Verodes sparsus (Champion)

Foley, Ian A. & Ivie, Michael A., 2008, A phylogenetic analysis of the tribe Zopherini with a review of the species and generic classification (Coleoptera: Zopheridae), Zootaxa 1928 (1), pp. 1-72 : 64-68

Verodes sparsus (Champion)

Fig. 62

Nosoderma sparsum Champion, 1884: 50 , pl. 2, fig. 12. García-París et al., 2006: 230.

Nosoderma sparsa ; Casey, 1907b: 481

Noserodes sparsus ; Gebien, 1936: 670.

Noserodes sparsa ; Blackwelder, 1945: 515.

Verodes sparsus ; Foley and Ivie, 2007: 72 .

Diagnosis: The species can be recognized based on its reduced elytral sculpture, with just a few weak ridges and nodules, and by the elongate body form with the elytra 1.95–2.05X longer than the widest point (some specimens of V. exsculptus also have an elongate body form, but that species has intertuberculate setae, and well defined ridges and nodules).

Distribution: Guatemala at elevations between 2,500 and 3,900 meters.

Recorded distribution: A summary of the 14 specimens examined is GUATEMALA: SAN MARCOS, SOLOLA, TOTONICAPÁN .

Types: Nosoderma sparsum : LECTOTYPE: ♀, missing left meso tarsus in BMNH. Orange ringed circle Type/Totonicapama,; 85– 10,500 ft. Champion./ Nosoderma sparsum Champ. / Sp. Figured/ B.C.A.Col.IV.1.; Nosoderma sparsum / red square Lectotypus Nosoderma ; sparsum ; Champion; García-París des 2000. PARA- LECTOTYPE: ♂, intact in BMNH. Same data as LECTOTYPE.

Zopherinus Casey, 1907 (Syn. Triplehorn 1972)

Zopherodes Casey, 1907 (Syn. Triplehorn 1972)

Zopherus chilensis Gray, 1832 C. & S. America

Zopherus insignis Blanchard, 1861 (Syn. Gebien 1936)

Zopherus bremei Guérin-Méneville (Syn. Gebien 1936)

Zopherus moreletii Lucas, 1852 (Syn. Gebien 1936)

Zopherus nervosus Solier, 1841: 42 . Mexico

Zopherus pectoralis LeConte, 1851 (Syn. Gebien 1936)

Zopherus reticulatus Champion, 1884 (Syn. Gebien 1936)

Zopherus compactus Champion, 1884 (Syn. Triplehorn 1972)

Zopherus marmoratus Casey, 1907 (Syn. Triplehorn 1972)

Zopherus nodulosus Solier, 1841: 43 . TX, Mexico

Zopherus variolosus Sturm, 1843 (Syn. Gebien 1936)

Zopherus sallaei Champion, 1884 (Syn. Gebien 1936)

Z. haldemani Horn, 1870 (Syn. Triplehorn 1972)

Zopherus verrucosus Champion, 1884 (Syn. Gebien 1936)

Zopherus jourdani Sallé, 1849 C. America

Zopherus mexicanus (auctorum, not Gray)

Zopherus costaricensis Champion, 1884 (Syn. Triplehorn 1972)

Zopherus jansoni Champion, 1884 C. America

Zopherus mexicanus Gray, 1832 Mexico

Zopherus maculatus Champion, 1884 (Syn. Gebien 1936)

Zopherus angulicollis Champion, 1884 Chihuahua

Zopherus laevicollis Solier, 1841 Mexico

Zopherus venosus Champion, 1884 (Syn. Triplehorn 1972)

Zopherus tuberculatus Champion, 1884 (Syn. Triplehorn 1972)

Zopherinus limbatus Casey, 1907a (Syn. Triplehorn 1972)

Zopherus xestus Triplehorn, 1972 TX

Zopherus solieri Triplehorn, 1972 Durango

Zopherus championi Triplehorn, 1972 Mexico

Zopherus tristis LeConte, 1851 SW USA, Baja

Zopheroides aequalis Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides variabilis Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides incrustans Casey, 1907b (Syn. Triplehorn 1972)

Zopherus concolor LeConte, 1851 SW USA

Zopherus guttulatus Horn, 1867 (Syn. Triplehorn 1972)

Zopheroides morosus Casey, 1907 (Syn. Triplehorn 1972)

Zopherus gracilis Horn, 1867 SW USA, Sonora

Zopheroides pudens Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides caudalis Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides lugubris Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides pruddeni Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides luctuosus Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides elongatus Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides germinatus Casey, 1907 (Syn. Triplehorn 1972)

Zopherus uteanus (Casey), 1907 SW USA

Zopheroides mormon Casey, 1907 (Syn. Triplehorn 1972)

Zopherus granicollis Horn, 1885 SW USA, Baja Zopheroides induratus Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides californicus Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides prominens Casey, 1924 . (Syn. Triplehorn 1972)

Zopheroides ventriosus Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides parvicollis Casey, 1907 (Syn. Triplehorn 1972)

Zopherus opacus Horn, 1867: 161 . SW USA

Zopherus elegans Horn, 1870: 272 . SW USA

Zopheroides otiosus Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides verrucipennis Casey, 1907 (Syn. Triplehorn 1972)

Zopheroides circumductus Casey, 1924 (Syn. Triplehorn 1972)

Zopheroides woodgatei Casey, 1924 (Syn. Triplehorn 1972)

Zopherus sanctaehelenae (Blaisdell), 1931 CA

Zopherosis White, 1859

Zopherosis georgei White 1859 E. Australia

Phloeodes LeConte, 1862

Noserus LeConte, 1862 (Syn. Ṡ lipi ń ski & Lawrence 1999)

Ageonoma Pascoe, 1866 (Syn. Horn 1870)

Ph. diabolicus (LeConte) 1851 CA, Baja

Nosoderma pustulosus (LeConte) 1859 [NEW SYNONYMY]

Ph. latipennis Casey 1907 (Syn. Gebien 1936)

Ph. ovipennis Casey 1907 (Syn. García-París et al. 2006)

Ph. elongatus Casey 1907 (Syn. García-París et al. 2006)

Ph. scaber Casey 1907 (Syn. García-París et al. 2006)

Ph. angustus Casey 1907 (Syn. García-París et al. 2006)

Ph. remotus Casey 1907 (Syn. García-París et al. 2006)

Ph. plicatus (LeConte) 1859 CA, Baja

Noserus torvus Casey 1907 (Syn. García-París et al. 2006)

Noserus collaris Casey 1907 (Syn. García-París et al. 2006)

Noserus corrosus Casey 1907 (Syn. García-París et al. 2006)

Noserus convexulus Casey 1907 (Syn. García-París et al. 2006)

Ph. venustus (Champion) 1884: 45 . [NEW COMBINATION] C. America

Sesaspis Casey, 1907b: 469 . [NEW STATUS, NEW SENSE]

Sesaspis denticulata (Solier), 1841 Mexico

Sesaspis emarginata (Horn), 1878 [NEW COMBINATION] SW USA, Tamaulipas Sesaspis triplehorni [NEW SPECIES] C. America

Sesaspis lutosa (Champion), 1884 [NEW COMBINATION] Mexico

Sesaspis sylvatica (García-París et al.), 2006 [NEW COMBINATION] Mexico

Sesaspis doyeni (García-París et al.), 2001 [NEW COMBINATION] Nuevo Leon

Sesaspis ashei [NEW SPECIES] Mexico

Verodes Casey, 1907 ( sensu Foley & Ivie 2007 )

Nosoderma Solier, 1841 not Guérin-Méneville 1838

Noserodes Casey, 1907 (Syn. Doyen & Lawrence 1979)

V. aequalis (Champion), 1884 Oaxaca

V. inaequalis (Say), 1835 Mexico

Nosoderma morbillosum Solier, 1841: 37 . (Syn. Gebien 1936)

Nosoderma vicinum Solier, 1841: 38 (Syn. Gebien 1936)

Nosoderma prominens Casey, 1907 [NEW SYNONYMY]

Nosoderma senex Casey, 1907 [NEW SYNONYMY]

Nosoderma brevicollis Casey, 1907 [NEW SYNONYMY]

Nosoderma subglabrus Casey, 1907 [NEW SYNONYMY]

V. championi Casey, 1907 Mexico

V. interruptus (Champion), 1884 Oaxaca

Nosoderma insigne Champion, 1884 [NEW SYNONYMY]

V. scabrosus (Solier), 1841 Oaxaca

Scoriaderma congolense Fairmaire 1894 [NEW SYNONYMY]

V. asperatus (Champion), 1884 Mexico

V. exsculptus (Champion), 1884 Mexico

Nosoderma anceps Champion, 1884 (Syn. García-París et al. 2006)

Nosoderma longipennis Casey, 1907b [NEW SYNONYMY]

V. carinatus (Champion), 1884 Veracruz

V. impressus (Champion), 1884 Veracruz

V. zunilensis (Champion), 1884 Honduras V. guatemalensis (Champion), 1884 C. America

Noserodes squalidus Casey, 1907 [NEW SYNONYMY]

V. sparsus (Champion), 1884 Guatemala

Nosoderma Guérin-Méneville, 1838

Meralius Casey, 1907 (Syn. Foley & Ivie 2007)

Nosoderma echinatum Guérin-Méneville, 1838

Nosoderma duponchelii Solier, 1841 (Syn. Gebien 1936)

Nosoderma turquinense (Garrido), 2004 Cuba

M. montanus Garrido, 2004 [NEW SYNONYMY]

Noserinus Casey, 1907

Noserinus dormeanus (Fairmaire) 1889 S. America

Noserinus annulatipes Casey (Syn. Gebien 1936)

Noserinus furcatus (Kirsch) [NEW COMBINATION] S. America

Scoriaderma Fairmaire, 1894

Scoriaderma cordicolle Waterhouse, 1880 E. Africa

Scoriaderma comoriense Fairmaire, 1894 [NEW SYNONYMY]

Zopher Š lipi ń ski & Lawrence, 1999

Zopher iviei Š lipi ń ski & Lawrence, 1999 Malaysia

Phellopsis porcata

Zopherus nodulosus

Zopherosis georgei

Zopher iviei

Noserinus dormeanus

Noserinus furcatus

Scoriaderma cordicolle Nosoderma echinatum Nosoderma turquenense Sesaspis emarginata

Sesaspis triplehorni

Sesaspis ashei

Sesaspis doyeni

Sesaspis sylvatica

Sesaspis lutosa

Sesaspis denticulata

Phloeodes venustus

Phloeodes diabolicus

Phloeodes plicatus

Verodes zunilensis

Verodes aequalis

Verodes sparsus

Verodes guatemalensis

Verodes inaequalis

Verodes championi

Verodes asperatus

Verodes carinatus

Verodes exsculptus

Verodes scabrosus

Verodes interruptus

Figures

FIGURES 61–64. Dorsal habiti. 61. Verodes interruptus, Mexico, Oaxaca; 62. V. sparsus, Guatemala; 63. V. zunilensis, Mexico, Chiapas; 64. V. guatemalensis, Mexico, Chiapas. Scale bars = 5mm.