Digitonthophagus Balthasar, 1959

Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed & Barbero, Enrico, 2019, An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae), Zootaxa 4658 (1), pp. 1-36 : 5-6

Genus Digitonthophagus Balthasar, 1959

Digitonthophagus aksumensis Génier in Génier & Moretto, 2017

Literature quotes. SAUDI ARABIA : Asir Province, 35 km N Mahayl [=Muhayil] (1850′N, 4158′E) . Madinah Prov- ince, Hedjaz (2430′N, 3830′E). Makkah Province, Lith, 10 mi. inland (209′N 4016′E); Wadi Jizan (2137′N 3910′E); 12 mi. N Jidda [=Jeddah] (2138′15′′N 3913′35’’E) ( Génier & Moretto, 2017) . YEMEN: Aden Governorate, Aden (1245′56’’N 451′1’’E). Al Hudaydah Governorate, Kamaran Island (1520′N 4235′E) . Lahij Governorate, Lahij (133′N 4453′E). Ta’izz Governorate, Taizz (1334′N 442′E) ( Génier & Moretto 2017).

Material examined. None.

Distribution. Arabian Peninsula: Saudi Arabia , Yemen. Africa: Djibouti, Eritrea, Ethiopia, Somalia ( Génier & Moretto 2017).

Digitonthophagus eucatta Génier in Génier & Moretto, 2017

[ Fig. 3 ]

Literature quotes. OMAN: Muscat governorate, “Mintaqat Masqat, Mascat (2336′48’’N 5835′36’’E)” ( Génier & Moretto, 2017).

Material examined. SAUDI ARABIA : Asir Province, Wadi Hali , 440 m (1830′N 4203′E), 11.ii.2016, H. Fadl, A. El Torkey, A. Al Ansi and M. Al Harbi leg. 2 ♂♂ ( KSMA) ; ibidem, 530 m (1830′N 4203′E), 11.ii.2016, D. Král leg 1 ex. ( NMPC) ; Wadi Marabah , 1159 m (1810′16″N 4222′15″E), 1.iv.2017, G. Magnani & D. Baiocchi leg. 7 exx. ( SZCM) ; ibidem, 4.iv.2017, G. Magnani & D. Baiocchi leg. 1 ex. ( SZCM) ; ibidem, 1197 m (1810′18″N 4222′12″E), 1/ 4.iv.2017, P. Rapuzzi leg. 7 exx. ( SZCM) ; Wadi Reem , 136 m (1752.551′N 4216.664′E), 9.ii.2016, H. Fadl, A. El Torkey, A. Al Ansi and M. Al Harbi leg. 2 ♀♀ ( KSMA) ; 10 km N Wadi Reem , 200 m (1755′28″N 4215′20″E), 30.iii.2017, G. Magnani & D. Baiocchi leg. 1 ex. ( SZCM) . Baha Province, 5 km N Al Makhwah , 464 m (1949′47′′N 4126′47′′E), 30.iii.2017, G. Magnani & D. Baiocchi leg. 3 exx. ( SZCM) ; ibidem, 30.iii.2017, P. Rapuzzi leg. 1 ex. ( SZCM) . Jizan Province, Abu Arish, Ali Abu Al-Shawari Farm , 73 m (1659.379′N 4249.662′E), 10.v.2018, H. Al Dhafer, M. Abdel-Dayem, A. El Gharbawy & I. Rasool leg. 2 ♀♀ ( KSMA) ; Wadi Atoud , 245 m (1748′N 4222′E), 8.ii.2016, H. Fadl, A. El Torkey, A. Al Ansi and M. Al Harbi leg. 10 ♀♀, 1 ♂ ( KSMA) ; ibidem, 8.ii.2016, D. Král leg. 4 exx. ( NMPC) ; 245 m (1748′N 4222′E),. Makkah Province, Wadi Ghanuna , 366 m (1925.341′N 4136.329′E), 11.v.2011, H. Fadl, A. El Torkey, M. Sharaf and H. Setyaningrum leg. 2 ♀♀, 1 ♂ ( KSMA) . Riyadh Province, Ri- yadh, vi.2009, 1 ex. ( SZCM) . YEMEN: Sana’a Governorate, Wadi Dhahr , 2240 m (1526′43″N 4467′34″E), 15/ 17. xi.2003, D. Král leg. 5 exx. ( NMPC) .

Distribution. Arabian Peninsula: Oman ( Génier & Moretto 2017). Africa: Benin, Burkina Faso, Cameroon, Chad, Eritrea, Ethiopia, Gambia, Mali, Mauritania, Niger, Nigeria, Senegal, Sudan ( Génier & Moretto 2017).

Remarks. First established records for Saudi Arabia and Yemen.

After the revision of Génier & Moretto (2017), the following citations:

- Onthophagus catta (Fabricius) : Arabia ( Arrow 1931) ;

- Onthophagus catta (Fabricius) : Saudi Arabia , Asir region, Ha’il region. Yemen ( Paulian 1948);

- Onthophagus gazella (Fabricius) : Saudi Arabia , “Hedjaz” (d’Orbigny 1898c). Yemen, “ Aden ” (d’Orbigny 1898c); “ Lahej ” ( Balthasar 1972). “Arabie” ( Baraud 1987);

- Onthophagus (Digitonthophagus) gazella (Fabricius) : “Arabien” ( Balthasar 1963). Saudi Arabia , “Wadi Hesua nr. Wadi ad Villa” ( Zunino 1979);

- Digitonthophagus gazella (Fabricius) : Saudi Arabia , “Bahara”; “Al Hakimah”; “Selouly Farm”; “Ad Darba”; “Gizan”; “Sharoura” ( Zunino 1981). Yemen (Rössner in Ziani et al. 2015),

should be referred to as either D. aksumensis or D. eucatta .

Figures

FIGURES 2–7. Onthophagini of Arabian Peninsula. 2, Cleptocaccobius viridicollis (Fåhraeus in Boheman, 1857): YE (Sana’a) Wadi Dhahr; 3, Digitonthophagus eucatta Génier in Génier & Moretto, 2017: SA (Asir) Wadi Marabah; 4, Euonthophagus carbonarius (Klug, 1855) sensu lato: YE (Sana’a) Wadi Dhahr; 5, Onthophagus (Eremonthophagus) infuscatus Klug, 1845: SA (Asir) 10 km N Wadi Reem; 6, Onthophagus (Eremonthophagus) sticticus Harold, 1867: OM (Dhofar) Ash-Shuwayiyah; 7, Onthophagus (Eremonthophagus) transcaspicus König, 1889: OM (Ad Dakhiliyah) Al Ayn. Photos by Gabriele Fiumi.