Anteon Jurine, 1807

Xu, Zaifu, Olmi, Massimo & He, Junhua, 2013, Dryinidae of the Oriental region (Hymenoptera: Chrysidoidea), Zootaxa 3614 (1), pp. 1-460 : 59-67

5. Genus Anteon Jurine, 1807

Anteon Jurine 1807: 302 ; R. Perkins 1907: 50; Kieffer 1914b: 140; Berland 1928: 157; Richards 1937: 118; Richards 1939: 245; J. Perkins 1976: 13; Krombein 1979: 1240; Olmi 1984: 286; Olmi 1994: 47; Olmi 1995g: 5; Olmi 1999a: 105; He & Xu 2002: 113; Virla & Olmi 2008: 370; Moya-Raygoza & Olmi 2010: 87.

Antaeon Haliday 1833: 275 , invalid emend.

Dryinus Latreille : Walker 1837: 413.

Chelogynus Haliday 1838: 518 (synonymized by Kieffer in Kieffer & Marshall 1905); type species: Chelogynus infectus (Haliday in Walker 1837), designated by Muesebeck & Walkley 1951 (designation of Ch. gaullei by Kieffer (1914b) invalid, because the type was not an originally included species); R. Perkins 1907: 48; Kieffer 1914b: 172; Berland 1928: 162; Richards 1937: 118; J. Perkins 1976: 18.

Anteon Jurine partim: Kieffer in Kieffer & Marshall 1905: 130.

Neochelogynus R. Perkins 1905: 60 (synonymized by R. Perkins 1907);

type species: Neochelogynus typicus R. Perkins 1905 , by original designation.

Prosanteon R. Perkins 1905: 66 (synonymized by R. Perkins 1907);

type species: Prosanteon chelogynoides R. Perkins 1905 , by monotypy and original designation.

Paranteon R. Perkins 1905: 67 (synonymized by Olmi 1984);

type species: Paranteon myrmecophilus R. Perkins 1905 , by original designation.

Lasianteon Kieffer 1913b: 300 (synonymized by Muesebeck & Walkley 1951);

type species: Lasianteon rubrifrons ( Kieffer, 1905) , designated by Muesebeck & Walkley 1951; Kieffer 1914b: 197.

Xenanteon Kieffer 1913b: 300 (synonymized by Richards 1939);

type species: Xenanteon reticulatum ( Kieffer, 1905) , designated by Richards 1939; Kieffer 1914b: 193.

Liodryinus Kieffer 1913b: 300 (synonymized by Richards 1953);

type species: Liodryinus doddi ( Kieffer 1911b) , by original designation.

Allanteon Kieffer 1914b: 198 (synonymized by Berland 1928);

type species: Allanteon punctatus ( Kieffer 1905) , by monotypy.

Anteon subg. Anteon Jurine : Richards 1939: 246; Krombein 1979: 1240.

Anteon subg. Chelogynus Haliday : Richards 1939: 246; Krombein 1979: 1241.

Hirtanteon Benoit 1951b: 163 (synonymized by Olmi 1984);

type species: Hirtanteon canabensis Benoit 1951b , by original designation.

Chelogynus (Chelogynus) : J. Perkins 1976: 18.

Chelogynus (Neochelogynus) : J. Perkins 1976: 19.

Type species. Anteon jurineanum Latreille, 1809 , by subsequent monotypy.

Diagnosis. Female: fully winged ( Plate 28D ); rarely brachypterous; occipital carina complete; palpal formula 6/3; antenna without rhinaria; fore wing with three cells enclosed by pigmented veins (costal, median and submedian) ( Plate 28D ); fore wing with stigmal vein and pterostigma ( Plate 28D ); distal part of stigmal vein much shorter than proximal part ( Plate 28D ), occasionally slightly shorter, as long as, or longer than proximal part; propodeum usually with transverse keel between dorsal and posterior surface ( Plate 28D ); protarsus chelate ( Plate 28D ); inner side of enlarged claw with proximal prominence bearing one long bristle ( Plate 17C ); tibial spurs 1/1/ 2. Male: fully winged ( Plate 43C ); rarely brachypterous; occipital carina complete; vertex of head usually without two oblique keels connecting posterior ocelli to occipital carina; palpal formula 6/3; fore wing with three cells enclosed by pigmented veins (costal, median and submedian) ( Plate 43C ); fore wing with stigmal vein and pterostigma ( Plate 43C ); distal part of stigmal vein much shorter than proximal part ( Plate 43C ), occasionally slightly shorter, as long as, or longer than proximal part; pterostigma less than four times as long as broad ( Plate 43C ); propodeum usually with transverse keel between dorsal and posterior surface; paramere without inner branch wrapping penis ( Plate 17A ); tibial spurs 1/1/2.

Distribution. Worldwide.

Hosts. Cicadellidae ( Guglielmino & Olmi, 1997, 2006, 2007), Eurymelidae ( Guglielmino & Olmi, 1997, as Eurymelinae ).

World species. Four hundred and twenty-two species are known, 148 in the Oriental region.

Remarks. Larvae of Anteon were described by Ponomarenko (1975) .

Key to the species

Females (unknown in the species not present in the key to the females)

1. Segment 4 of protarsus much shorter than segment 1, at most about 0.5 as long as segment 1......................... 2

- Segment 4 of protarsus slightly shorter, as long as, or longer than segment 1..................................... 45

2. Posterior surface of propodeum without longitudinal keels, or rarely with two incomplete longitudinal keels less than 0.6 as long as posterior surface of propodeum.................................................................... 3

- Posterior surface of propodeum with two complete longitudinal keels........................................... 12

3. Scutum completely reticulate rugose and granulated, or sculptured by many irregular and longitudinal keels on whole surface.. 4

- Scutum smooth, at most with some irregular keels on sides of notauli............................................ 6

4. Segment 5 of protarsus with two rows of small lamellae and distal group of small lamellae ( Plate 34C ).. A. maai Olmi

- Segment 5 of protarsus with one row of long bristles or lamellae and one–two long distal lamellae ( Plate 35E )........... 5

5. Notauli absent or hardly shortly visible near anterior margin of scutum............................ A. metuendum Olmi

- Notauli reaching about 0.3–0.5 length of scutum.......................................... A. jurineanum Latreille

6. Fore wing with one dark transverse band or one dark spot beneath pterostigma..................... A. borneanum Olmi

- Fore wing hyaline, without dark transverse bands............................................................ 7

7. Arolium very large, approximately as long as enlarged claw ( Plate 50A )............................ A. wushense Olmi

- Arolium small, much shorter than enlarged claw ( Plates 20A, B , 25F , 45D )....................................... 8

8. Segment 5 of protarsus with basal region about as long as distal part ( Plate 25F )................................... 9

- Segment 5 of protarsus with basal region much longer than distal part ( Plates 20A, B , 45D )......................... 10

9. Head granulated and with very slight tracks of numerous irregular keels........................... A. dravidicum Olmi

- Head completely strongly reticulate rugose..................................................... A. crassum Mita

10. Segment 5 of protarsus with large inner expansion ( Plate 45D ).................................... A. spenceri Olmi

- Segment 5 of protarsus without large inner expansion ( Plates 20A, B , 139B )..................................... 11

11. Notauli very short, reaching about 0.3 length of scutum........................................ A. borneanum Olmi

- Notauli longer, reaching about 0.6 length of scutum........................................... A. zopyrum sp. nov.

12. Head black or brown, except mandible testaceous; clypeus totally testaceous; occasionally also with part of face testaceous 13

- Head black, except mandible testaceous; clypeus black, rarely with anterior half testaceous, never totally testaceous...... 18

13. Thorax totally black; notauli reaching approximately 0.5 length of scutum............................ A. dignum Olmi

- Thorax partly black, with at least pronotal tubercle testaceous; notauli reaching 0.6–0.9 length of scutum.............. 14

14. Segment 1 of protarsus approximately three times as long as segment 4.............................. A. lesagei Olmi

- Segment 1 of protarsus approximately twice as long as segment 4.............................................. 15

15. Scutellum testaceous-reddish............................................................. A. subdignum Olmi

- Scutellum black..................................................................................... 16

16. Face completely finely punctate, smooth; prothorax black; notauli reaching approximately 0.6–0.7 length of scutum........................................................................................ A. xuexini Xu, He & Olmi

- Anterior half of face dull, rugose, posterior half punctate, unsculptured among punctures; prothorax with at least pronotal tubercle testaceous or completely testaceous; notauli reaching approximately 0.8–0.9 length of scutum................ 17

17. Head black, except mandible testaceous....................................................... A. medium Olmi

- Head black, except mandible, clypeus and part of face testaceous....................... A. nanlingense Xu, Olmi & He

18. Head completely or almost completely strongly reticulate rugose.............................................. 19

- Head at most with irregular keels, or only with part of head reticulate rugose, never completely reticulate rugose........ 29

19. Scutum sculptured by longitudinal keels; scutellum smooth, slightly punctate..................... A. philippinum Olmi

- Scutum and scutellum without longitudinal keels........................................................... 20

20. Scutum completely reticulate rugose...................................................... A. rugosum Xu & He

- Scutum not reticulate rugose; occasionally partly reticulate rugose............................................. 21

21. Notauli absent.................................................................. A. clavatum Olmi & Currado

- Notauli present, occasionally very short.................................................................. 22

22. Scutum granulated and strongly punctate, or not granulated, but strongly punctate and with very wide impressions similar to areolae............................................................................................ 23

- Scutum punctate, smooth, unsculptured among punctures; rarely with few areolae on lateral regions.................. 24

23. Scutum granulated and strongly punctate..................................................... A. heppneri Olmi

- Scutum finely punctate or strongly punctate and with very wide impressions similar to areolae, not granulated. A. thai Olmi

24. Notauli very short, reaching approximately 0.15 length of scutum.................. A. confertilaminarum Xu, He & Olmi

- Notauli reaching approximately 0.3–0.7 length of scutum.................................................... 25

25. Notauli reaching about 0.6–0.7 length of scutum........................................................... 26

- Notauli reaching about 0.3–0.4 length of scutum........................................................... 28

26. Fore wing with one transverse dark band beneath pterostigma.......... A. pteromaculatum Xu, Olmi, Guglielmino & Chen

- Fore wing hyaline, without dark transverse bands........................................................... 27

27. Segment 5 of protarsus with large lamellae and distal part slightly shorter than segment 4 ( Plate 33F )...................................................................................................... A. liui Xu, Olmi & He

- Segment 5 of protarsus with small lamellae and distal part slightly longer than segment 4 ( Plate 47B )................................................................................................. A. tenuitarse Xu, Olmi & He

28. Segment 5 of protarsus with proximal and medial region provided of lamellae ( Plate 29C )............ A. hirashimai Olmi

- Segment 5 of protarsus with proximal and medial region provided of bristles, without lamellae ( Plates 18D , 42A )......................................... A. austini Olmi (Oriental region) and A. reticulatum Kieffer (Palaearctic) (see remarks)

29. Posterior surface of propodeum with median area as rugose as lateral areas...................................... 30

- Posterior surface of propodeum with median area at least partly smooth and not rugose............................. 37

30. Head with POL approximately as long as OOL, or shorter, or slightly longer..................................... 31

- Head with POL approximately twice as long as OOL........................................................ 36

31. Chela with arolium very large, approximately as long as enlarged claw ( Plate 50A ).................. A. wushense Olmi

- Chela with arolium much smaller than enlarged claw ( Plates 18B, C , 47A )....................................... 32

32. Segment 5 of protarsus with distal part much shorter than basal part ( Plates 18B , 47A )............................. 33

- Segment 5 of protarsus with distal part approximately as long as basal part, or slightly shorter, or slightly longer ( Plates 18C , 38C , 44B ).......................................................................................... 34

33. Notauli short, reaching approximately 0.15 length of scutum......................... A. tengchongense Xu, He & Olmi

- Notauli long, reaching approximately 0.4 length of scutum................................. A. amritodiphagum Olmi

34. with anterior half of face strongly rugose and with remaining surface partly rugose and partly strongly punctate and unsculptured among punctures...................................................................... A. atrum Olmi

- Head almost completely smooth, punctate, unsculptured among punctures; occasionally vertex slightly rugose.......... 35

35. Head, including vertex, completely punctate and unsculptured among punctures..................... A. nepalense Olmi

- Head punctate and unsculptured among punctures, except vertex sculptured by irregular keels and partly reticulate rugose......................................................................................... A. silvicolum Olmi

36. Notauli short, approximately 0.3 length of scutum........................................... A. viraktamathi Olmi

- Notauli long, approximately 0.6 length of scutum............................................. A. yoshimotoi Olmi

37. Notauli complete or almost complete.......................................................... A. sinicum Olmi

- Notauli incomplete, reaching approximately 0.3–0.7 length of scutum.......................................... 38

38. Head punctate, unsculptured among punctures............................................................. 39

- Head granulated, or punctate and with more or less numerous irregular keels or areolae; occasionally head slightly granulated or partly reticulate rugose.............................................................................. 41

39. Segment 5 of protarsus without distinct basal and distal parts ( Plate 34A )..................... A. lochmocolum He & Xu

- Segment 5 of protarsus with distinct basal and distal parts( Plates 27E , 30B )...................................... 40

40. Segment 5 of protarsus with basal part much longer than distal part ( Plate 30B )........................ A. indicum Olmi

- Segment 5 of protarsus with basal part slightly shorter than distal part ( Plate 27C )........ A. gaoligongense Xu, Olmi & He

41. Frontal line absent; head smooth, completely granulated.................................. A. brachycerum (Dalman)

- Frontal line present; head at least partly rugose............................................................. 42

42. Forewing with one large brown transverse band on distal third................................ A. zoroastrum sp. nov.

- Forewing hyaline, without dark transverse bands........................................................... 43

43. Segment 5 of protarsus with inner margin straight ( Plate 33F ); notauli reaching about 0.65 length of scutum................................................................................................. A. liui Xu, Olmi & He

- Segment 5 of protarsus with inner margin not straight ( Plates 18B , 37C ); notauli shorter, reaching about 0.3–0.5 length of scutum............................................................................................ 44

44. Notauli reaching approximately 0.3 length of scutum........................................... A. mysorense Olmi

- Notauli reaching approximately 0.4–0.5 length of scutum.................................. A. amritodiphagum Olmi

45. Posterior surface of propodeum without longitudinal keels.................................................... 46

- Posterior surface of propodeum with two longitudinal keels................................................... 58

46. Enlarged claw much longer than segment 5 of protarsus ( Plate 41E ).................................. A. raptor Olmi

- Enlarged claw approximately as long as segment 5 of protarsus, or slightly shorter ( Plates 17C , 32E , 51A, B )........... 47

47. Segment 5 of protarsus laterally expanded and forming wide lamina ( Plate 32E ).................... A. laminatum Olmi

- Segment 5 of protarsus laterally not expanded ( Plates 17C , 43A , 51A, B )........................................ 48

48. Head granulated, not rugose, without areolae and keels (at most with frontal line and two frontal lateral keels along orbits)....................................................................................... A. yasumatsui Olmi

- Head punctate and unsculptured among punctures, or reticulate rugose, or more or less rugose and granulated or not granulated among keels........................................................................................ 49

49. Pronotum raised into transverse carina between anterior and posterior half ( Plate 17B )............... A. achterbergi Olmi

- Pronotum not raised into a transverse carina between anterior and posterior half................................... 50

50. Segment 1 of protarsus approximately as long as, or longer than segment 4...................................... 51

- Segment 1 of protarsus much shorter than segment 4........................................................ 56

51. Notauli long, almost reaching posterior margin of scutum; head mostly rugose........................ A. sarawaki Olmi

- Notauli short, reaching at most 0.5 length of scutum; head almost completely smooth, or completely or partly reticulate rugose.................................................................................................. 52

52. Head with OPL longer than POL; notauli short, at most reaching 0.2 length of scutum.................. A. vistosum Olmi

- Head with OPL shorter or longer than POL; notauli long, reaching 0.3–0.5 length of scutum......................... 53

53. Segment 5 of protarsus with basal part much longer than distal part ( Plate 35A ).................................................................................................. A. mangshanense Xu, Olmi, Guglielmino & Chen

- Segment 5 of protarsus with basal part much shorter than distal part ( Plates 23B , 50C )............................. 54

54. Scutum almost completely reticulate rugose, with anterior fourth unsculptured....................... A. zoilum sp. nov.

- Scutum punctate and unsculptured among punctures, or weakly granulated...................................... 55

55. Head granulated, with numerous irregular keels and areolae................................ A. yangi Xu, He & Olmi

- Head punctate and unsculptured among punctures, with numerous areolae near clypeus and inner margin of eyes.................................................................................................... A. choui Olmi

56. Head smooth, finely punctate, unsculptured among punctures, without keels, except frontal line and tracks of lateral keels near orbits.................................................................................. A. malaisei Olmi

- Head completely or almost completely reticulate rugose..................................................... 57

57. Notauli short, faintly visible near anterior margin of scutum; head not reticulate rugose on sides of ocellar triangle.............................................................................................. A. bengalense Olmi

- Notauli long, reaching approximately 0.5 length of scutum; head completely reticulate rugose.......... A. krombeini Olmi

58. Head smooth, more or less punctate, unsculptured among punctures, not reticulate rugose........................... 59

- Head completely or partly reticulate rugose............................................................... 75

59. Segment 4 of protarsus approximately twice as long as segment 1.............................................. 60

- Segment 4 of protarsus as long as, or less than twice as long as segment 1....................................... 61

60. Posterior surface of pronotum strongly punctate........................................ A. punctum Xu, He & Olmi

- Posterior surface of pronotum finely punctate.................................................... A. hilare Olmi

61. Mesosoma mostly or totally testaceous................................................................... 62

- Mesosoma mostly or totally black....................................................................... 64

62. Fore wing with two dark transverse bands; hindwing with distal third darkened................ A. wengae Xu, Olmi & He

- Fore wing and hind wing hyaline, without dark transverse bands............................................... 63

63. Head almost totally testaceous, partly darkened; notauli reaching about 0.9 length of scutum.............. A. quatei Olmi

- Head black, except mandible, clypeus and anterior region of face between antennal toruli brown-testaceous; notauli reaching about 0.25 length of scutum............................................................. A. zonarium sp. nov.

64. Head with OL much longer than POL....................................................... A. lankanum Olmi

- Head with OL as long as, or shorter than POL............................................................. 65

65. Head with OPL twice or more than twice as long as greatest breadth of posterior ocelli............................. 66

- Head with OPL less than twice as long as greatest breadth of posterior ocelli..................................... 71

66. Mesosoma completely black (occasionally with posterior margin of pronotum brown).............................. 67

- Mesosoma partly testaceous............................................................................ 70

67. Posterior surface of propodeum with median region smooth, shiny, punctate and unsculptured among punctures..................................................................................................... A. kresli Olmi

- Posterior surface of propodeum with median region dull and rugose............................................ 68

68. Segment 5 of protarsus with very large lamellae ( Plate 26D )........................................ A. fidum Olmi

- Segment 5 of protarsus with small lamellae ( Plates 19F , 22A )................................................. 69

69. Segment 5 of protarsus very broad ( Plate 22A )............................................. A. caii Xu, He & Olmi

- Segment 5 of protarsus very slender ( Plate 19F )..................................... A. blanduscutum Xu, He & Rui

70. Head and prothorax totally testaceous.............................................. A. multicolor Xu, He & Olmi

- Head black (with mandible testaceous); prothorax totally testaceous.................... A. chonganense Xu, He & Olmi

71. Head totally testaceous, or black, with mandible, clypeus, genae and anterior half of face testaceous.................. 72

- Head black, with only mandible testaceous................................................................ 73

72. Notauli reaching about 0.5 length of scutum............................................... A. bui Xu, Olmi & He

- Notauli reaching about 0.65–0.80 length of scutum............................................... A. hilare Olmi

73. Posterior surface of pronotum shorter than half of scutum.......................................... A. fidum Olmi

- Posterior surface of pronotum longer than half of scutum..................................................... 74

74. Segment 5 of protarsus with distal apex rounded and distinct from remaining part of segment ( Plate 19F )............................................................................................ A. blanduscutum Xu, He & Rui

- Segment 5 of protarsus with inner margin almost straight ( Plate 40D )............................. A. provinciale Olmi

75. Fore wing hyaline, without dark transverse bands........................................................... 76

- Fore wing with 1–2 dark transverse bands................................................................. 99

76. Head completely reticulate rugose....................................................................... 77

- Head partly reticulate rugose and partly smooth............................................................ 82

77. Body totally testaceous.................................................................... A. peculiare Olmi

- Body almost totally black.............................................................................. 78

78. Segment 5 of protarsus with basal part about as long as, or much longer than distal part ( Plates 22B , 30A , 46A )......... 79

- Segment 5 of protarsus with basal part much shorter than distal part ( Plates 36D , 51E )............................. 81

79. Segment 5 of protarsus with more than 50 distal lamellae ( Plate 30A )........ A. hunanense Xu, Olmi, Guglielmino & Chen

- Segment 5 of protarsus with less than ten distal lamellae ( Plates 22B , 46A )...................................... 80

80. Segment 5 of protarsus with basal part about as long as distal part ( Plate 22B ); face with two strong lateral keels located near orbits and directed towards antennal toruli....................................................... A. cheni Olmi

- Segment 5 of protarsus with basal part much longer than distal part ( Plate 46A ); face without lateral keels located near orbits and directed towards antennal toruli........................................................ A. superbum Olmi

81. Notauli reaching about 0.65 length of scutum; segment 5 of protarsus with two rows of lamellae, one of rows composed of lamellae slightly longer than others ( Plate 36D )................................................... A. muiri Olmi

- Notauli reaching about 0.8 length of scutum; segment 5 of protarsus with two–three rows of lamellae, one of rows composed of lamellae much longer thghan others ( Plate 51E )...................................... A. zhangae Xu, Olmi & He

82. Segment 5 of protarsus with basal region approximately as long as distal region, or slightly shorter, or slightly longer ( Plates 17G , 19D , 28C , 31A )................................................................................. 83

- Segment 5 of protarsus with basal region much longer or much shorter than distal region ( Plates 24C , 27A )............ 87

83. Posterior surface of propodeum with median area shiny, smooth, almost completely unsculptured..................... 84

- Posterior surface of propodeum with median area dull and rugose.............................................. 85

84. Vertex without two short oblique keels connecting posterior ocelli to occipital carina; notauli reaching approximately 0.8–0.9 length of scutum; segment 1 of protarsus more than 1.5 times as long as segment 4..................... A. becvari Olmi

- Vertex behind ocelli with two short oblique keels connecting posterior ocelli to occipital carina; notauli reaching approximately 0.7 length of scutum; segment 1 of protarsus slightly longer than segment 4...................... A. heveli Olmi

85. Segment 4 of protarsus as long as segment 1................................................... A. insertum Olmi

- Segment 4 of protarsus shorter than segment 1............................................................. 86

86. Posterior surface of pronotum shorter than half of scutum; head with OPL much shorter than OOL........... A. acre Olmi

- Posterior surface of pronotum longer than half of scutum; head with OPL longer than OOL...... A. longum Xu, Olmi & He

87. Segment 5 of protarsus with basal part much longer than distal part ( Plate 24C )...................... A. confusum Olmi

- Segment 5 of protarsus with basal part much shorter than distal part ( Plates 18A , 27A )............................. 88

88. Posterior surface of propodeum with median area smooth and unsculptured...................................... 89

- Posterior surface of propodeum with median area as rugose as lateral areas...................................... 91

89. Posterior surface of pronotum transverse, shorter than half of scutum...................... A. funiuense Xu, He & Olmi

- Posterior surface of pronotum longer than half of scutum..................................................... 90

90. Head with face completely reticulate rugose.......................................... A. actuosum Xu, Olmi & He

- Head with anterior half of face reticulate rugose and posterior half punctate and unsculptured among punctures................................................................................................... A. munitum Olmi

91. Posterior surface of pronotum much longer than anterior surface............................................... 92

- Posterior surface of pronotum about as long as anterior surface................................................ 97

92. Ocelli large, so that OPL is shorter than greatest breadth of posterior ocelli; mesosoma black, with some regions, including ventral surface, testaceous; head testaceous, with brown ocellar spot..................... A. yunnanense Xu, Olmi & He

- Ocelli small, so that OPL is much longer than greatest breadth of posterior ocelli.................................. 93

93. Mesosoma black, at most with lateral margins of pronotum slightly testaceous.................................... 94

- Mesosoma almost completely testaceous-ferruginous; head testaceous-ferruginous................................ 96

94. Head black, except mandible, clypeus and anterior third of face testaceous............................ A. takenoi Olmi

- Head black, except mandible testaceous.................................................................. 95

95. Distal part of segment 5 of protarsus forming obtuse angle with rest of segment ( Plate 37E )............ A. naduense Olmi

- Distal part of segment 5 of protarsus forming right angle with rest of segment ( Plate 140B )............. A. tadauchii Mita

96. Notauli incomplete, reaching about 0.7 length of scutum................................... A. tongi Xu, Olmi & He

- Notauli complete, posteriorly separated.................................................. A. phuphayonense Olmi

97. Notauli almost complete.................................................................... A. flaccum Olmi

- Notauli reaching approximately 0.6 length of scutum........................................................ 98

98. Face punctate, unsculptured among punctures; posterior surface of pronotum about as long as scutum.................................................................................................... A. wangi Xu, He & Olmi

- Face rugose, with posterior third punctate, unsculptured among punctures; posterior surface of pronotum about 0.5 as long as scutum............................................................ A. semirugosum Xu, Olmi & Guglielmino

99. Temples absent......................................................................... A. laotianum Olmi

- Temples distinct......................................................................... A. fyanense Olmi

Remarks. The females of A. austini Olmi (Oriental region) and A. reticulatum Kieffer (Palaearctic) are not separable on the basis of the present knowledge.

Males (unknown in the species not present in the key to the males)

1. Posterior surface of propodeum without longitudinal keels..................................................... 2

- Posterior surface of propodeum with two complete longitudinal keels; occasionally longitudinal keels incomplete, at least as long as half of posterior surface of propodeum............................................................. 15

2. Antennae pectinate................................................................... A. ctenoides Xu & He

- Antenna filiform...................................................................................... 3

3. Paramere with distal inner pointed or rounded process ( Plates 17D , 25B , 38A, B ).................................. 4

- Paramere without distal inner process ( Plates 21C , 31E , 42G ).................................................. 6

4. Scutum completely reticulate rugose and granulated.............................................. A. devriesi Olmi

- Scutum punctate, unsculptured among punctures, occasionally partly very slightly granulated......................... 5

5. Head granulated and completely reticulate rugose; distal inner process of paramere without bristles near distal apex ( Plate 38A, B )................................................................................... A. nemorale Olmi

- Head sculptured by numerous irregular keels, unsculptured among keels; distal inner process of paramere with many bristles near distal apex ( Plate 17D ).............................................................. A. achterbergi Olmi

6. Scutum completely reticulate rugose, or completely sculptured by irregular keels; notauli absent...................... 7

- Scutum unsculptured, or punctate, or granulated, not completely reticulate rugose, not completely sculptured by irregular keels; notauli present.................................................................................. 9

7. Scutellum reticulate rugose................................................................ A. silvestre Olmi

- Scutellum unsculptured................................................................................ 8

8. Head and scutum granulated and sculptured by numerous irregular keels; POL more than three times as long as OPL............................................................................................. A. cerberum Olmi

- Head and scutum completely reticulate rugose; POL less than twice as long as OPL................ A. khaokhoense Olmi

9. Scutum granulated................................................................................... 10

- Scutum punctate, or unsculptured, not granulated; occasionally scutum partly reticulate rugose....................... 11

10. Paramere much shorter than penis ( Plate 20C )............................................... A. borneanum Olmi

- Paramere about as long as penis ( Plate 31C, D )........................................... A. jurineanum Latreille

11. Head punctate, unsculptured among punctures................................................. A. insertum Olmi

- Head granulated, or rugose, or with irregular keels.......................................................... 12

12. Head dull, smooth, granulated............................................................ A. yasumatsui Olmi

- Head shiny, rugose, with areolae and irregular keels......................................................... 13

13. Notauli almost reaching posterior margin of scutum............................................. A. sarawaki Olmi

- Notauli reaching at most 0.5 length of scutum.............................................................. 14

14. Notauli reaching approximately 0.5 length of scutum........................................................................................... A. krombeini Olmi (Oriental) and A. jurineanum Latreille (Palaearctic) (see remarks)

- Notauli reaching approximately 0.3–0.4 length of scutum............................................................................................ A. thai Olmi (Oriental) and A. jurineanum Latreille (Palaearctic) (see remarks)

15. Posterior surface of propodeum with median area smooth, almost completely not rugose............................ 16

- Posterior surface of propodeum with median area as rugose as lateral areas...................................... 50

16. Paramere with distal inner pointed or rounded process ( Plates 19C , 41D , 45F , 46D )................................ 17

- Paramere without distal inner process ( Plates 25E , 30E , 38D )................................................. 31

17. Notauli reaching approximately 0.6–0.8 length of scutum.................................................... 18

- Notauli reaching approximately 0.3–0.5 length of scutum.................................................... 24

18. Paramere with distal inner process small and rounded ( Plate 41D ).................................... A. quatei Olmi

- Paramere with distal inner process large and usually pointed ( Plates 19C , 45F , 46D )............................... 19

19. Paramere with two distal processes ( Plate 19C ); distal inner processes of paramere less pointed ( Plate 19C )................................................................................................ A. biprocessum Xu & He

- Paramere with only one distal process ( Plates 21D , 45F , 46D ); distal inner process of paramere more pointed ( Plates 21D , 45F , 46D ).............................................................................................. 20

20. Head with POL approximately as long as OOL............................................. A. sulawesianum Olmi

- Head with POL shorter than OOL....................................................................... 21

21. Head with two keels connecting posterior ocelli to occipital carina............................................. 22

- Head without two keels connecting posterior ocelli to occipital carina........................................... 23

22. Distal inner pointed process of paramere narrower ( Plate 46D ).................................. A. taiwanense Olmi

- Distal inner pointed process of paramere broader ( Plate 21D ).................................... A. chaoi Xu & He

23. Frontal line absent.................................................................. A. sylvaticum Xu & He

- Frontal line present in front of anterior ocellus.............................................. A. cacumen Xu & He

24. Frontal line absent; head smooth, completely granulated.................................. A. brachycerum (Dalman)

- Frontal line present, complete or incomplete; rarely frontal line absent, then head punctate or rugose, never smooth and completely granulated.................................................................................... 25

25. Distal inner process of paramere medially situated ( Plates 32D , 37D )........................................... 26

- Distal inner process of paramere distally situated ( Plates 29D, F , 33E , 35C )...................................... 27

26. Frontal line absent; head with POL shorter than OOL...................................... A. laeve Xu, He & Olmi

- Frontal line complete; head with POL longer than OOL......................................... A. mysorense Olmi

27. Distal inner process of paramere rounded ( Plates 33E , 35C ).................................................. 28

- Distal inner process of paramere pointed ( Plates 29D , 33C , 41B ).............................................. 29

28. Distal inner process of paramere large and with inner ventral margin serrate ( Plate 33E ); head black, except mandible, clypeus and narrow stripe between and on sides of antennal toruli testaceous-reddish............... A. maculatum Xu, He & Olmi

- Distal inner process of paramere small and without inner ventral serrate margin ( Plate 35C ); head black, with only mandible testaceous; clypeus and face black....................................................... A. meifenganum Olmi

29. Head dull, completely reticulate rugose........................................................ A. huangi Olmi

- Head partly or completely punctate, unsculptured among punctures, not reticulate rugose........................... 30

30. Head completely punctate, unsculptured among punctures............................................ A. lini Olmi

- Face with anterior half granulated and posterior half punctate and unsculptured among punctures.... A. qinlingense He & Xu

31. Distivolsella with two lateral processes ( Plate 25E ).............................................. A. doiense Olmi

- Distivolsella simple ( Plate 30E , 38D , 50D )................................................................ 32

32. Distal part of stigmal vein as long as proximal part, or slightly shorter.......................................... 33

- Distal part of stigmal vein much shorter than proximal part................................................... 34

33. Head punctate, unsculptured among punctures........................................ A. isoneuron Xu, He & Olmi

- Head completely reticulate rugose....................................... A. ningshanense Xu, Olmi & Guglielmino

34. Notauli very short, hardly visible near anterior margin of scutum.................................... A. yasiri Olmi

- Notauli reaching at least 0.4 length of scutum.............................................................. 35

35. Notauli reaching approximately 0.40–0.55 length of scutum.................................................. 36

- Notauli reaching at least 0.6 length of scutum.............................................................. 41

36. Head completely or almost completely reticulate rugose..................................................... 37

- Head completely or almost completely punctate, not reticulate rugose........................................... 39

37. Paramere without proximal membranous processes, or with two very short processes ( Plate 26A, B )..... A. expolitum Olmi

- Paramere with two long proximal membranous processes ( Plates 23C, D , 38H )................................... 38

38. Paramere with inner band covered with papillae ( Plate 38H ).............................. A. papillum Xu, He & Olmi

- Paramere without inner band covered with papillae ( Plates 23C, D )................................ A. chui Xu & He

39. Paramere much shorter than penis ( Plate 19A, B ).............................................. A. autumnale Olmi

- Paramere approximately as long as penis ( Plate 39B )........................................................ 40

40. Face with two lateral keels along orbits directed towards antennal toruli............................ A. ingenuum Olmi

- Face without two lateral keels along orbits directed towards antennal toruli.......................... A. peterseni Olmi

41. Paramere approximately as long as penis ( Plates 38H , 42F , 45A, B )............................................ 42

- Paramere much shorter than penis ( Plates 25A , 36C )........................................................ 49

42. Head completely reticulate rugose................................................... A. papillum Xu, He & Olmi

- Head not or partly reticulate rugose...................................................................... 43

43. Head with vertex reticulate rugose; rest of head punctate and unsculptured among punctures.......... A. semipolitum Olmi

- Head completely punctate and unsculptured among punctures................................................. 44

44. Proximal membranous process of paramere not longer than volsellae ( Plates 40A , 45C , 50F )........................ 45

- Proximal membranous process of paramere longer than volsellae ( Plates 37B , 42E )................................ 48

45. Notauli reaching approximately 0.7–0.8 length of scutum.................................................... 46

- Notauli reaching approximately 0.6 length of scutum........................................................ 47

46. Proximal membranous process of paramere much shorter than volsellae, with margins hairless ( Plate 45A )......................................................................................... A. songyangense Xu, He & Olmi

- Proximal membranous process of paramere slightly shorter than volsellae, with inner margin provided of numerous hairs ( Plate 40A )..................................................................... A. pilosum Xu, Olmi & He

47. Inner side of paramere with mosaic drawing ( Plate 45C ).................................. A. striolaforceps Xu & He

- Inner side of paramere without mosaic drawing ( Plate 50F )................................. A. yuani Xu, He & Olmi

48. Inner side of distal apex of paramere without papillae ( Plate 37B ); proximal region of paramere without inner expansion showing mosaic drawing ( Plate 37B )............................................................. A. munitum Olmi

- Inner side of distal apex of paramere with numerous papillae ( Plate 42E ); proximal region of paramere with inner expansion showing mosaic drawing ( Plate 42E )........................................................ A. sabahnum Olmi

49. Paramere without proximal membranous process ( Plate 25A ); POL approximately as long as OOL......... A. debile Olmi

- Paramere with proximal membranous process ( Plate 36C ); POL much shorter than OOL.......... A. montanum Xu & He

50. Head punctate or granulated, not reticulate rugose.......................................................... 51

- Head completely or partly reticulate rugose............................................................... 61

51. Head completely or partly granulated.................................................................... 52

- Head completely punctate, not granulated................................................................. 54

52. Paramere with pointed distal inner process ( Plate 36B )........................................... A. mixtum Olmi

- Paramere without distal inner process ( Plates 17A , 33D )..................................................... 53

53. Paramere approximately as long as penis ( Plate 33D )........................................... A. luzonense Olmi

- Paramere much shorter than penis ( Plate 17A )............................................... A. abatanense Olmi

54. Paramere without distal inner process ( Plates 17F , 30D , 31C, D , 35B ).......................................... 55

- Paramere with pointed or rounded distal inner process ( Plates 26C, E , 38I , 40C ).................................. 58

55. Inner side of paramere sculptured by numerous papillae ( Plate 35B )................................ A. martiale Olmi

- Inner side of paramere not sculptured by papillae ( Plates 17F , 30D , 31C, D )...................................... 56

56. Notauli reaching approximately 0.6–0.8 length of scutum.......................................... A. hilare Olmi

- Notauli shorter, reaching less than 0.6 length of scutum...................................................... 57

57. Head black, except mandible testaceous; head usually with POL as long as, or longer than OOL; distal part of stigmal vein much shorter than proximal part............................................................ A. insertum Olmi

- Head black, except mandible, clypeus and anterior margin of face testaceous; head with POL shorter than OOL; distal part of stigmal vein slightly shorter than proximal part......................................... A. amabile Xu, Olmi & He

58. Head with face partly yellow-whitish and partly black................................. A. flavoniger Xu, He & Olmi

- Head totally black, or brown, or brown-reddish; at most with mandible testaceous, rarely with clypeus testaceous........ 59

59. Paramere with distal inner rounded process ( Plate 38I )........................................ A. parapriscum Olmi

- Paramere with distal inner process more or less pointed ( Plates 26C , 40B, C )..................................... 60

60. Distal inner process of paramere with inner distal margin slightly excavated ( Plate 26C ).................. A. fidum Olmi

- Distal inner process of paramere with inner distal margin not excavated ( Plate 40B, C ).................. A. priscum Olmi

61. Paramere much shorter than penis ( Plate 48A )..................................................... A. thai Olmi

- Paramere approximately as long as penis ( Plates 38G , 42F , 46E , 48C )........................................... 62

62. Head mostly punctate and unsculptured among punctures, rugose only near orbits or behind posterior ocelli............ 63

- Head completely reticulate rugose, or with face sculptured by numerous longitudinal or irregular keels and rest of surface reticulate rugose; occasionally face with smooth and shiny area in front of anterior ocellus, rest of head reticulate rugose...... 66

63. Inner side of paramere with numerous papillae ( Plate 38G )..................................... A. papillatum Olmi

- Inner side of paramere without papillae ( Plates 42F , 46E , 48C )................................................ 64

64. Frontal line incomplete, only shortly present in front of anterior ocellus.......................... A. semipolitum Olmi

- Frontal line complete................................................................................. 65

65. Head with OOL about 0.5 as long as POL.................................................. A. viraktamathi Olmi

- Head with OOL about as long as POL, or slightly longer than POL.................................. A. takenoi Olmi

66. Paramere with distal inner rounded or pointed process ( Plates 17E , 28B , 29D, E , 36A , 38F )......................... 67

- Paramere without distal inner rounded or pointed process ( Plates 38E , 50D )...................................... 71

67. Distal inner process of paramere pointed ( Plate 29D – F )........................................... A. huangi Olmi

- Distal inner process of paramere rounded ( Plates 17E , 28B , 36A , 38F ).......................................... 68

68. Scutum rugose.......................................................................... A. heppneri Olmi

- Scutum punctate, unsculptured among punctures, not rugose.................................................. 69

69. Head with OOL shorter than OL............................................................ A. adebratti Olmi

- Head with OOL much longer than OL.................................................................... 70

70. Notauli reaching approximately 0.6–0.7 length of scutum............................................ A. mite Olmi

- Notauli reaching approximately 0.4 length of scutum...................................... A. pangi Xu, He & Olmi

71. Scutum mostly reticulate rugose or granulated and sculptured by many irregular keels or areolae...... A. pahanganum Olmi

- Scutum punctate, unsculptured among punctures........................................................... 72

72. Head with anterior half of face sculptured by numerous longitudinal keels; rest of face reticulate rugose; notauli very short, only shortly visible near anterior margin of scutum................................................ A. yasiri Olmi

- Head with face completely reticulate rugose, at most with smooth and shiny area in front of anterior ocellus; notauli reaching 0.3–0.5 length of scutum.............................................................................. 73

73. Head with OOL more than twice as long as OPL................................................. A. gauldi Olmi

- Head with OOL as long as, or less than twice as long as OPL................................................. 74

74. Inner side of paramere with numerous papillae ( Plate 39C )................................. A. phetchabunense Olmi

- Inner side of paramere without papillae ( Plates 25C , 27F, G , 42B ).............................................. 75

75. Distal apex of paramere slightly curved towards penis forming a very small rounded distal process ( Plate 42B )............................................................................................... A. reticulatum Kieffer

- Distal apex of paramere not curved towards penis, not forming very small rounded distal process ( Plates 25C , 27F, G ).... 76

76. Paramere without membranous process ( Plate 27F, G )................................ A. guangxiense Xu, He & Olmi

- Paramere with membranous process ( Plates 25C , 27G )...................................................... 77

77. Head with OOL much longer than OPL............................................. A. henanense Xu, He & Olmi

- Head with OOL about as long as OPL............................ A. diaoluoshanense Xu, Olmi, Guglielmino & Chen

Remarks. The male of Anteon jurineanum Latreille is very similar to those of A. borneanum Olmi , A. krombeini Ponomarenko and A. thai Olmi. It can be mistook for them, although the females of these four species are very different. However, A. jurineanum is a Palaearctic species present in the Oriental region only in transition areas between the two zoogeographical regions ( Nepal, Sichuan of China).

Figures

PLATE 17. A, D–F. Male genitalia. B. Pronotum and scutum. C, G. Chelae, holotypes. A. Anteon abatanense, holotype (from Olmi 1991). B. A. achterbergi, female from UAE, dorsal view. C. A. achterbergi (from Olmi 1991). D. A. achterbergi from UAE (from Olmi 2008a). E. A. adebratti, holotype (from Olmi 1993d). F. A. amabile, holotype (from Xu et al. 2006a). G. A. acre (from Olmi 1991). Scale bar 0.11 mm for A, 0.17 mm for C and G, 0.14 mm for D and E, 0.19 mm for F.

PLATE 18. Chelae, holotypes. A. Anteon actuosum (from Xu et al. 2006b). B. A. amritodiphagum (from Olmi 1991). C. A. atrum (from Olmi 1998e). D. A. austini (from Olmi 1991). Scale bar 0.22 mm for A, 0.21 mm for B and C and 0.18 mm for D.

PLATE 19. A–C. Male genitalia. D–F. Chelae. A. Anteon autumnale, holotype (from Olmi 1991). B. A. autumnale from China. C. A. biprocessum, holotype (from Xu & He, 1997b). D. A. becvari, holotype (from Olmi 2000b). E. A. bengalense, holotype (from Olmi 1984). F. A. blanduscutum, holotype (from Xu et al. 1996). Scale bar 0.20 mm for A, B and C, 0.26 mm for D, 0.30 mm for E, 0.10 mm for F.

PLATE 20. A, B, D. Chelae. C, E. Male genitalia. A. Anteon borneanum, holotype (from Olmi 1984). B. A. flaccum, holotype (=borneanum) (from Olmi 1992a). C. A. borneanum from the Philippines (from Olmi 1991). D, E. A. brachycerum from Ireland (D) and UK (E) (from Olmi 1984). Scale bar 0.14 mm for A, 0.20 mm for B, 0.12 mm for C and 0.37 mm for D and E.

PLATE 21. A. Chela, holotype. B–D. Male genitalia, holotypes. A. Anteon bui (from Xu et al. 2003). B. A. cacumen (from Xu & He, 1997b). C. A. cerberum (from Olmi 1992a). D. A. chaoi (from Xu & He, 1997b). Scale bar 0.14 mm for A, 0.18 mm for B, 0.16 mm for C and 0.17 mm for D.

PLATE 22. Chelae, holotypes. A. Anteon caii (from Xu et al. 2001a). B. A. cheni (from Olmi 1996c). Scale bar 0.22mm for A and 0.10 mm for B.

PLATE 23. A, B. Chelae, holotypes. C, D. Male genitalia. A. Anteon chonganense (from He & Xu, 2002). B. A. choui, holotype (from Olmi 1996c). C. A. chui, holotype (from Xu & He, 1998a). D. A. chui from China, Nanling. Scale bar 0.17 mm for A, 0.26 mm for B, 0.28 mm for C and D.

PLATE 24. A–C. Chelae, holotypes. D. Antenna. E. Male genitalia. A. Anteon clavatum (from Olmi & Currado, 1979a). B. A. confertilaminarum (from Xu et al. 1998). C. A. confusum (from Olmi 1991). D, E. A. ctenoides, holotype (from Xu & He, 1997a). Scale bar 0.43 mm for A, 0.22 mm for B, 0.17 mm for C and 0.18 mm for E.

PLATE 25. A–C, E. Male genitalia, holotypes. D, F. Chelae, holotypes. A. Anteon debile (from Olmi 1984). B. A. devriesi (from Olmi 1998e). C. A. diaoluoshanense (from Xu et al. 2011e). D. A. dignum (from Olmi 1991). E. A. doiense (from Olmi 2008c). F. A. dravidicum (from Olmi 2010). Scale bar 0.07 mm for A, 0.09 mm for B, 0.21 mm for C and D, 0.23 mm for E and 0.19 mm for F.

PLATE 26. A–C, E. Male genitalia. D, F. Chelae, holotypes. A. Anteon expolitum, holotype (from Olmi 1984). B. A. expolitum from Indonesia. C. A. fidum from China, Xizang (from Xu et al. 2006a). D. A. fidum (from Olmi 1991). E. A. flavoniger, holotype (from Xu et al. 1998). F. A. flaccum (from Olmi 1987a). Scale bar 0.18 mm for A and B, 0.07 mm for C, 0.13 mm for D, 0.15 mm for E and 0.31 mm for F.

PLATE 27. A–C. Chelae, holotypes. D–G. Male genitalia. A. Anteon funiuense (from Xu et al. 2001a). B. A. fyanense (from Olmi 1984). C. A. gaoligongense (from Xu et al. 2009a). D. A. gauldi, holotype (from Olmi 1987c). E. A. gauldi from Indonesia. F. A. guangxiense, holotype (from Xu et al. 1998). G. A. henanense, holotype (from Xu et al. 2001a). Scale bar 0.41 mm for A and B, 0.20 mm for C, 0.15 mm for D, 0.16 mm for E and G, 0.34 mm for F.

PLATE 28. A. Chela of Anteon heppneri, holotype (from Olmi 1992a). B. Male genitalia of A. heppneri, paratype (from Olmi 1992a). C. Chela of A. heveli, holotype (from Olmi 1998a). D. A. hilare (=A. javanum), Female. Length 2.5 mm. Scale bar 0.24 mm for A, 0.18 mm for B and 0.25 mm for C.

PLATE 29. A, C. Chelae, holotypes. B, D–F. Male genitalia. A. Anteon hilare (=Anteon javanum) (from Olmi 1984). B. A. hilare from Indonesia (from Olmi 1984). C. A. hirashimai (=A. tarsale Ponomarenko), holotype (from Olmi 1993b). D, E. A. huangi, paratype (D) and holotype (E) (from Olmi 1996c). F. A. huangi from China. Scale bar 0.18 mm for A, 0.12 mm for B, 0.38 mm for C, 0.13 mm for D and E, 0.15 mm for F.

PLATE 30. A, B. Chelae, holotypes. C–E. Male genitalia. A. Anteon hunanense (from Xu et al. 2012c). B. A. indicum (from Olmi 1984). C. A. ingenuum, holotype (from Olmi 1984). D. A. insertum, paratype (from Olmi 1991). E. A. isoneuron, holotype (from Xu et al. 1998). Scale bar 0.29 mm for A, 0.17 mm for B, 0.18 mm for C, 0.19 mm for D and 0.20 mm for E.

PLATE 31. A, B. Chelae. C–E. Male genitalia. A. Anteon insertum, holotype (from Olmi 1991). B. A. jurineanum (from Olmi 1984). C. A. jurineanum from UK, England (from Olmi 1984). D. A. jurineanum from UK, Scotland (from Olmi 1984). E. A. khaokhoense, holotype (from Olmi 2008c). Scale bar 0.18 mm for A, 0.16 mm for B and C, 0.15 mm for D and E.

PLATE 32. A, B, E, F. Chelae, holotypes. C, D. Male genitalia. A. Anteon kresli (from Olmi 2008b). B. A. krombeini (from Olmi 1984). C. A. krombeini from Sri Lanka (from Olmi 1984). D. A. laeve, holotype (from Xu et al. 1998). E. A. laminatum (from Olmi 1987a). F. A. lankanum (from Olmi 1984). Scale bar 0.32 mm for A, 0.34 mm for B, 0.33 mm for C, 0.23 mm for D, 0.26 mm for E and F.

PLATE 33. A, B, F. Chelae, holotypes. C–E. Male genitalia, holotypes. A. Anteon laotianum (from Olmi 1984). B. A. lesagei (from Olmi 1998e). C. A. lini (from Olmi 1996c). D. A. luzonense (from Olmi 1991). E. A. maculatum (from Xu et al. 2001a). F. A. liui (from Xu et al. 2010a). Scale bar 0.40 mm for A, 0.30 mm for B, 0.18 mm for C and D, 0.14 mm for E and 0.31 mm for F.

PLATE 34. Chelae, holotypes. A. Anteon lochmocolum (from He & Xu, 2002). B. A. longum (from Xu et al. 2011b). C. A. maai (from Olmi 1991). D. A. malaisei (from Olmi 1993d). Scale bar 0.32 mm for A, 0.21 mm for B, 0.15 mm for C and 0.18 mm for D.

PLATE 35. A, D, E. Chelae, holotypes. B, C. Male genitalia, holotypes. A. Anteon mangshanense (from Xu et al. 2012c). B. A. martiale (from Olmi 1992a). C. A. meifenganum (from Olmi 1991). D. A. medium (from Olmi 1998e). E. A. metuendum (from Olmi 1987a). Scale bar 0.32 mm for A, 0.36 mm for B, 0.26 mm for C, 0.24 mm for D and 0.16 mm for E.

PLATE 36. A–C. Male genitalia, holotypes. D, E. Chelae, holotypes. A. Anteon mite (from Olmi 1996c). B. A. mixtum (from Olmi 1996c). C. A. montanum (from Xu & He, 1994b). D. A. muiri (from Olmi 1984). E. A. multicolor (from Xu et al. 1998). Scale bar 0.17 mm for A, 0.11 mm for B, 0.22 mm for C, 0.24 mm for D and 0.13 mm for E.

PLATE 37. A, C, E, F. Chelae. B, D. Male genitalia. A, B. Anteon munitum (=A. bauense), female from China (A) (from Olmi 1991) and holotype (B) (from Olmi 1984). C, D. A. mysorense, holotype (C) and paratype (D) (from Olmi 1984). E. A. naduense, holotype (from Olmi 1987a). F. A. nanlingense, holotype (from Xu et al. 2011b). Scale bar 0.27 mm for A and B, 0.11 mm for C, 0.17 mm for D, 0.26 mm for E and 0.24 mm for F.

PLATE 38. A, B, D–I. Male genitalia, holotypes. C. Chela. A, B. Anteon nemorale (from Olmi 1984). C. A. nepalense, holotype (from Olmi 1984). D. A. ningshanense (from Xu et al. 2012a). E. A. pahanganum (from Olmi 1991). F. A. pangi (from Xu et al. 1998). G. A. papillatum (from Olmi 2000b). H. A. papillum (from He & Xu, 2002). I. A. parapriscum (from Olmi 1991). Scale bar 0.13 mm for A and B, 0.16 mm for C and I, 0.28 mm for D and H, 0.21 mm for E and G, 0.23 mm for F.

PLATE 39. A, D, E. Chelae, holotypes. B, C. Male genitalia. A. Anteon peculiare (from Olmi 1993d). B. A. peterseni from China, Hainan. C. A. phetchabunense, holotype (from Olmi 2008c). D. A. philippinum (from Olmi 1984). E. A. phuphayonense (from Olmi 2008c). Scale bar 0.16 mm for A, 0.15 mm for B and C, 0.21 mm for D and 0.13 mm for E.

PLATE 40. A–C. Male genitalia. D, E. Chelae, holotypes. A. Anteon pilosum, holotype (from Xu et al. 2010b). B. A. priscum, paratype (proximal membranous process removed) (from Olmi 1991). C. A. priscum from China, Ningxia. D. A. provinciale (from Olmi 1991). E. A. pteromaculatum (from Xu et al. 2011d). Scale bar 0.22 mm for A, 0.10 mm for B and C, 0.16 mm for D, 0.35 mm for E

PLATE 41. A, C, E. Chelae. B, D. Male genitalia. A. Anteon punctum, holotype (from He & Xu, 2002). B. A. qinlingense, holotype (from He & Xu, 2002). C. A. quatei from Malaya (from Olmi 1998e). D. Anteon quatei, paratype (from Olmi 1991). E. A. raptor, holotype (from Olmi 1998e). Scale bar 0.26 mm for A, 0.23 mm for B, 0.13 mm for C, 0.17 mm for D, 0.26 mm for E.

PLATE 42. A, C. Chelae, holotypes. B, D, E–G. Male genitalia. A. Anteon reticulatum (from Olmi 1984). B. A. reticulatum from Italy (from Olmi 1991). C. A. rugosum (from Xu & He, 1997d). D. A. sabahnum from Brunei. E. A. sabahnum, holotype (from Olmi 1998a). F. A. semipolitum, holotype (from Olmi 2008c). G.A. silvestre, holotype (from Olmi 1984). Scale bar 0.34 mm for A, 0.25 mm for B, 0.16 mm for C and F, 0.18 mm for D, 0.23 mm for E and 0.12 mm for G.

PLATE 43. A, B. Anteon sarawaki. A. Chela of holotype (from Olmi 1984). B. Male genitalia of paratype (from Olmi 1984). C. A. semipolitum, holotype. Length 2.62 mm. Scale bar 0.22 mm for A and 0.26 mm for B

PLATE 44. Chelae, holotypes. A. Anteon semirugosum (from Xu et al. 2012a). B. A. silvicolum (from Olmi 1984). C. A. sinicum (from Olmi 1996c). Scale bar 0.17 mm for A, 0.20 mm for B and 0.18 mm for C

PLATE 45. A–C, F, G. Male genitalia. D, E. Chelae. A. Anteon songyangense, holotype (from Xu et al. 1998). B. A. songyangense from China. C. A. striolaforceps from China. D. A. spenceri (from Olmi 1991). E. A. subdignum, holotype (from Olmi 1992a). F. A. sulawesianum, holotype (from Olmi 1991). G. A. sulawesianum from China. Scale bar 0.22 mm for A and B, 0.18 mm for C and D, 0.15 mm for E, 0.13 mm for F and G.

PLATE 46. A, B. Chelae, holotypes. C–E. Male genitalia. A. Anteon superbum (from Olmi 1984). B. A. takenoi (from Olmi 1995f). C. A. sylvaticum, holotype (from Xu & He, 1997b). D. A. taiwanense, holotype (from Olmi 1991). E. A. takenoi from China. Scale bar 0.12 mm for A, 0.19 mm for B, C and D, 0.20 mm for E.

PLATE 47 . Chelae , holotypes . A . Anteon tengchongense ( from Xu et al . 1998 ) . B . A . tenuitarse ( from Xu et al . 2010 b ) . C . A . thai ( from Olmi 1984 ) . Scale bar 0.17 mm for A , 0.19 mm for B and 0.28 mm for C .

PLATE 48. A–C. Male genitalia. D, E. Chelae, holotypes. A. Anteon thai from Thailand. B. A. thai from China. C. A. viraktamathi, paratype (from Olmi 1987c). D. A. tongi (from Xu et al. 2006b). E. A. viraktamathi (from Olmi 1987c). Scale bar 0.14 mm for A and B, 0.17 mm for C, 0.09 mm for D and 0.24 mm for E.

PLATE 50. A–C. Chelae, holotypes. D–F. Male genitalia. 147. A. Anteon wushense (from Olmi 1991). B. A. xuexini (from Xu et al. 2001a). C. A. yangi (from Xu et al. 1998). D. A. yasiri, holotype (from Olmi 1998e). E. A. yasumatsui from China. F. A. yuani, holotype (from Xu et al. 1998). Scale bar 0.34 mm for A, 0.19 mm for B, 0.12 mm for C, 0.13 mm for D, 0.18 mm for E and 0.22 mm for F.

PLATE 51. Chelae, holotypes. A. Anteon yasumatsui (from Olmi 1984). B. A. malaysianum (=A. yasumatsui) (from Olmi 1987c). C. A. yoshimotoi (from Olmi 1991). D. A. yunnanense (from Xu et al. 2006b). E. A. zhangae (from Xu et al. 2010a). Scale bar 0.14 mm for A and B, 0.18 mm for C, 0.15 mm for D and 0.16 mm for E.

PLATE 139. Chelae, holotypes. A. Anteon zonarium sp. nov. B. A. zopyrum sp. nov. C. A. zoroastrum sp. nov.. Scale bar 0.06 mm for A and B, 0.08 mm for C.

PLATE 140. Chelae, holotype. A. Anteon crassum. B. A. tadauchii (from Mita 2012). Scale bar 0.11 mm for A and B.