Endroedyolini Davis, Deschodt & Scholtz, 2019

Davis, Adrian L. V., Deschodt, Christian M. & Scholtz, Clarke H., 2019, Defining new dung beetle tribes to resolve discrepancies between phylogeny and tribal classification in the subfamily Scarabaeinae (Coleoptera: Scarabaeidae), Zootaxa 4608 (1), pp. 131-144 : 138-139

Tribe Endroedyolini Davis, Deschodt & Scholtz , new tribe

Type genus. Endroedyolus Scholtz & Howden, 1987 , here designated ( Figs. 3 , 6 ).

Head. Clypeus bidentate with a notch to the side of each denticle. Antennae nine segmented.

Pronotum. Very convex with lateral margins explanate, sometimes carinate laterally.

Elytra. Strongly convex; setae in straight and slightly converging lines; metathoracic wings always absent.

Sterna. Smooth to punctate with punctures of varying density and size; prosternum fairly well developed; metasternum smooth to punctate, sometimes with deep fovea extending from anterior to sometimes half way between mesocoxae.

Protibia. With three small denticles on outside margin.

Meso- and metatibia. Unmodified with setae.

Pygidium. With small setiferous punctures; with or without depression or fovea.

Aedeagus. Parameres symmetrical.

Body size. Very small to small: 1.5mm to 4.8 mm.

Diagnosis and known distribution. The combination of extreme small size, very convex body, unique clypeal shape and explanate lateral pronotal margins, distinguishes members of this tribe from any other. They are restricted to forest patches of south east South Africa where they are associated with forest litter.

Figures

FIGURE 3. Endroedyolus paradoxus Scholtz & Howden, 1987, type species of the type genus of the tribe Endroedyolini Davis, Deschodt & Scholtz, new tribe.

FIGURE 6. Map showing the distribution of the known species in the tribe, Endroedyolini Davis, Deschodt & Scholtz, new tribe.