Coptoborus furiosa, Smith & Cognato, 2021

Smith, Sarah M. & Cognato, Anthony I., 2021, A revision of the Neotropical genus Coptoborus Hopkins (Coleoptera, Curculionidae, Scolytinae, Xyleborini), ZooKeys 1044, pp. 609-720 : 609

Coptoborus furiosa sp. nov. Figure 6J-L, P

Type material.

Holotype, female, Ecuador: Los Ríos Prov., Canton La Clementina, Samama Nature Reserve, 01°38.852'S, 79°19.867'W, 381-430 m, 13-15.v.2015, Cognato, Smith, Osborn, Martinez et al., sample EC 15, ex 7 cm diameter bole from large tree fall (PUCE). Paratypes, female, as holotype (MSUC, 1; PUCE, 1); as holotype except: sample EC 32, ex 3 cm dia. branches of tree fall (NMNH, 1: PUCE, 1).

Diagnosis.

2.1-2.2 mm (mean = 2.12 mm; n = 5), 3-3.14 × as long as wide. This species is distinguished by the elytral apex attenuate and strongly emarginate, declivity convex, declivital interstriae 2 denticulate, elytral apex with interstriae 3 and 9 joining, forming a crenulate carina that continues submarginally to apex, declivital interstriae 3 densely denticulate with more than ten denticles, and elytral apices sharply acute.

Similar species.

C. inornatus , C. janeway , C. martinezae , C. tolimanus , C. vasquez .

Description

(female). 2.1-2.2 mm (mean = 2.12 mm; n = 5), 3.0-3.14 × as long as wide (holotype 2.1 mm, 3.0 × as long as wide). Body brown, elytra darker, antennae and legs lighter. Head: epistoma smooth. Frons subshiny, finely punctate, setose; each puncture bearing a long, erect hair-like seta. Eyes broadly and moderately emarginate. Submentum narrow, triangular, deeply impressed. Antennal scape short and thick, much shorter than club. Pedicel shorter than funicle. Club longer than wide, flat, type 3; segment 1 corneous, convex on anterior face, occupying basal ~1/3; segment 2 narrow, transverse, corneous; segments 1 and 2 present on posterior face. Pronotum: 1.1 × as long as wide. In dorsal view long and rounded frontally, type 7, sides parallel in basal 3/4, rounded anteriorly; anterior margin without serrations. In lateral view elongate, disc longer than anterior slope, type 7, summit prominent, on anterior 3/5. Anterior slope with densely spaced, broad fine asperities, becoming lower and more strongly transverse towards summit. Disc strongly shiny with sparse, minute punctures, some longer hair-like setae at margins. Lateral margins obliquely costate. Elytra: 1.9 × as long as wide, 1.6 × as long as pronotum. Scutellum small. Elytra attenuate, parallel-sided in basal 62%, then acutely tapered to apex, apex acutely produced, strongly emarginate. Disc smooth, strongly shiny; strial punctures large, deep, glabrous; interstriae flat, sparsely, minutely punctate, unarmed, each puncture bearing a long semi-erect seta. Declivity gradual, occupying ~1/3 of elytra, smooth, shining, declivital face convex; striae not impressed, strial punctures larger, deeper than those of disc, each puncture bearing a semi-erect seta as long as two punctures; interstriae flat, interstriae denticulate along their entire lengths, interstriae 3 very densely denticulate, denticles separated by no more than the width of two denticles and with at least ten denticles, interstrial setae erect, bristle-like, uniseriate, interstriae 1 with an additional row of slightly shorter erect hair-like setae. Posterolateral margin with interstriae 3 and 9 joining, forming an acutely denticulate carina and continuing submarginally to apex. Legs: protibiae obliquely triangular, broadest at apical 1/4; apical 1/2 of outer margin with five large, socketed denticles, their length longer than basal width. Meso- and metatibiae flattened; outer margin evenly rounded with six and seven large, socketed denticles, respectively.

Etymology.

Portrayed by Charlize Theron, Imperator Furiosa is the heroine in the movie 'Mad Max: Fury Road’ (2015). The “spiny” elytra give the species a fierce appearance. Noun in apposition.

Distribution.

Ecuador (Los Ríos).

Biology.

This species was found in a bole and branches of an unidentified tree 3-7 cm in diameter.

Figures

Figure 6. Dorsal, lateral, frontal and declivital view of Coptoborus erwini holotype, 1.4 - 1.5 mm (A-C, M), C. exilis 2.3 - 2.4 mm (D-F, N), C. exutus holotype, 1.9 mm (G-I, O), C. furiosa holotype, 2.1 - 2.2 mm (J-L, P). All photographs by SMS, G-I, O copyright National Museum of Natural History, Smithsonian Institution, Washington, D. C., published by permission.