Ocys tachysoides (Antoine, 1933)

Maddison, David R. & Anderson, Roy, 2016, Hidden species within the genus Ocys Stephens: the widespread species O. harpaloides (Audinet-Serville) and O. tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini), Deutsche Entomologische Zeitschrift 63 (2), pp. 287-301 : 296-297

Ocys tachysoides (Antoine, 1933) Figs 2B; 3B; 6C, D; 7C, D; 8B, 9C, D; 10C, D; 11C, D

Bembidium harpaloides v. tachysoides Antoine, 1933:79. Holotype female, in the MNHN, examined, with three labels: "O. Nefifik (Maroc) Antoine I.1928" [partly handwritten], “Holotype” [handwritten on red paper], " tachysoides Antoine det. m." [partly handwritten]. Type locality: Mouth of the Oued Nefifikh, east of Casablanca, Morocco (in the vicinity of 33.72°N, 7.34°W).

Nomenclatural notes.

The type of Bembidium harpaloides v. tachysoides Antoine belongs to this species. We have examined external features of the holotype, including elytral microsculpture (which has a density of 21 lines per 0.1 mm), as well as shape of the spermatheca. Both second gonocoxites are broken in the holotype, and thus we were not able to measure their lengths.

Diagnosis.

Body length 4.0-5.8 mm ( Toribio 2013 and our observations; average length of males 4.79 mm (n=7) of females 5.16 mm (n=7)). Head and pronotum a deep red-brown contrasting with the dark brown to blackish elytra (Fig. 3), only the epipleural gutters and sometimes sutures paler, light reddish-brown. Occasionally with a bluish lustre. Microsculpture of elytral disc less transverse, with more of tendency to form sculpticells (Fig. 7C, D); density of microsculpture lines 20-21 per 0.1 mm (n=4 males). Hind margin of prothorax with a slight emargination laterally, such that the hind margin at the hind angles is directed posteriorly (Fig. 10C, D). Elytra less parallel-sided, greatest width around middle. Basal margin at shoulder in most specimens more or less straight (Fig. 11C), relatively few specimens slightly arcuate with a forward-directed concavity (Fig. 11D). Elytral striae 2 through 4 more marked in the apical third. Aedeagus with ventral margin bend more or less straight (Fig. 6C, D); apex less rounded, blunt. Anterior edge of central sclerite complex of internal sac angulate; brush sclerite smaller; dorsal membranes of internal sac paler, less evident. Gonocoxite relatively short (Fig. 8B), GCR=0.55-0.60 (average 0.58, n=5); spermathecal margin opposite the efferent duct of the spermathecal gland more or less straight (Fig. 8C,D; n=5).

Geographic distribution

(Fig. 12). In Africa, known from Morocco ( Antoine 1955). In Europe, from Portugal ( Sciaky 1998), Spain ( Toribio 2013), France, Belgium, Germany, and the United Kingdom. Examination of specimens in additional collections will likely show it to be more widely distributed.

Specimens examined.

In addition to the type specimen, and those listed in Table 1, we examined specimens from France: Vaires (MNHN).