Amara (Harpaloamara) latithorax Baliani, 1934

Kavanaugh, David H., Hieke, Fritz, Liang, Hongbin & Dong, Dazhi, 2014, Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Zabrini (Coleoptera, Carabidae), ZooKeys 407, pp. 55-119 : 76-77

7. Amara (Harpaloamara) latithorax Baliani, 1934 Figs 5 b–c, 5e, 6b, 15, 22b, 23b, 27-30

Amara (Harpaloamara) latithorax Baliani, 1934a: 198. Type material: Holotype male, allotype, and 1 additional paratype in BMNH, 1 paratype in MCSNG, and 3 paratypes in FRSDD. Type locality: India, United Provinces, Kumaon, West Almora and Chakrata, Mundali.

Amara (Bradytus) interrupta Landin, 1955: 417. Type material: Holotype female in NHRS. Type locality: Myanmar (Burma), Kambaiti, 7000 ft. Synonymized by Hieke (1997: 227).

Amara (Bradytus) neglecta Landin, 1955: 416. Type material: Holotype male in NHRS. Type locality: Myanmar (Burma), Kambaiti, 2000 m. Synonymized by Hieke (1997:227).

Diagnosis.

Adults of this species can be distinguished from those of all other species in the region by the following combination of character states: body length 8.5-9.0 mm; dorsal surface black, with very faint metallic blue or green metallic reflection in most individuals, more vivid (as in Fig. 15a) in or lacking from a few specimens, at least femora and outer antennomeres dark (piceous to black); pronotum (Fig. 15a) only slightly narrowed anteriorly with anterior margin nearly straight and clearly narrower than posterior margin, without or with only very slightly projected anterior angles, posterior angles distinctly rounded (narrowly so in some individuals), slightly to moderately obtuse, pronotal base moderately coarsely punctate, outer basal impressions either not sharply delimited laterally by raised areas or, if so, then the raised area broader; elytra elytra with slight sub-basal depressions centered on striae 6 (visible on right elytron in Fig. 15a), also on striae 4 and/or 5 in some individuals, parascutellar pore puncture absent; medial protibial spurs simple; tarsomere 5 of hind tarsi with two or (in a few specimens) three pairs of setae ventrally (Fig. 6b); last abdominal sternite of male with one pair (Fig. 5e) and female with two pairs (Fig. 5c) of setiferous punctures near hind margin.

Habitat distribution.

Specimens of this species were collected in daytime from under stones and other cover in open roadside areas (Figs 22b, 23b) and meadows with scattered grasses and shrubs, at the edges of agricultural fields, including wet and dry rice paddies and on open sandy banks of streams, and in these same habitats at night, when beetles were found active on bare substrate. Members of this species were found at elevations ranging from 1195 to 2022 m, syntopic with adults of Amara chalciope , Amara congrua , Amara dissimilis , Amara lucidissima and Amara sikkimensis at one or more sites.

Geographical distribution within the Gaoligong Shan.

Fig. 15d. We examined a total of 37 specimens (12 males and 25females) from the following localities: Fugong County: Aludi Village (Nu Jiang, 27.10834°, 98.87218°, 1195-1250 m, 22 April 2004, D.H. Kavanaugh collector [1 female; CAS]); Lishadi Township (Shilajia Village at Shibali Road on North Fork Yamu He, 27.13947°, 98.82184°, 1800-1900 m, 24-25 April 2004, D.H. Kavanaugh & C.E. Griswold collectors [1 female; CAS]), (11 km above Nu Jiang on Yaping Road at Shimowa Village, 27.13839°, 98.82147°, 1850-1928 m, 25 April 2004, H.B. Liang collector [1 male and 2 females; CAS, IOZ, ZMHB]); Shangpa Township (road on west side of Nu Jiang S of Shangpa, 26.88952°, 98.86539°, 1223 m, 22 and 27 April 2004, D.H. Kavanaugh & C.E. Griswold collectors [1 male and 1 female; CAS, IOZ]). Gongshan County: Cikai Township (15 km W of Cikai on Dulong Valley Road, 27.79584°, 98.58443°, 2022 m, 10 October 2002, D.H. Kavanaugh, P.E. Marek & H.B. Liang collectors [1 female; CAS]), (Heiwadi Village, 27.72250°, 98.59902°, 1965 m, 14 November 2004, H.B. Liang, D.Z. Dong & G. Tang collectors [2 males; IOZ, ZMHB]), (Pula He just above Nu Jiang Road, 27.74861°, 98.66675°, 1440 m, 23 October 2004, D.H. Kavanaugh & H.B. Liang collectors [1 female; IOZ], 11 November 2004, D.H. Kavanaugh, H.B. Liang, D.Z. Dong & G. Tang collectors [1 male and 2 females; CAS, IOZ]); Dulongjiang Township (Bapo, 27.73902°, 98.34975°, 1412 m, 20 October 2004, H.B. Liang collector [1 female; IOZ], 3 November 2004, H.B. Liang collector [2 males and 1 female; CAS, IOZ], at Miliwang, 27.72383°, 98.36117°, 1956 m, 31 October 2004, H.B. Liang collector [6 females; CAS, IOZ]), (Dulong Jiang at Elideng Village, 28.00287°, 98.32145°, 1640 m, 3 November 2004, D.H. Kavanaugh, D.Z. Dong & G. Tang collectors [1 female; CAS]), (2.3-3.3 airkm S of Longyuan Village on Dulong Jiang, 28.00532°, 98.32145°, 1685-1720 m, 2 November 2004, D.H. Kavanaugh, D.Z. Dong & G. Tang collectors [1 female; IOZ], 2.8 km S of Longyuan Village on Dulong Jiang, 28.00905°, 98.32204°, 1660 m, 2 November 2004, D.H. Kavanaugh & D.Z. Dong collector [1 male; CAS]), (main road between Mabidang and Kongdang, 27.76361°, 98.34111°, 1329 m, 5 November 2004, V.F. Lee & D.G. Long collectors [1 female; CAS]), (Maku, 27.68553°, 98.30425°, 1823 m, 2 November 2004, H.B. Liang collector [2 males and 3 females; CAS, IOZ]), (Moqie Wang at KM 91 on Gongshan-Dulong Road, 27.89934°, 98.34999°, 1550 m, 6 November 2004, D.H. Kavanaugh & H.B. Liang collectors [1 male and 1 female; CAS, IOZ]).

Members of this species were collected only in the northern half of the study area (Core Areas 1, 2 and 3), on both sides of the mountain range (Core Areas 1 and 2).

Overall geographical distribution.

Fig. 27. This species has been recorded from Bhutan, China (Yunnan Province), India (Sikkim, Uttar Pradesh and West Bengal), Myanmar and Nepal. Its occurrence in the study area represents the southern and eastern limits of its known geographical range.