Amara (Curtonotus) macronota (Solsky, 1875)

Li, Yihang, Li, Haoyuan & Shi, Hongliang, 2024, Revision of the macropterous subgenus Curtonotus from east China, with the description of a new species (Carabidae, Zabrini, Amara), ZooKeys 1190, pp. 39-73 : 39

Amara (Curtonotus) macronota (Solsky, 1875)

Fig. 11

Curtonotus macronotus Solsky, 1875: 265 (type locality: “Nikolskoje” [= Nikolskoye, Kamchatka Krai, Russia]; holotype in ZRAS); Matsumura 1929: 194; Lafer 1989: 180; Hieke 1995: 322; Sasakawa 2009: 107; Hieke et al. 2012: 61.

Curtonotus nitens Putzeys, 1866: 234 (type locality: "Chine boréale” [= northern China]; holotype in MNHN); Solsky 1875: 265; Lewis 1879: 189; Bates 1888: 370; Tschitschérine 1894: 385; Habu 1953: 43; Junior secondary homonym of Amara nitens Letzner, 1852.

Amara (Curtonotus) jureceki Jedlička, 1957: 29 (type locality: “Wladiwostok” [= Vladivostok, Russia]; holotype in NMPC); synonymized by Lafer 1989: 180.

Amara (Curtonotus) ovalipennis Jedlička, 1957: 30 (type locality: “Kyoto” [in Japan]; holotype in NMPC); synonymized by Hieke 1995: 322.

Specimens examined.

1 male (CLYH), China, Beijing, Haidian district, Baiwangshan Forest Park, 40.033893°N, 116.256957°E, 100 m, 2021.03.12, Yihang Li leg. ; 2 males (CLYH), China, Beijing, Changping district, Hedi Road , 40.139454°N, 116.305624°E, 40 m, 2022.06.13, Yihang Li leg. ; 3 males (CLYH), China, Sichuan, Mianning County, Tuowu Mountain , 2200 m, 2022.04.15, Yuan Li leg. ; 1 female (CLHY), China, Beijing, Haidian district, Yuanmingyuan Park , 40 m, 2022.02.06, Haoyuan Li leg. ; 3 males, 2 females (CBJFU), China, Beijing, Shunyi District, Hanshiqiao Wetland Reserve , 2016.10.17, Pingzhou Zhu leg. ; 2 males (CBJFU), China, Shanxi, Gujiao, Yunding Mountain Preserve , 35.5246°N, 111.3553°E, 1790 m, 2021.09.10, Xiaojie Sun leg. ; 1 female (CLYH), China, Hunan, Huaihua, Subaoding Mt , 1800 m, 2022. 07. 14, Yihang Li leg. ; 1 male (CLYH), China, Guizhou, Changshun county, Changshun No. 1 Primary School , 2020.08.25, local collector leg .; 4 males, 5 females (CCJH), China, Guangxi, Ziyuan, Shilipingtan, Zijinshan Mt , 26.169585°N, 110.499607°E, 1734 m, Jiaheng Chen leg.

Chinese common name.

巨胸暗步甲.

Diagnosis.

Large-sized species, BL = 10.5-13.5 mm; body completely black, legs reddish brown to black; head medium sized, more than half length of pronotum, with two supraorbital setae. Pronotum (Fig. 11B ) cordate, widest at middle; densely and very coarsely punctate at basal region, very sparsely punctate at anterior portion; lateral margins strongly sinuate before posterior angles; posterior angle strongly protruded laterally, apex sharp, nearly rectangular. Elytra oblong, widest after middle; coarsely punctate on basal two-thirds of striae; humeral tooth protruded, but smaller than the previously one, apex a little bent backward; microsculpture isodiametric in both sexes; lateral sides of abdominal sternites punctate and wrinkled. Male mesotibiae projection composed of two denticles (Fig. 11C ); proximal denticle acutely pointed, near apical third of tibiae; distal denticle slightly smaller, present between the proximal one and tibiae apex. Male genitalia with relatively long apical lamella (Fig. 11F ), nearly straight and widely triangular, gradually narrowed to apex, apex rounded; gonocoxite 2 (Fig. 11G ) of ovipositor stout, length subequal to greatest width, apex widely rounded.

Comparison.

This species can be distinguished from related species by having very strongly sinuate lateral margin before posterior angles, nearly straight elytra basal border, heavy punctate on pronotum basal region and elytra striae, and very short gonocoxite.

Distribution.

Beijing, Fujian, Gansu, Guangdong, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Jiangsu, Jilin, Jiangxi, Liaoning, Inner Mongolia, Sichuan, Shaanxi, Shanxi, Shanghai, Shandong, Tianjin, Yunnan, Zhejiang, Japan, North Korea, South Korea, Russia (Far East), Russia (east Siberia).

Figures

Figure 11. Amara (Curtonotus) macronota A dorsal habitus, male (Gujiao, Shanxi) B pronotum posterior angle C male mesotibia D lateral view of aedeagus E right paramere F apical lamella G female gonocoxite (Shunyi, Beijing). Scale bars: 5 mm (A); 1 mm (B-E); 0.5 mm (F, G).