Reserve, 8 km NW Amboasary, Harin’Hala , 13 (00006417) ( CAS) ; 04 Dec 2001 –11 25.00666 ° S 46.30333 ° E, 85 m, 25 Oct 2002 –26 Oct Dec 2001, R. Harin’Hala, 23 (00006345, 00006415) 2002, M. Irwin, F. Parker, R. Harin’Hala, 13 (00006340) ( SU) ; 11 Dec 2001 – 18 Dec 2001, R. Harin’Hala, 23 ( CAS) ; 26 Oct 2002 – 02 Nov 2002, M. Irwin, F. Parker, (00006132, 00006379) ( AMNH) ; 18 Dec 2001 –25 Dec R. Harin’Hala, 13 (00006349) ( CAS), 13 (00006357) 2001, R. Harin’Hala , 13 (00006388) ( CAS) ; 25 Dec ( BMNH) ; 30 Nov 2002 – 07 Dec 2002, M. Irwin, F. 2001– 02 Jan 2002, R. Harin’Hala, 13 (00006548) Parker, R. Harin’Hala, 13 (00005359) ( SU), 33 ( BMNH) ; 18 Jan 2002 – 25 Jan 2002, M.E. Irwin, F.D. (00006353–00006355) ( CAS) ; 26 Jan 2003 – 05 Feb 2003, Parker, R. Harin’Hala, 13 (00045278) ( BMNH) ; 25 Jan M. Irwin, F. Parker, R. Harin’Hala, 13 (00006529) 2002– 01 Feb 2002, R. Harin’Hala , 23 (00006389, ( UCR); 24 Mar 2003 – 03 Apr 2003, M. Irwin, F. Parker, 00006390) ( USNM) ; 08 Feb 2002 – 15 Feb 2002, R. R. Harin’Hala , 23 (00006532, 00045392) ( AMNH) ; 03 Harin’Hala, 23 (00006342, 00006402) ( CAS) ; 14 Mar May 2003– 14 May 2003, M. Irwin, F. Parker, R. 2002– 22 Mar 2002, R. Harin’Hala , 13 (00006420) Harin’Hala, 13 (00006374) ( CAS) ; 22 Nov 2003 –30 ( CAS) ; 29 Mar 2002 – 10 Apr 2002, R. Harin’Hala , 13 Nov 2003, M. Irwin, F. Parker, R. Harin’Hala, 13 (00006555) ( CAS) ; 29 Apr 2002 – 19 May 2002, R. (00006534) ( BMNH) ; 07 Dec 2003 – 14 Dec 2003, M. Harin’Hala, 13 (00006386) ( CAS) ; 19 May 2002 –08 Irwin, F. Parker, R. Harin’Hala, 23 (00006541, Jun 2002, R. Harin’Hala, 13 (00006401) ( CAS); 08 Jun 00006542) ( USNM) ; 02 Jan 2004 – 13 Jan 2004, M. 2002– 18 Jun 2002, R. Harin’Hala, 23 (00006421, Irwin, F. Parker, R. Harin’Hala, 13 (00006547) ( CAS); 00006423) ( BMNH) ; 18 Jun 2002 – 28 Jun 2002, R. 23 Jan 2004 – 04 Feb 2004, M. Irwin, F. Parker, R. Harin’Hala, 13 (00006393) ( CAS) ; 07 Jul 2002 –18 Jul Harin’Hala, 23 (00006543, 00006546) ( MNHN) ; 04 2002, R. Harin’Hala, 13 (00006412) ( CAS) ; 28 Jul Feb 2004– 15 Feb 2004, M. Irwin, F. Parker, R. 2002– 09 Aug 2002, R. Harin’Hala , 13 (00006338) Harin’Hala, 23 (00006528, 00006536) ( AMNH) ; 15 ( CAS); 09 Aug 2002 – 16 Aug 2002, R. Harin’Hala , 13 Feb 2004 – 02 Mar 2004, M. Irwin, F. Parker, R. (00006427) ( CAS) ; 16 Aug 2002 – 28 Aug 2002, R. Harin’Hala, 13 (00006358) ( SU) ; 27 May 2004 –06 Jun Harin’Hala, 23 (00006403, 00006404) ( USNM) ; 09 Sep

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2002– 20 Sep 2002, R. Harin’Hala, 13 (00006414) head dorsally, scape, labium except segment II ( CAS); 20 Sep 2002 – 05 Oct 2002, R. Harin’Hala, 13 basally, scutellum medially, hemelytron except (00006422) ( UCR); 20 Oct 2002 – 28 Oct 2002, R. posteromedial spot, femora apically, tibiae, tarsi, Harin’Hala, 13 (00006380) ( CAS); 28 Oct 2002 –10

laterotergites II – V, anterior half of laterotergite

Nov 2002, M.E. Irwin, F.D. Parker, R. Harin’Hala, 23 (00006409, 00006410) ( BMNH); 02 Dec 2002 –12 Dec VI, sternites III – V, and anterior half of sternite 2002, R. Harin’Hala, 23 (00006347, 00006348) ( CAS); VI except posterolateral stripes. Distal half of 12 Dec 2002 – 17 Dec 2002, R. Harin’Hala, 13 antennal segment V and segments VI and VII (00006378) ( CAS); 17 Dec 2002 – 20 Dec 2002, R. white. VESTITURE: As in generic description. Harin’Hala, 13 (00006419) ( CAS); 20 Dec 2002 –24

STRUCTURE: HEAD: Ventrally flat; clypeal apex

Dec 2002, R. Harin’Hala, 13 (00006095) ( CAS) ; 24 Dec 2002 – 02 Jan 2003, R. Harin’Hala, 13 (00006413) dorsally elevated relative to labrum (pl. 13G, H); ( AMNH) ; 09 Jan 2003 – 23 Jan 2003, R. Harin’Hala, 43 postclypeus with relatively deep, broad medial (00006405–00006408) ( CAS) ; 23 Jan 2003 – 04 Feb 2003, longitudinal depression to middle of interocular R. Harin’Hala, 53 (00006395–00006399) ( MNHN) ; 04 area; synthlipsis about 1.5 times width of eye; Feb 2003– 16 Feb 2003, R. Harin’Hala , 13 (00006400) interocular sulcus near hind margin of eye (pl. (SU); 16 Feb 2003 – 02 Mar 2003, R. Harin’Hala, 13

13B); postocular narrow in dorsal view (pl. 13A);

(00006556) ( BMNH); 13 Mar 2003 – 23 Mar 2003, R. Harin’Hala, 23 (00006381, 00006382) ( USNM); 23 ocelli large, separated by less than diameter of Mar 2003– 02 Apr 2003, R. Harin’Hala, 13 (00006339) ocellus, located on distinct median tubercle (pl. ( CAS); 02 Apr 2003 – 10 Apr 2003, R. Harin’Hala, 13 13H); eye about one-fourth of head length, not (00006352) ( CAS); 29 Apr 2003 – 07 May 2003, R. reaching dorsal and ventral head surfaces; pedi- Harin’Hala, 13 (00006416) ( UCR). Parc National

cel about one-fourth longer than scape; labial

d’Andohahela, Ihazofotsy Parcelle III, 24.83083 ° S 46.53616 ° E, 80 m, 06 May 2003 – 13 May 2003, M. segment III longer than II. THORAX (pl. 15F): Irwin, F. Parker, R. Harin’Hala , 13 (00006343) (SU) ; 29 Anterior pronotal lobe without distinct antero- Jun 2003– 06 Jul 2003, M. Irwin, F. Parker, R. Harin’Hala , lateral projections, laterally carinate ; posterior 13 (00044843) ( UCR); 03 Aug 2003 – 13 Aug 2003, M. pronotal lobe transversely striated; pronotal Irwin, F. Parker, R. Harin’Hala, 23 (00044809, transverse furrow divided by paramedian longi- 00044837) ( CAS). Unknown : 53 (00006383, 00006384,

tudinal ridges (pl. 15B); legs slender; tarsomeres

00006535, 00006538, 00006539) ( CAS).

I and II combined subequal to III; hemelytron

reaching or surpassing abdominal apex. ABDO-

Tanindrazanus irwini , new species MEN: Apex rounded; dorsal laterotergite II not

Plates 2, 6, 9, 15F; map 7 expanded; intersegmental sutures carinulate

DIAGNOSIS: Males are recognized among between sternite II–IV and laterally between IV– other species in Tanindrazanus by the large body VI; pygophore process subtriangular in lateral size, black coloration with orange markings, flat view (pl. 20H, I); BPE as long as basal plate; area ventral head surface, labial segment II shorter of endosomal struts–DPS fusion ovate (pl. 21H). than III, laterally carinate anterior pronotal lobe, ETYMOLOGY: The species epithet is a noun in slender legs, and intersegmental sutures carinu- the genitive case and is named after Mike Irwin, late between sternite II–IV and laterally between who helped collect a majority of the specimens IV–VI. This species is similar to T. varicolor , but examined for this species. differs by the larger body size, flat ventral head DISTRIBUTION: Marojejy Nature Reserve in surface, labial segment II shorter than III, later- the Antsiranana province and near the Andaally carinate anterior pronotal lobe, and interseg- sibe-Mantadia National Park in the Toamasina mental sutures carinulate between sternite II–VI province (map 7). and laterally between VI and VII. DISCUSSION: Coloration is variable with the

DESCRIPTION: Large body size (length: 22.76 orange areas reduced or absent on the dorsal mm, holotype; 18.95–22.76 mm). COLOR- head surface, labium, scutellum, femora, hind

ATION: Dark brown-black with bright orange tibia, and abdomen in some specimens.

2016 FORTHMAN ET AL.: MILLIPEDE ASSASSIN BUG FAUNA 79

HOLOTYPE: Male : Madagascar: Antsiranana: distinguished by dark brown corium, ventrally

Marojejy National Park, 5 km W Manantenina flat head surface, clypeal apex not elevated rela-

village, Camp Mantella, 14.43816 ° S 49.774 ° E, tive to the labrum, larger synthlipsis width, sub-

490 m, 30 May 2005 – 11 Jun 2005, M. Irwin, R. equal lengths of labial segments II and III,

Harin’Hala (00045088) ( CAS). smooth posterior pronotal lobe, divided pronotal

PARATYPES: Madagascar: Antsiranana: Marojejy transverse suture, and intersegmental sutures

National Park, 5 km W Manantenina village, Camp carinulate between sternites II – VI.

Mantella, 14.43816 ° S 49.774 ° E, 490 m, 09 Dec 2004 –15

DESCRIPTION: Medium body size (length:

Dec 2004, M. Irwin, R. Harin’Hala, 23 (00045220,

00045375) ( CAS); 25 Dec 2004 – 30 Dec 2004, M. Irwin, 11.19 mm, holotype). COLORATION: Dull

R. Harin’Hala, 13 (00006224) (SU), 13 (00006225) orange with thoracic sterna pale and hemelytron,

(UCR); 05 Jan 2005 – 10 Jan 2005, M. Irwin, R. except corium basally, brown. Antennal segments

Harin’Hala, 23 (00006096, 00006097) (CAS); 10 Jan V and VI white. VESTITURE: As in generic

2005– 15 Jan 2005, M. Irwin, R. Harin’Hala, 13 description. STRUCTURE: HEAD: Ventrally flat;

(00006235) (UCR); 15 Jan 2005 – 26 Jan 2005, M. Irwin,

R. Harin’Hala, 13 (00006135) (AMNH); 04 Feb 2005 – clypeal apex not elevated relative to labrum (pl.

11 Feb 2005, M. Irwin, R. Harin’Hala, 33 (00006233, 13C–F, I–K); postclypeus with relatively shallow,

00006234, 00006239) (CAS); 11 Feb 2005 – 18 Feb 2005, broad medial longitudinal depression to middle of

M. Irwin, R. Harin’Hala, 33 (00006217, 00006241, interocular area (pl. 13B); synthlipsis about two

00006242) (AMNH); 18 Feb 2005 – 25 Feb 2005, M. times width of eye; interocular sulcus posterior to

Irwin, R. Harin’Hala, 33 (00045603, 00045612, hind margin of eye; postocular narrow in dorsal

00045712) (BMNH); 25 Feb 2005 – 04 Mar 2005, M.

Irwin, R. Harin’Hala, 53 (00005366, 00006226– view (pl. 13A); ocelli large, separated by less than

00006229) (CAS); 11 Mar 2005 – 18 Mar 2005, M. Irwin, diameter of ocellus, located on distinct median

R. Harin’Hala, 13 (00006232) (UCR); 04 Apr 2005 –16 tubercle (pl. 13H); eyes about one-fourth of head

Apr 2005, M. Irwin, R. Harin’Hala, 23 (00006499, length, not reaching dorsal and ventral head sur-

00045339) (SU); 16 Apr 2005 – 28 Apr 2005, M. Irwin, faces; pedicel about one-fourth longer than scape;

R. Harin’Hala, 63 (00007087, 00045004, 00045048,

00045072, 00045405, 00045475) (CAS); 28 Apr 2005 – labial segment III subequal II. THORAX: Anterior

07 May 2005, M. Irwin, R. Harin’Hala, 13 (00006221) pronotal lobe without distinct anterolateral pro-

(UCR); 07 May 2005 – 18 May 2005, M. Irwin, R. jections; posterior pronotal lobe smooth; pronotal

Harin’Hala, 33 (00006134, 00006237, 00006238) transverse furrow divided by paramedian longitu-

(USNM); 18 May 2005 – 30 May 2005, M. Irwin, R. dinal ridges (pl. 15B); forefemur slightly incras-

Harin’Hala, 13 (00006216) (BMNH), 13 (00006243) sate; tarsomeres I and II combined subequal to III;

(USNM); 30 May 2005 – 11 Jun 2005, M. Irwin, R.

Harin’Hala, 43 (00007211, 00044957, 00044982, hemelytron surpassing abdominal apex. ABDO-

00045462) (CAS); 28 Jun 2005 – 13 Jul 2005, M. Irwin, MEN: Apex rounded; dorsal laterotergite II not

R. Harin’Hala, 13 (00006218) (SU), 13 (00006222) expanded; intersegmental sutures carinulate

(UCR), 43 (00006223, 00006230, 00006231, 00006236) between sternites II–VI; pygophore process very

(MNHN), 43 (00006240, 00006452–00006454) (CAS). flattened in lateral view (pl. 20F); BPE as long as

Toamasina: Lakato env. Near Andasibe-Mantadia basal plate; area of endosomal struts–DPS fusion

National Park, 19.18651 ° S 48.43856 ° E, 635 m, 09 Jan

2007– 10 Jan 2007, Z. Mráček, 13 (MMBC). elongate subquadrate (pl. 21G).

ETYMOLOGY: The species epithet is a noun in

Tanindrazanus joffrevillus , new species apposition and is named after the type locality, Plates 2, 6, 9; map 6 Joffreville , Madagascar .

DIAGNOSIS: Males recognized among other DISTRIBUTION: Only one of the two known

species in this genus by the dull orange color- specimens has locality data. It is known from a

ation, ventrally flat head surface, clypeal apex not locality north of Joffreville in the Antsiranana

elevated relative to the labrum, smooth posterior province (map 6).

pronotal lobe, and slightly incrassate forefemur. HOLOTYPE: Male : Madagascar: Antsiranana:

This species is similar to T. andohahela , but is 7 km N of Joffreville, 12.33333 ° S 49.25 ° E, 360 m,

80 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 400

20 Mar 2001 – 07 Apr 2001, M. Irwin, R. 15D): Anterior pronotal lobe with very small Harin’Hala (00007256) ( CAS). anterolateral projections (pl. 15B) ; posterior pro- PARATYPES: Unknown, 13 (00044998) ( CAS). notal lobe transversely striated ; pronotal trans-

verse furrow continuous; legs slender; tarsomeres Tanindrazanus kathrynae , new species I and II combined subequal to III; hemelytron Plates 2, 6, 9, 13G, 15D, 19F, 20G; map 7 reaching or surpassing abdominal apex. ABDO- DIAGNOSIS: Males recognized among other MEN: Apex rounded; dorsal laterotergite II not Tanindrazanus species by the red and black col- expanded; intersegmental sutures carinulate oration, elevated clypeal apex relative to the between sternites II–V and laterally between V labrum, broad postocular in dorsal view, and and VI; pygophore process subtriangular in latsynthlipsis about two times the width of the eye. eral view (pl. 20G); BPE longer than basal plate; This species is very similar to T. anjozorobeus , area of endosomal struts–DPS fusion ovate (pl. but is distinguished by the smaller body size, 21H). smaller eyes that do not surpass the dorsal and ETYMOLOGY: The species epithet is a noun ventral margins of the head in lateral view, larger in the genitive case and is named in memowidth of the synthlipsis, and the very small riam of the senior author’s partner’s mother, anterolateral projections on the pronotum. Kathryn Carroll. DESCRIPTION: Medium body size (length: DISTRIBUTION: Miandritsara Forest in the 12.67 mm, holotype; 11.62–12.67 mm). COLOR- Fianarantsoa province (map 7). ATION: Red with pale orange-brown mandibu- DISCUSSION: In some specimens, the head lar and maxillary plates, clypeus, anterolateral and anterior pronotal lobe have reduced red col- and dorsal margin of eye, labium, anterior pro- oration or are entirely black and the median lonnotal lobe except lateral margins, and corium gitudinal suture on the pronotum and the medial except distally. Remainder of head except two area of abdominal sternite VII are black. paramedian dorsal stripes posterior to ocelli, lat- HOLOTYPE: Male : Madagascar: Fianaranteral margins of anterior pronotal lobe, anterior soa: Miandritsara Forest , 40 km S Ambositra, part of propleura, meso- and metapleura, scutel- 20.79266 ° S 47.17566 ° E, 825 m, 13 Nov 2006 – lum, corium distally, forewing membrane, fem- 22 Nov 2006, M. Irwin, R. Harin’Hala ora and apical one-third to one-half of fore- and (00006103) ( CAS). mid tibiae, hind tibiae except small apical part, PARATYPES: Madagascar: Fianarantsoa: Andringiventrolateral spot on abdominal sternites II – VI, tra Est, Ambalamarovandana, 22.22595 ° S 46.93418 ° E, and pygophore dark brown to black. VESTI- 1500–1600m, 15 Jan 1971 – 25 Jan 1971, 13 ( MNHN) .

Miandritsara Forest, 40 km S Ambositra, 20.79266 ° S TURE: As in generic description. STRUCTURE:

47.17566 ° E, 825 m, 23 Oct 2004 – 03 Nov 2004, M. HEAD (pl. 13G): Ventrally with shallow antero- Irwin, R. Harin’Hala , 13 (00045053) ( UCR) ; 05 Jan medial depression; clypeal apex dorsally elevated 2005– 18 Jan 2005, M. Irwin, R. Harin’Hala , 23 relative to labrum ; postclypeus with relatively (00044858, 00045506) ( CAS), 13 (00045608) (SU) ; 09 deep, broad medial longitudinal depression to Feb 2005– 22 Feb 2005, M. Irwin, R. Harin’Hala , 13 middle of interocular area ; synthlipsis about two (00045383) (AMNH); 22 Feb 2005 – 05 Mar 2005, M.

Irwin, R. Harin’Hala, 23 (00044863, 00045360) (CAS), times width of eye; interocular sulcus posterior 13 (00045626) (USNM); 13 Apr 2005 – 27 Apr 2005, M. to hind margin of eye; postocular broad in dorsal Irwin, R. Harin’Hala, 23 (00007062, 00044984) (CAS), view (pl. 13B); ocelli large, separated by less than 13 (00045549) (MNHN); 05 Jan 2006 – 12 Jan 2006, M. diameter of ocellus, located on slightly elevated Irwin, R. Harin’Hala, 23 (00044818, 00045566) median tubercle; eye about one-fourth of head (BMNH); 16 Feb 2006 – 27 Feb 2006, M. Irwin, R.

Harin’Hala, 13 (00044832) (BMNH); 27 Feb 2006 –08 length, not reaching dorsal and ventral head sur-

Mar 2006, M. Irwin, R. Harin’Hala, 13 (00045632) faces ; pedicel about two-fifths longer than scape; (SU); 14 Apr 2006 – 02 May 2006, M. Irwin, R. labial segment III subequal to II. THORAX (pl. Harin’Hala, 23 (00006094, 00006298) ( CAS), 13

2016 FORTHMAN ET AL.: MILLIPEDE ASSASSIN BUG FAUNA 81

(00006312) (MNHN); 13 Nov 2006 – 22 Nov 2006, M. interocular area (pl. 13B); synthlipsis about 1.5 Irwin, R. Harin’Hala, 23 (00006293, 00006299) times width of eye; interocular sulcus near hind (AMNH); 01 Dec 2006 – 10 Dec 2006, M. Irwin, R. margin of eye (pl. 13B); postocular narrow in

Harin’Hala, 13 (00006302) (UCR); 10 Dec 2006 – 19 Dec 2006, M. Irwin, R. Harin’Hala, 23 (00006313, dorsal view (pl. 13A); ocelli large, separated by 00006314) (CAS); 19 Dec 2006 – 28 Dec 2006, M. Irwin, less than diameter of ocellus, located on dis- R. Harin’Hala, 43 (00007053, 00007202, 00045201, tinct median tubercle (pl. 13H); eye about one- 00045365) (CAS); 15 Jan 2007 – 24 Jan 2007, M. Irwin, third of head length, not reaching dorsal and R. Harin’Hala, 23 (00045083, 00045482) (CAS); 24 Jan ventral head surfaces; pedicel about one-fourth 2007– 31 Jan 2007, M. Irwin, R. Harin’Hala, 13 longer than scape; labial segment III longer

(00006294) (BMNH); 31 Jan 2007 – 11 Feb 2007, M. Irwin, R. Harin’Hala , 23 (00006315, 00006316) (SU) ; than II. THORAX: Anterior pronotal lobe with- 11 Feb 2007 – 20 Feb 2007, M. Irwin, R. Harin’Hala , 13 out distinct anterolateral projections ; posterior (00006301) (MNHN), 33 (00006309–00006311) pronotal lobe transversely striated; pronotal (CAS); 20 Feb 2007 – 27 Feb 2007, M. Irwin, R. transverse furrow divided by paramedian lon- Harin’Hala, 13 (00006297) ( USNM) ; 08 Mar 2007 –17 gitudinal ridges (pl. 15B); forefemur slightly Mar 2007, M. Irwin, R. Harin’Hala, 43 (00006303– 00006306) ( CAS) ; 17 Mar 2007 – 26 Mar 2007, M. Irwin , incrassate ; tarsomeres I and II combined sub- R. Harin’Hala , 13 (00006300) ( AMNH) ; 26 Mar 2007 – equal to III on fore- and mid legs, longer on 05 Apr 2007, M. Irwin, R. Harin’Hala , 23 (00006307, hind legs; wings spread in specimens. ABDO- 00006308 ) ( USNM) ; 05 Apr 2007 – 14 Apr 2007, M. MEN: Apex rounded ; dorsal laterotergite II not Irwin, R. Harin’Hala , 23 (00006295, 00006296) ( UCR). expanded ; intersegmental sutures carinulate between sternites II and III; pygophore process

Tanindrazanus mahafaly , new species subtriangular in lateral view (pl. 20H, I); BPE Plates 3, 6, 9; map 7 as long as basal plate; area of endosomal struts–

DIAGNOSIS: Males are recognized among other DPS fusion circular. species in this genus by a combination of the fol- ETYMOLOGY: The species epithet is a noun in lowing characters: the large body size, distinct apposition and is named after the type locality, dark orange-brown to red-orange coloration, Beza Mahafaly Reserve, Madagascar. labial segment II shorter than III, slightly incras- DISTRIBUTION: Beza Mahafaly Reserve in the sate forefemur, and only the intersegmental suture Toliara province (map 7). between sternites II and III carinulate. This spe- DISCUSSION: In some specimens, the head, cies is similar to T. irwini , from which it differs by scutellum, fore- and mid legs, and abdomen are the dark orange-red color pattern, anteromedially bright red-orange, while the remaining strucdepressed ventral head surface, and carinulation tures are dull orange. pattern on the sternal intersegmental sutures.

HOLOTYPE: Male: Madagascar: Toliara: Beza

DESCRIPTION: Large body size (length: 19.57 Mahafaly Reserve , Parcelle I near research stamm, holotype; 18.03–19.92 mm). COLOR- tion, 23.6865 ° S 44.591 ° E, 165 m, 15 Oct 2001 –10

ATION: Dark orange with apical half of pedicel Nov 2001, M. Irwin, F. Parker, R. Harin’Hala and basiflagellomere, anterior pronotal lobe, (00045290) (CAS).

metapleuron, and abdomen, except lateroter- PARATYPES: Madagascar: Toliara: Beza Mahafaly gites, dark reddish brown. Hemelytron bright Reserve , Parcelle II near Bellevue , 23.68983 ° S 44.5755 ° E, orange and antennal segments VI and VII pale 180 m, 21 Nov 2001 – 28 Nov 2001, R. Harin’Hala, 13 orange to white. VESTITURE: As in generic (00006244) ( CAS). Beza Mahafaly Reserve , Parcelle I description. STRUCTURE: HEAD: Ventrally near research station, 23.6865 ° S 44.591 ° E, 165 m, 04 Dec 2001 – 11 Dec 2001, R. Harin’Hala, 13 (00006245) ( CAS) ;

with shallow anteromedial depression; clypeal 08 Feb 2002 – 15 Feb 2002, R. Harin’Hala, 13 (00006133)

apex dorsally elevated relative to labrum (pl. ( CAS); 22 Feb 2002 – 01 Mar 2002, R. Harin’Hala, 13 13G, H) ; postclypeus with shallow, broad (00006220) (SU); 18 Jun 2002 – 28 Jun 2002, R. medial longitudinal depression to middle of Harin’Hala, 13 (00006219) ( UCR) .

82 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 400