Heth, Cobb, 1898

Phillips, Gary, Moulton, John K. & Bernard, Ernest C., 2020, Heth pivari n. sp. (Nematoda: Ransomnematoidea: Hethidae) from the indigenous North American millipede Narceus gordanus (Spirobolida: Spirobolidae), with keys for worldwide Heth spp., Zootaxa 4861 (4), pp. 486-514 : 506

Key to Heth species groups

1. With continuous collar of cervical spines, having only a minor interruption of larger lateral spines; lappets present or absent ( Figs. 9 , 10 ).......................................................................................... 2

- Cervical collar of spines discontinuous or absent; lappets absent................................................ 3

2. Lappets absent ( Figs. 9 , 10 , 11D )........................................................... Group 1 Heth spp.

- Lappets present ( Figs. 11 , excluding 11D, 12)............................................... Group 2 Heth spp.

3. With discontinuous cervical collar ( Figs. 13 , 14 )............................................... Group 3 Heth spp.

- Without clear cervical collar; lappets absent: not conforming to any other group description ( Fig. 15 )..... Group 4 Heth spp.

Figures

FIGURE 9. Worldwide Heth spp., continuous collar and no shield (Group 1). A) H. albertoi (after García & Fontana 2004). B) H. amazonensis (after Kloss 1965). C) H. bifidispiculum (after Adamson 1982). D) H. clunyi (after Adamson 1985). E) H. coyi (after García et al. 1997). F) H.gordae (after Mejía-Madrid 2014). G) H.hexaspinosum (after Chitwood 1935).H) H. hispaniolae (after García et al. 2001). I) H. konoplevi (after Malysheva & Sumaya 2017). J) H. maicuru (after Adamson 1983).

FIGURE 10. Worldwide Heth spp. Group 1 (continued). A) H. mauriesi (after Adamson 1982). B) H. perarmatum (after Dollfus 1952). C, D) H. pinnatum, head end and neck region (after García & Coy 1995). E) H. spinosum (after Adamson 1983). F) H. xarochae (after Mejía-Madrid 2014). G) H. zeuglocantha (after Hunt 1994).

FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).

FIGURE 13. Worldwide Heth spp., discontinuous collar, no shield (Group 3). A) H. artigasi (after Dollfus 1952). B) H. artigasi (after Adamson 1983). C) H. baracoa (after Spiridonov 1989). D) H. duvidosum (after Artigas 1929). E) H. imias (after Spridonov 1989). F) H. macrocephala (after Kloss 1965). G) H. multiplus (after Kloss 1965). H) H. parartigasi (after Adamson 1985).

FIGURE 14. Worldwide Heth spp. Group 3 (continued). A) H. sinediscus (after Kloss 1965). B) H. spinalatum (after Kloss 1965). C) H. spoliatus (after García & Coy 1995). D) H. travassosi (Dollfus 1952). E) H. travfilhoi (after Adamson 1983). F) H. tuzetae (after Adamson 1983).

FIGURE 15. Worldwide Heth spp. not conforming to any other group (Group 4). A) H. adolphi (after Sanchez-Velazquez 1979). B) H. josephinae (after Sanchez-Velazquez 1979). C) H. lamothei (after Sanchez-Velazquez 1979). D) H. poeyi (after Otero et al. 1993).