Tragardhus (Tragardhus) jani, Kamiński, 2017

Kamiński, Marcin Jan, 2017, Afrotropical Melambiina: revision of the genus Tragardhus Koch, 1956 and redescription of Pseudemmallus Koch, 1956 (Tenebrionidae: Pedinini), Zootaxa 4347 (1), pp. 71-93 : 84-85

Tragardhus (Tragardhus) jani sp. nov.

http://zoobANk.oRg/URN:lsiD:zoobANk.oRg:Act:2F8C2643-843D-4B99-8271-CDAFA2BD7C97 ( Figs 4C, F , 7 )

Studied type material. Holotype, femAle (TMNH): “S. AfR.: KWZ NAtAl / NtiNiNi NAt. Res. 1022m / 28.16S– 30.55E”, “ 30.11.2010; E-Y: 3887 / UNDeR stoNes & gRoUND / leg. RUth MülleR” ( Fig. 4C ).

Etymology. NAmeD AfteR JAN ANtoNi KAwecki (boRN oN MARch 20th, 2015), soN of mY fRieNDs AgNieszkA KRAssowskA-KAweckA AND TomAsz KAwecki.

Diagnosis. This species is similAR to T. (T.) majae sp. nov. bY shARiNg A commoN stRUctURe of elYtRA (thiRD elYtRAl iNteRvAl with A shARplY RAiseD coNtiNUoUs costA). Both species cAN be DistiNgUisheD bY the DiffeReNt stRUctURes of the pRoNotUm (ANteRioR pRoNotAl ANgles obtUse iN jani , pRotRUDiNg oUtwARDs iN majae ; lAteRAl siDes of pRoNotUm RoUNDeD iN bAsAl pARt iN jani , stRAight iN majae ) AND elYtRA (eighth iNteRvAl with A shARplY RAiseD coNtiNUoUs costA iN jani , composeD of shoRt tUbeRcles iN majae ).

Description. BoDY—leNgth cA. 5.5 mm. HAbitUs As iN Fig. 4C .

DoRsAl siDe of heAD shiNY, with coNspicUoUs pUNctURes (0.1 DiAmeteRs ApARt). FRoNtoclYpeAl sUtURe NoticeAble; with A pAiR of Deep DepRessioNs locAteD NeAR meDiAN pARt. CANthUs sUbpARAllel At bAse. ClYpeAl emARgiNAtioN Deep (clYpeAl emARgiNAtioN wiDth: Depth RAtio cA. 5.5). LAbRUm wiDe (wiDth: leNgth RAtio cA. 3.1); DelicAtelY emARgiNAte iN miDDle; posteRioR thiRD coveReD with DeNse AND coNspicUoUs setAe. MeNtUm wiDest iN miDDle; flAt, Not coveRiNg cARDo oR stipes; ApicAllY with shAllow meDiAN iNDeNtAtioN; lAteRAl lobes, iN veNtRAl view, coveReD bY miDDle pARt. SUbmeNtUm tRiANgUlAR. ApicAl segmeNt of mAxillARY pAlpUs tRApezoiDAl; with seNsoRY fielD occUpYiNg whole pRoximAl eDge. SecoND segmeNt of mAxillARY pAlpUs eloNgAteD (leNgth: wiDth RAtio cA. 3.8). PAlpifeR with pAtch of loNg setAe locAteD NeAR bAse; with iNteRNAl sUlcUs foR mAxillARY pAlpUs visible fRom veNtRAl view. ANteRioR teNtoRiAl pit ciRcUlAR. EYe DoRsAllY shielDeD bY loNgitUDiNAl keel. DoRsAl AND veNtRAl pARts of eYes composeD of similAR NUmbeR of fAcets; thRee fAcets pReseNt betweeN cANthUs AND temple. ANteNNA RelAtivelY shoRt (ANteNNA: pRoNotUm leNgth RAtio cA. 0.8); thiRD ANteNNomeRe shoRt (3RD: 2ND ANteNNomeRe leNgth RAtio cA. 2.0); fifth ANteNNomeRe shoRt (leNgth: wiDth RAtio cA. 0.7); ANteNNomeRes 8–10 tRANsveRse.

PRoNotUm NARRoweR thAN elYtRA (elYtRA: pRoNotUm wiDth RAtio cA. 1.1); RelAtivelY wiDe (leNgth: wiDth RAtio cA. 0.7); Disc shiNY, coveReD with coARse pUNctURes (0.1 DiAmeteRs ApARt); with two pAiRs of RoUNDeD cARiNAe. LAteRAl siDes of pRoNotAl Disc RoUNDeD; wiDest iN miDDle; withoUt ApophYseAl DepRessioNs. LAteRAl emARgiNAtioN of pRoNotUm NARRow AND complete; bAsAl emARgiNAtioN AbseNt; ANteRioR wiDelY iNteRRUpteD miDDle. BAse of pRoNotUm slightlY bisiNUAte. MiDDle of pRosteRNUm coveReD bY pRomiNeNt setAe. PRoNotAl hYpomeRoN shiNY, with coARse pUNctURes (1.0 DiAmeteRs ApARt). INteRcoxAl pRocess of pRosteRNUm DepResseD ApicAllY, howeveR with shARplY pRojectiNg tip. PRocoxAe NARRowlY sepARAteD (pRocoxA: iNteRcoxAl pRocess wiDth RAtio cA. 2.0).

ElYtRAl stRiAe Not impResseD, visible As teN Rows of coNspicUoUs pUNctURes oN eAch elYtRoN (siNgle DiAmeteR ApARt). INteRvAls 1, 3, 5, 7, 8, 9 iN foRm of shARplY RAiseD coNtiNUoUs costAe; 2, 4, 6, 10 flAt AND DUll, coveReD with micRoscUlptURe. ElYtRAl bAse v-like; Not emARgiNAte. EpipleURoN DilAteD ApicAllY, with RoUNDish AlveolA At level of foURth veNtRite; slightlY wiDeR thAN NeighboRiNg elYtRAl iNteRvAl; sitUAteD veNtRAllY, iNvisible iN DoRsAl Aspect. ElYtRAl hUmeRUs DeNtAte, slightlY pRotRUDiNg lAteRAD. ScUtellUm RelAtivelY smAll; ciRcUlAR. MetAthoRAcic wiNgs AbseNt.

MesoveNtRite withoUt pRocesses. MetAveNtRite RelAtivelY shoRt (metAcoxAl cAvitY: metAveNtRite (betweeN iNseRtioNs of meso- AND metAcoxAe) leNgth RAtio cA. 1.9). PRocess of fiRst AbDomiNAl veNtRite wiDe (DistANces betweeN mesocoxAe: metAcoxAe RAtio cA. 0.7). Fifth AbDomiNAl veNtRite withoUt sUbmARgiNAl sUlcUs; coveReD bY fiNe pUNctURes (siNgle DiAmeteR ApARt).

Meso- AND metAtRochANteRs NeARlY eNtiRelY coveReD bY Respective femoRA; eqUippeD with A siNgle setAe. MetAcoxA RoUNDeD (wiDth: leNgth RAtio cA. 1.1). FemAle legs simple (pRotibiAe eveNlY wiDeNeD towARD Apex). MoRphologY of mAle legs UNkNowN.

MoRphologY of mAle teRmiNAliA UNkNowN.

MoRphologY of ovipositoR (boDY: ovipositoR leNgth RAtio cA. 7.0) AND femAle geNitAl tUbes coNsisteNt with those ReveAleD foR T. glandipleurum ( Fig. 5A, C, D ).

Distribution. This species hAs beeN collecteD iN the followiNg ecoRegioNs of SoUth AfRicA (MoUNtAiN ZoNe): DRAkeNsbeRg moNtANe gRAsslANDs, wooDlANDs AND foRests ( Fig. 7 ).

Figures

FIGURE 4. Habitus images of the studied species. Tragardhus (Tragardhus) biaplicalis Koch, 1956, holotype (A), T. (T.) glandipleurum Koch, 1956, syntype (B), T. (T.) jani sp. nov., holotype (C, F), T. (T.) majae sp. nov., paratype (D) and T. (T.) stigmaticus Koch, 1956, syntype (E, G).

FIGURE 5. Male and female terminalia of Tragardhus and Pseudemmallus. Ovipositor (A – D), genital tubes (C, D), aedeagal tegmen (E, G) and penis (F). Tragardhus (Tragardhus) glandipleurum Koch, 1956 (A, C – F), T. (Mitragardhus) nodosus (B) and Pseudemmallus aspericollis (G). Abbreviations: C 1 – C 3 — subsequent lobes of coxites, val—valvifer, ov—oviduct, bcbursa copulatrix, sp—spermatheca, par—paraproct, p—penis, cl—clavae.

FIGURE 7. Distribution of the genus Tragardhus.