Trogossitini Latreille, 1802, 1876

Kolibac, Jiri, 2013, Trogossitidae: A review of the beetle family, with a catalogue and keys, ZooKeys 366, pp. 1-194 : 37-38

Tribe Trogossitini Latreille, 1802

Trogossitini Latreille, P. A. 1802: 110.

Type genus:

Trogossita Olivier, 1790 (= Temnoscheila Westwood, 1830)

Burakowski, B. et al. 1986: 115 ( Nemosomatinae ). Kolibáč, J. 2006: 120 (diagnosis, stat. n.). Kolibáč, J. 2007a: 364 (phylogeny). Kolibáč, J. 2008: 118-119 (phylogeny)

Remarks.

Two character analyses of Trogossitini ( Kolibáč 2006, 2008) separate off a monophyletic group composed of the genera Temnoscheila , Nemozoma , Tenebroides , Corticotomus , and Leipaspis . The other two genera analyzed, Airora and Alindria , are more primitive. Their position in the cladogram of 2008 (p. 119) makes Trogossitini paraphyletic with reference to Gymnochilini but the original analysis (2006) unambiguously set the two groups appart as distinct monophyletic clades. A classification of Seidlitzella has been discussed above, in the “Remarks” section of the Gymnochilini entry.

There are also some genera that are not included in the two character analyses because of insufficient data sets, namely Dupontiella , Elestora , Eupycnus , Euschaefferia , and Parallelodera . The classification of all these rather advanced genera within Trogossitini is undeniable, apart from the monotypic Elestora which is obviously related to Melambia , for which the systematics are quite complicated and in need of revision.

Most of the members of Trogossitini lead the kind of life typical of predatory Cleridae , especially of the subfamilies Clerinae and Tillinae . Adults hunt for xylophagous insects (e.g. Curculionidae : Scolytinae , Bostrichidae ) on branches and logs while larvae dwell and hunt under bark or in galleries. However, some trogossitine adults live in insect galleries together with their larvae (e.g. Nemozoma ). The trogossitins are not as efficient in the air as the gymnochilins, and neither do they move so swiftly on the ground.

Key to the recent genera

1	Frons with conspicuous sharp longitudinal groove	2
-	Frons without groove, with shallow depression at most	3
2	Anterior part of cranium (frons) with two large horn-like processes; body extremely elongate, small (about 3-6 mm)	Nemozoma
-	Anterior part of cranium (frons) without distinct processes; body not extremely elongate, larger (about 7-25 mm)	Temnoscheila
3	Pronotum conspicuously elongate, weakly narrowed at base; body elongate and cylindrical	4
-	Pronotum transverse or quadrate or narrowed at base; elytra widest in apical third and somewhat flattened	8
4	Pronotum somewhat cordate; elytra with carinae; large species (about 10-35 mm)	Alindria
-	Sides of pronotum nearly parallel; elytra without carinae; smaller species (about 2-15 mm)	5
5	Outer margins of all tibiae with large spines; antennae reach backwards anterior margin of pronotum; larger species (about 7-15 mm)	Airora
-	Outer margin of tibiae with 2-3 spines at apex at most; antennae reach back to beyond anterior margin of pronotum; smaller species (about 2-5 mm)	6
6	Pronotum conspicuously narrowed (constricted) at base	Dupontiella
-	Pronotum not narrowed at base, oblong	7
7	Pronotum with distinctly raised lateral margins; submentum distinctly separated from gula in front, outer angles not prominent; at least front tibiae with spines at apex	Corticotomus
-	Pronotum without distinctly raised lateral margins, apical angles obliterated; submentum not distinctly separated from gula in front, outer angles prominent and produced apically at least to base of mandibles; tibiae without spines	Euschaefferia
8	Elytra with conspicuous carinae and regular punctation	9
-	Elytra without carinae, with regular sculpture only	10
9	Dorsal body surface distinctly flattened; very wide, black species, elytra with four striking orange spots; mesonotum with long orange hairs	Elestora
-	Dorsal body surface not distinctly flattened, almost cylindrical; elongate, almost cylindrical, unicolorous (black or brown) species without colour pattern	Melambia
10	Body including head and pronotum distinctly elongate; pronotum constricted at base	Leipaspis
-	Body not so elongate; sides of pronotum subparallel or cordate	11
11	Tarsal pattern 4-4-4: 1st tarsomere coalescent with 2nd tarsomere in all pairs of legs; elytra rather convex	Parallelodera
-	Tarsal pattern 5-5-5; elytra rather flattened	12
12	All tibiae with about 3-6 conspicuous spines along outer margin; pronotum subparallel, elongate; labrum retracted, hardly visible; body more coarsely sculptured	Eupycnus
-	All tibiae with about 2-4 spines along outer margin; pronotum cordate, approximately as long as wide; labrum distinctly visible; body sculpture finer	Tenebroides