Rhaucus marmoratus (Roewer, 1912) Pinzón-M & Pinto-Da-Rocha, 2024

Pinzón-M, Conchita & Pinto-Da-Rocha, Ricardo, 2024, Two new species of Rhaucus Simon, 1879 reveal putative synapomorphies related to the microstructural morphology of male genitalia (Opiliones: Cosmetidae), Zootaxa 5555 (1), pp. 35-56 : 47-48

Rhaucus marmoratus (Roewer, 1912) New Combination

( Figs 8 , 12 )

Pararhaucus marmoratus Roewer, 1912c: 142 , pl. 7, Figs. 2–3 ; Flórez & Sánchez, 1995: 368. Meterginus marmoratus : Roewer, 1912b: 106; 1923: 383, Fig. 467; Mello-Leitão, 1932: 87; Kury, 2003: 72, 280.

Type Material: Colombia, Cundinamarca, Between Tambo and Boca del Monte (4 males syntypes MNHN, not examined; 1 male syntype SMF 285-32 examined by photos) .

Other Examined Material: Colombia, Cundinamarca: 3 males and 3 females ( ICN-Ar-1659 ), Parque Nacional Natural Chicaque , 4°36’5.6’’N, 94°18’46’’W. Grupo Biología leg. 15.X.2016 .

Diagnosis: Ozopore dorsally located, with the ID mostly projected and an elongated slit of the ozopore extending towards the lateral margin of the DS (with a rounded ozopore in R. artifex sp. nov., R. papilionaceus , R. quinquelineatus , R. serripes , and R. vulneratus ; in R. serratus comb. nov., the integumentary dome covers the ozopore and is well projected dorsally; in R. gaiterus sp. nov., there is no dorsal fold). The ocularium has a well-marked medial depression (not well-marked in R. artifex sp. nov., R. florezi , R. gaiterus sp. nov., R. robustus , R. quinquelineatus , and R. serratus comb. nov.). Area III of the DS has a pair of prominent mammiliform tubercles close to each other (spiniform tubercles in R. artifex sp. nov. and R. gaiterus sp. nov.; mammiliform and smaller in R. florezi and R. vulneratus ). Cx IV has clavi inguines formed by a mound with few granules (strongly granulated in R. florezi , R. quinquelineatus , R. serripes , and R. vulneratus ). The dorsal apical apophysis on Cx IV is multicapitate, forming a keel slightly flattened laterally (not laterally flattened in the remaining species). The color pattern consists of two lateral longitudinal patches extending from the prosoma to the posterior areas of the DS (see Fig. 8A–B ).

Penis Description: ( Fig. 8C–F ) VP is short and squared. The glans has a dorsal process that is elongated and cylindrical, with wattle extending ventrally and bearing stylar barbs. Two pairs of MS C: C1 is mostly more curved than C2, with both being long and flattened. Two pairs of MS D: D1 is cylindrical and straight, closer to the pair C, while D2 is dorsally attached, short, and located in the middle portion of the VP. Two pairs of MS A: dorsally attached, straight, cylindrical, and bifurcated at the top. Two pairs of MS E are volcano-shaped, and a pair of areolate MS B is located at the base of the VP. The lateral fields of the VP are covered by microsetae type 4, and the medial portion has some microsetae T2. The lateral corners of the VP are curved dorsally.

Figures

FIGURE 2. Rhaucus artifex sp.nov. A–B: Male holotype (minor male) (MUSENUV-Ar-2618). A. Habitus, dorsal view. B. Habitus, lateral view. C. Male Paratype (major male) (MNRJ 1594). Dorsal view. D. Female paratype (MUSENUV-Ar 2619). Habitus, dorsal view. Foto C courtesy of Daniela Ahumada, scale bar 5mm.

FIGURE 3. Rhaucus artifex sp. nov. A–J: Male holotype (MUSENUV-Ar-2618).A. Habitus, dorsal view; B. Right chelicera, dorsal view; C. Right hand of chelicera, anterior view; D. Right leg, Pa, dorsal view; E. Left leg I, tarsomeres, mesal view; F–H. Left leg IV: F. Ti, ventral view; G. Fe, ventral view; H. Fe, dorsal view; I–J. Left pedipalp: I. Tro, Fe, Pa, Ti, Ta, Dorsal view; J. Same, Mesal view. Scales: 1 mm.

FIGURE 8. Rhaucus marmoratus comb. nov. (ICN-Ar-1659) A. Male. Habitus, dorsal view; B. Female. Habitus, dorsal view; C–F. Penis: C. Dorsal view; D. Ventral view, arrow showing the areolated MS B.; E–F. Lateral view. Scales: 0.05 mm. Micrographs C–F. Courtesy of B. N. Damrom.

FIGURE 12. Distribution map of Rhaucus spp.