Jerseucoila Liu and Engel, 2007

LIU, ZHIWEI, ENGEL, MICHAEL S. & GRIMALDI, DAVID A., 2007, Phylogeny and Geological History of the Cynipoid Wasps (Hymenoptera: Cynipoidea), American Museum Novitates 3583, pp. 1-48 : 21-23

Jerseucoila Liu and Engel , new genus

TYPE SPECIES: Jerseucoila plesiosoma Liu and Engel , new species.

DIAGNOSIS: Like Anteucoila (see above) with freely articulated metasomal T2–4, this new genus can be readily excluded from crown-group Eucoilinae . The presence of a mesoscutellar plate, as in the former genus, supports its position as a stem-group eucoiline. Jerseucoila differs from Anteucoila by the apically less expanded antenna, the weakly carinate and sculptured mesoscutellar plate, the arching of the distal abscissa of Rs, the elongate metabasitarsus, the larger second metasomal tergum, and the broader petiole ( fig. 13 ) (refer also to the Diagnosis for Anteucoila ).

ETYMOLOGY: The new genus-group name is a combination of Jersey (a reference to the amber deposit in which it was discovered) and Eucoila , type genus of the subfamily. The name is feminine .

Jerseucoila plesiosoma Liu and Engel , new species figures 13 , 14

Cynipoid sp.: Grimaldi and Engel, 2005: 422 [figured].

DIAGNOSIS: As for the genus (see above).

DESCRIPTION: Female. Body length 0.60 mm; forewing length 0.51 mm. Body mostly black; wings hyaline, without macula or band. Antenna cylindrical, with distalmost flagellomeres more elongate than preceding flagellomeres; pedicel almost spherical, two-thirds as long as scape; F1 about as long as F2; elongate placodeal sensilla distinct on nearly all flagellomeres except basal two. Frons, vertex, gena, and malar space apparently glabrous (difficult to integument in holotype owing to layer of microscopic bubbles); gena not expanded behind compound eye; compound eye much longer than malar space, nearly five times longer than malar length. Lateral surface of pronotum apparently glabrous. Mesoscutum weakly curved dorsally in lateral view; mesoscutellum slightly sloped posteriorly and with weakly defined central plate; mesopleuron apparently glabrous; mesocoxa directed vertically and not inserted on separate, oblique posterior area. Propodeum devoid of processes. Wings hyaline, with scattered pubescence; all wing margins, especially posterior margins, ciliate. Forewing with marginal cell closed, about three times as long as wide; bulla in Sc+R 1 absent; 2r-rs oblique, sloping outward posteriorly; areolet absent; Rs+M reduced, scarcely traceable, apparently arising from posterior end of first free abscissa of M (5 basal vein). Metabasitarsus nearly as long as combined lengths of second to fifth metatarsomeres; pretarsal claws simple, without basal lobe. Postpetiolar metasomal terga not fused, all subequal in length along middorsal line except metasomal T2 more elongate; hypopygium of female long and slender. Male. Unknown.

HOLOTYPE: Female, AMNH NJ-1006, Late Cretaceous (Turonian), White Oak Pit , Sayreville , Middlesex County, New Jersey; deposited in the Amber Fossil Collection , Division of Invertebrate Zoology , AMNH.

ETYMOLOGY: The specific epithet is a combination of the Greek words plesios (meaning ‘‘near’’) and soma (meaning ‘‘body’’).

Figures

Fig. 13. Photomicrograph of holotype female of Jerseucoila plesiosoma Liu and Engel, new genus and species (NJ-1006).

Fig. 14. Holotype female of Jerseucoila plesiosoma Liu and Engel, new genus and species (NJ-1006).