Carabus (Bothriopterus), Chaudoir, 1835

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 506

Subgenus Bothriopterus Chaudoir, 1835

Tilodes Fischer von Waldheim, 1829a: 19 [potential nomen oblitum, see Bousquet (2002c: 179)]. Type species: Carabus oblongopunctatus Fabricius, 1787 designated by Bousquet (1999: 104). Etymology. Unknown [masculine].

Bothriopterus Chaudoir, 1835: 447 [potential nomen protectum]. Type species: Carabus oblongopunctatus Fabricius, 1787 designated by Chaudoir (1838: 9). Etymology (see Chaudoir 1838: 9). From the Greek bothros (pit, dimple) and pteron (wing, by extension elytron), probably alluding to the foveolate discal setigerous punctures on the third elytral interval in adults of Pterostichus oblongopunctatus and related species [masculine].

Dysidius Chaudoir, 1838: 8. Type species: Feronia morosa Dejean, 1828 (= Feronia muta Say, 1823) by original designation. Synonymy established by Goulet (1974a: 27). Etymology (original). From the Greek dys - (bad, with difficulty; referring to by Chaudoir as deplorable, sad) and eidos (form, appearance), alluding to the specific name of the sole species ( Feronia morosa ) Chaudoir attributed to this genus-group taxon [masculine].

Geopezus Gistel, 1856: 358. Type species: Carabus oblongopunctatus Fabricius, 1787 by monotypy.

Parargutor Casey, 1918: 324. Type species: Pterostichus lustrans LeConte, 1851 by original designation. Synonymy established, under the name Dysidius Chaudoir, by Lindroth (1966: 489). Etymology. From the Greek para (near, beside) and the generic name Argutor [q.v.] [masculine].

Diversity.

Eighteen species (Lorenz 2005: 273) in the arctic (marginal), subarctic, boreal, and temperate areas of the Nearctic (six species, one of them circumboreal) and Palaearctic (12 species, including two in the Himalayas) Regions, with one species ( Pterostichus tropicalis Bates) in the mountains of northern Mexico.

Identification.

Tschitschérine (1900d) published a key to the species but his work is outdated. Lindroth (1966: 484-491) reviewed the North American species under his adstrictus and mutus groups.