Tragardhus, Koch, 1956

Kamiński, Marcin Jan, 2017, Afrotropical Melambiina: revision of the genus Tragardhus Koch, 1956 and redescription of Pseudemmallus Koch, 1956 (Tenebrionidae: Pedinini), Zootaxa 4347 (1), pp. 71-93 : 83

Key to the species of subgenus Tragardhus

1. Third elytral interval forming a sharp and strongly raised continuous costa ( Fig. 4F )............................... 2

-. Third elytral interval composed of more or less elongated tubercles ( Fig. 4G )..................................... 3

2. Anterior pronotal angles protruding outwards. Lateral sides of pronotum, at base, straight. Eighth elytral interval composed of short tubercles ( Fig. 4D )............................................................... T. (T.) majae sp. nov.

-. Anterior pronotal angles not protruding outwards. Lateral sides of pronotum rounded. Eighth elytral interval, at least at middle, with a sharply raised continuous costa ( Fig. 4C ).............................................. T. (T.) jani sp. nov.

3. Fifth, seventh and ninth elytral intervals with a sharply raised continuous costae ( Fig. 2H ). Male protibia with a prominent, obtusely dentiform, arcuate median dilatation ( Fig. 2J )....................................... T. (T.) glandipleurum

-. Fifth, seventh and ninth elytral intervals composed of more or less elongated tubercles ( Fig. 2I ). Male protibia without median dilatation........................................................................................... 4

4. Pronotum cordiform, sinuate anterior to basal angles ( Fig. 4E ). Third, fifth and seventh elytral intervals covered with coarse, dense tubercles; second and fourth intervals covered with sparse tubercles........................... T. (T.) stigmaticus

-. Pronotum narrowed, subparallel in basal half ( Fig. 4A ). Third, fifth and seventh elytral intervals covered with sparse tubercles; second and fourth intervals not covered with tubercles........................................... T. (T.) biapicalis

Figures

FIGURE 2. Diagnostic characters proposed for Mitragardhus, Tragardhus and chosen species. Pronotal disc (A, B, C), epipleuron (C), 5 th abdominal ventrite (D, E), elytral disc (F, H, I), metatibia (G) and protibia (J). Tragardhus (Tragardhus) biapicalis Koch, 1956 (I), T. (T.) glandipleurum Koch, 1956 (A, E, H, J), T. (Mitragardhus) ewae sp. nov. (B, F) and T. (M.) nodosus Koch, 1956 (D, G). White arrow indicates a characteristic apex of the metatibia of T. (M.) nodosus.

FIGURE 4. Habitus images of the studied species. Tragardhus (Tragardhus) biaplicalis Koch, 1956, holotype (A), T. (T.) glandipleurum Koch, 1956, syntype (B), T. (T.) jani sp. nov., holotype (C, F), T. (T.) majae sp. nov., paratype (D) and T. (T.) stigmaticus Koch, 1956, syntype (E, G).