Rodrigues, Bruno V. B., Cizauskas, Igor & Rheims, Cristina A., 2018, Description of Paracymbiomma gen. nov., a new genus of prodidomid spiders from the Neotropical region (Araneae: Prodidomidae) including a new troglobite species, Zootaxa 4514 (3), pp. 301-331: 303-304

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gen. nov.

Paracymbiomma  gen. nov.

Etymology. The generic name is a contraction of paracymbium and Lygromma  , since the paracymbium is one of the diagnostic structures of the genus. Gender is neuter.

Type species. Paracymbiomma angelim  (original designation).

Diagnosis. Species of Paracymbiomma  gen.nov. are distinguished from those of other Prodidomidae  genera by the combination of the following characters: male palp with paracymbium and RTA single ( Figs 45View FIGURES 43–48, 54View FIGURES 53–56, 61View FIGURES 61–64), embolus long and filiform with curved tip ( Figs 43, 47View FIGURES 43–48, 49View FIGURES 49–52, 61View FIGURES 61–64), conductor sclerotized ( Figs 53, 55View FIGURES 53–56, 57) and median apophysis membranous with sclerotized tip ( Figs 53View FIGURES 53–56, 49View FIGURES 49–52, 61View FIGURES 61–64, 69–74View FIGURES 69–74); female epigyne with posteromedian atrium and sclerotized margins ( Figs. 51View FIGURES 49–52, 59, 63View FIGURES 61–64, 67View FIGURES 65–68), vulva with copulatory ducts long, strongly convoluted laterally with proximal part (closest to copulatory openings) translucent ( Figs 52View FIGURES 49–52, 68View FIGURES 65–68); secondary spermathecae elongated ( Figs 60, 64View FIGURES 61–64).

Description. Total length males 1.86–3.9 and females 2.25–4.15. Carapace and legs pale yellow, and abdomen weakly pigmented. Carapace longer than wide, narrower at cephalic region ( Fig. 7View FIGURES 7–12). Fovea present as a narrow dark longitudinal line on posterior half of carapace ( Figs 1–7View FIGURES 1–6View FIGURES 7–12). Eyes variable in number and size ( Figs 13–18View FIGURES 13–18). Chilum present, entire ( Fig. 8View FIGURES 7–12). Chelicerae relatively long (0.27–0.9) with a convex boss ( Fig. 8View FIGURES 7–12); promargin with 1–4 teeth, retromargin with 3–6 teeth; one large promarginal escort setae at base of fang ( Fig. 31View FIGURES 31–36); comb of whisker setae at cheliceral promargin near the base of fang on all females collected in caves ( Fig. 20View FIGURES 19–24). Endites slightly convergent anteriorly ( Fig. 9View FIGURES 7–12), with dense scopulae on internal margin; serrula present in single row ( Fig. 21View FIGURES 19–24); labium longer than wide ( Fig. 9View FIGURES 7–12). Sternum shield shaped, anteriorly straight, longer than wide, rebordered anteriorly and laterally; posterior region strongly protruding between coxae IV with numerous long and erect setae ( Figs 10View FIGURES 7–12, 22View FIGURES 19–24, 33View FIGURES 31–36); precoxal triangles present ( Fig. 10View FIGURES 7–12). Pedicel with ventral sclerite triangular. Leg formula 4123. Spination variable. All tarsi scopulate ( Figs 23View FIGURES 19–24, 25View FIGURES 25–30). Trichobothria present on dorsal tibia, metatarsi and tarsi, arranged in two rows on tarsi ( Fig. 25View FIGURES 25–30) and one on metatarsi ( Fig. 34View FIGURES 31–36); bothrium with 5–9 deep ridges ( Fig. 26View FIGURES 25–30). Tarsal organ capsulate with oval opening, located dorsally on tarsi. Two pectinate claws; tarsi I–II with long toothed claws, with 6–13 teeth ( Figs. 27View FIGURES 25–30, 35View FIGURES 31–36) tarsi III–IV with short toothed claws, with 3–4 teeth ( Fig. 36View FIGURES 31–36). Claw tufts composed of slightly widened tenent setae ( Fig. 27View FIGURES 25–30); presence of classic claw tuft clasper ( Figs 27View FIGURES 25–30, 35, 36View FIGURES 31–36) Abdomen oval, longer than wide, covered with sheet-shaped translucent scales ( Figs 29, 30View FIGURES 25–30); sparse tactile hairs with several barbs present; dorsum of abdomen anteriorly with long, dark, posteriorly curved setae; postepigastric invaginations present ( Figs 12View FIGURES 7–12, 37View FIGURES 37–42). Six spinnerets ( Figs 11View FIGURES 7–12, 38, 39View FIGURES 37–42); ALS elongated (0.6–0.13), separated from each other by one diameter or more ( Fig. 11View FIGURES 7–12), piriform gland spigots elongated with few associated setae ( Fig. 40View FIGURES 37–42), male with one major ampullate gland spigot ( Fig. 40View FIGURES 37–42) and female with two; PLS and PMS conical and short ( Figs. 41, 42View FIGURES 37–42), PLS larger than PMS ( Figs 11View FIGURES 7–12, 38View FIGURES 37–42); PLS in male with one minor ampular gland spigot and four aciniform gland spigots ( Fig. 41View FIGURES 37–42), females apparently with one minor ampular gland spigot, five aciniform gland spigots and two cylindrical gland spigots; PMS in male with one minor ampular gland spigot and two aciniform gland spigots, females apparently with one minor ampular gland spigot, four aciniform gland spigots and two cylindrical gland spigots ( Fig. 42View FIGURES 37–42). Female pedipalp with one pectinate claw with 6–8 teeth ( Fig. 28View FIGURES 25–30). Male palp: tibia generally short, except in P. caecus  sp. nov., in which it is more elongated ( Figs 65, 66View FIGURES 65–68, 74View FIGURES 69–74), RTA single, directed apically ( Figs 54, 56View FIGURES 53–56); cymbium oval with apical scopula ( Figs 62View FIGURES 61–64, 66View FIGURES 65–68), paracymbium present ( Figs 43, 45View FIGURES 43–48, 54View FIGURES 53–56, 61View FIGURES 61–64); bulb rounded ( Figs 53, 55View FIGURES 53–56, 69–72View FIGURES 69–74) or ovoid ( Figs 61View FIGURES 61–64, 65View FIGURES 65–68, 73, 74View FIGURES 69–74); embolus long with curved tip and wide base, inserted between 9–12 o’clock position ( Figs 55View FIGURES 53–56, 71View FIGURES 69–74), or 3 o’clock position ( Figs 61View FIGURES 61–64, 74View FIGURES 69–74); conductor sclerotized, with curved tip, inserted

prolaterally ( Figs 47View FIGURES 43–48, 55View FIGURES 53–56, 70View FIGURES 69–74); median apopohysis inserted distally and with sclerotized tip( Figs 69–74View FIGURES 69–74). Epigyne: posterior margin of epigynal plate anteriorly curved ( Figs 51View FIGURES 49–52, 67View FIGURES 65–68) or truncate ( Fig. 59); conspicuous posteromedian atrium with well-defined margins ( Figs 59, 63View FIGURES 61–64). Vulva: secondary spermathecae elongated ( Figs 60, 64View FIGURES 61–64); fertilization ducts antero-lateral.

Distribution. Brazil: states of Pará (exclusively in ferruginous caves), Paraíba, Pernambuco, Bahia and São Paulo ( Fig. 75View FIGURES 75).

Composition. Six species: Paracymbiomma angelim  sp. nov.; P. doisirmaos  sp. nov.; P. pauferrense  sp. nov.; P. carajas  sp. nov.; P. bocaina  sp. nov.; P. caecus  sp. nov. ( Figs 1–6View FIGURES 1–6).