Helicopsyche (Feropsyche) sanblasensis, Johanson, Kjell Arne & Malm, Tobias, 2006

Johanson, Kjell Arne & Malm, Tobias, 2006, Seven new Helicopsyche (Feropsyche) Johanson, 2002 from the Neotropical region and Nearctic Mexico (Insecta: Trichoptera: Helicopsychidae), Zootaxa 1208, pp. 1-24 : 19-22

Helicopsyche (Feropsyche) sanblasensis , new species

Fig. 52–59 , 61

Helicopsyche sanblasensis , new species is unique in having a combination of a very short segment IX, a nearly 3­branched primary branch of the gonocoxite, and a very long, coneshaped basimesal lobe. With respect to the genitalia, especially the shape of the gonocoxite, H. sanblasensis is similar to H. extensa Ross and H. dampfi .

Male head. Cephalic warts oval and flat. Postantennal setal warts not observed. Antennal scape about as long as each maxillary palp segment and eye diameter.

Male wings. Forewing ( Fig. 52 ). Forewing brownish, length about 3.2 mm; hind wing length 2.3 mm. Hind wing venation and hamuli not observed.

Male abdomen and genitalia (53–59). Segment IV with broken process ( Fig. 53–54 ). Segment IX ( Fig. 55 ) with anterior lobe triangular in lateral view, oriented anteriad and located dorsally on segment; anterodorsal margin nearly straight, about as long as shallowly concave anteroventral margin; in dorsal view with inner margin widely ellipsoid ( Fig. 56 ); in ventral view with wide central posterior process ( Fig. 57 ). Lateral apodeme curving slightly anterodorsad in lateral view ( Fig. 55 ), tapering and fading before anterior margin; sub­marginal line absent; tergal transverse apodeme absent; sternal transverse apodemes short. Tergum X oriented posteroventrally in lateral view ( Fig. 55 ), nearly straight and cone­shaped, with apex pointed; widening at midlength and narrowing toward rounded apex in dorsal view ( Fig. 56 ), apical notch shallow, with about 7 pairs long dorsal and 5 pairs short lateral megasetae. Superior appendage thick, slightly club­shaped in dorsal view ( Fig. 55 ). Primary branch of gonocoxite with deeply concave posterodorsal margin in lateral view ( Fig. 55 ) resulting in 3­branched gonocoxite (basimesal lobe included); narrowest distally to base, with weakly undulate margins; anterodorsal margin smooth, apex cone­shaped in lateral view and strongly curving mesally in dorsal and ventral view ( Fig. 56, 57 ); narrowest part of primary branch as wide as height of tergum X and narrower than tergum X width ( Fig. 55, 56 ); anterodorsal margin nearly concave, posteroventral margin shallowly concave along its length, with 2 prominently erect setal bases ( Fig. 55 ). Basimesal lobe broadly cone­shaped and apically pointed in lateral and ventral view ( Fig. 55, 57 ), with median margins basally divergent and distally parallel ( Fig. 57 ), about 6 megasetae on dorsal face. Basal plate narrow, slightly undulate in lateral view ( Fig. 55 ); strongly pointed anteriorly in ventral view ( Fig. 57 ). Phallus with ventral margin curved at middle in lateral view ( Fig. 58 ); in lateral and ventral view ( Fig. 58, 59 ) basal part slightly thicker than central region. Phallic basis narrow ( Fig. 59 ). Endotheca produced, posteroventral part strongly sclerotized. Sperm channel divided inside phallus into slender anterior part and thicker posterior part ( Fig. 58, 59 ).

Male holotype: PANAMA: San Blas: 2 km S. Nusagandi, 3.iii.1985 [Flint & Louton] (NMNH, pinned, genitalia in glycerol).

Distribution: Panama (San Blas) ( Fig. 61 ).

Etymology: sanblasensis , derived from the type locality of the species, San Blas in Panama.

Remarks: Helicopsyche sanblasensis can be separated from H. extensa by the shorter, cone­shaped apical branch of the primary gonocoxite branch, the cone­shaped tergum X in lateral view, and the long and cone­shaped basimesal lobes. It is distinguished from H. dampfi by the shorter segment IX, the cone­shaped apical branch of the primary branch of the gonocoxite, and the longer and cone­shaped basimesal lobe.

Figures

FIGURE 52. Helicopsyche sanblasensis, new species, holotype. 52 — habitus, lateral.

FIGURES 53 – 59. Helicopsyche sanblasensis, new species, holotype. 53 — VIth sternal process, lateral; 54 — VIth sternal process, ventral; 55 — genitalia, lateral; 56 — genitalia, dorsal; 57 — genitalia, ventral; 58 — phallus, lateral; 59 — phallus, ventral.

FIGURES 60 – 62. Distribution maps showing the localities of the new species and the new records. 60 — Mexico, showing the distributions of H. curvipalpia (filled square) and H. sinuata (filled circle); 61 — Panama, showing the distributions of H. incisa (filled square), H. chiriquensis, H. linguata, H. woldai and H. incisa (filled circle), H. blantoni and H. woldai (open square), and H. sanblasensis (cross); 62 — Brazil, showing the distributions of H. paprockii and H. cipoensis (filled square).