Cletocamptinae, Gómez & Yáñez-Rivera, 2022

Gomez, Samuel & Yanez-Rivera, Beatriz, 2022, The genus Cletocamptus (Harpacticoida, Canthocamptidae): a reappraisal, with proposal of a new subfamily, a new genus, and a new species, ZooKeys 1080, pp. 165-208 : 165

Subfamily Cletocamptinae subfam. nov.

Type genus.

Cletocamptus Schmankevitsch, 1875.

Other genera.

Amphibiperita Fiers & Rutledge, 1990, Cletocamptoides gen. nov.

Diagnosis.

Canthocamptidae . Body fusiform, without clear distinction between prosome and urosome. Without nuchal organs on cephalothorax or body somites. Female rostrum distinct, rarely fused to cephalothorax, large and triangular with broad proximal margin; ornamented with ventral (sub)distal spinules or (occasionally) setules and with rounded or (rarely) bilobed tip in both sexes. Cephalothorax and/or prosomites with posterior margins serrated or ornamented with short or long slender spinules; posterior margin of urosomites except for anal somite serrated or with short spinules; posterior margin of cephalothorax and body somites without or (rarely) with cuticular sensillum-bearing socles. Anal operculum without dorsal ornamentation or with transverse row of strong large or short, small spinules; posterior margin unornamented, serrated, or with small or large spinules. Female genital somite and third urosomite separated dorsolaterally, completely fused ventrally forming genital double-somite. Female antennule six-, rarely seven-segmented. Antenna with allobasis, with one or two abexopodal setae (proximal element basal, distal seta endopodal); exopod one-segmented, longer than wide or minute, or absent. Mandibular palp one-segmented, very small and wider than long, or longer than wide; basis and endopod incorporated to basis or (rarely) with basis and endopod distinct (uniramous); exopod absent or (rarely) represented by single seta; when palp uniramous, then basis unarmed or with one seta, endopod with three setae at most; when palp one-segmented and longer than wide, then with two basal and two endopodal setae at most, and exopod (when present) represented by single seta; when palp one-segmented very small and wider than long, then basis and endopod not discernible, with one or two (most probably endopodal) setae, with or without surface (most probably exopodal) seta on coxa. Maxillule with endopod and exopod incorporated to basis; praecoxal arthrite with ventral seta thick and strongly spinulose, or slender and pinnate or smooth. Maxilla with two syncoxal endites; endopod completely incorporated to allobasis. Maxilliped subchelate; syncoxa with one seta; basis unarmed; claw with accessory seta. P1 not prehensile, rarely prehensile; P1-P4EXP three-segmented; female P1-P3ENP two-segmented, P4ENP two- or one-segmented; inner exopodal and endopodal setae with or without comb tip. Female P5EXP and BENP fused, occasionally separated; both baseoendopods of P5 separated. Armature formulae as follows:

Female P6 with one or two setae. Caudal rami with six or seven setae; setae IV and V fused basally or separated.

Sexual dimorphism can be expressed in a) rostrum slenderer in the male, b) the male antennule (chirocer or subchirocer), c) basis of P1 (with or without inner distal process), d) outer spine on P2 ENP2 (thicker and/or shorter than in the female), e) P2EXP and/or P3 and P4EXP (segments longer than in the female, outer spines stronger than in the female, rami curved inwards,), f) shape of P2-P4ENP (segments thicker than in female), g) P3ENP (two- or three-segmented; when three-segmented then inner apophysis on second segment; when two-segmented then inner apophysis medially or subdistally on second segment, occasionally with additional outer apophysis; inner apophysis simple, without arrow-head tip, variable in length; with or without asprothekes on second segment), h) P5 (both legs fused medially or separated; EXP and BENP fused; both legs separated or fused to somite), i) P6 (composed of two lappets articulated to somite or asymmetrical in which case only one leg functional, the other fused to somite; unarmed or (occasionally) with one or two setae), j) caudal rami (longer than in the female).