Onthophagus (Palaeonthophagus) medius ( Kugelann, 1792 )

Rössner, Eckehard, Schönfeld, Joachim & Ahrens, Dirk, 2010, Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) — a good western palaearctic species in the Onthophagus vacca complex (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini), Zootaxa 2629, pp. 1-28 : 20-27

Onthophagus (Palaeonthophagus) medius ( Kugelann, 1792) (revised status)

Copris medius Kugelann, 1792: 271 (Type locality: "Preußen").

Scarabaeus medius Panzer, 1796: 1 (Type locality: Germany).

Copris affinis Sturm, 1800: 102 (Type locality: "Michaelsberg bei Herspruck" [probably Bavaria]). Onthophagus vacca var. vicinus Mulsant, 1842: 133 (Type locality: France). Onthophagus vacca var. intermedius Mulsant, 1842: 134 (Type locality: France). Onthophagus vacca var. propinquus Mulsant, 1842: 134 (Type locality: France). Onthophagus vacca var. sublineolatus Mulsant, 1842: 134 (Type locality: France). Onthophagus vacca var. basalis Mulsant, 1842: 134 (Type locality: France). Onthophagus confluens Gistel, 1857: 61 (Type locality: Germany).

Type material examined: Neotype ( Copris medius , here designated) [Figs. 2L, 2P, 2AF, 2AJ, 3A, 3B, 3AG, 3AP]: "Lyck, Ostpreussen [black handwritten] B.v.Bodemeyer [black printed on white paper] | Onthophagus vacca v. medius [black handwritten on white paper] | Neotypus Copris media Kugelann, 1792 , E. Rößner, 2008 [black handwritten on red paper] | Onthophagus (Palaeonthophagus) medius ( Kugelann, 1792) , det. E. Rößner, 2008 [black handwritten on white paper]" (ɗ, ZMHB) [specimen glued on carton, genital extracted]. Denmark: "Helnaes, Bobakker 8.v.1999 | DK.F.NG61, P. Jörum leg." (2 ΨΨ, CPJ).- "Rösnaes (Rösnaesgd.) 11.vi.2005 | DK.NWZ.PG17, P. Jörum leg." (1 ɗ, 1 Ψ, CPJ).- "Mögelskår 19.v.2004 | DK.EJ.NH90, P. Jörum leg." (2 ΨΨ, CPJ).- "Kopenhagen | coll. Leonhard" (2 ɗɗ, 2 ΨΨ, DEI). Netherlands: "Wateringen, 4.V.1954, R. Bouwer" (1 ɗ, CJS).- "Opheusten, 20.VI.1957, R. Bouwer" (1 Ψ, CJS).- "Maastricht, 5.1921, [leg.] Rüschkamp, Sammlung F. Rüschkamp. Eing. Nr. 1.1964" (2 ΨΨ, ZMFK). Luxemburg: " Luxemburg, Differdingen, 4.[19]15, P. Heymes" (1 Ψ, coll. Heymes, NME).- " Luxembourg: Remich, Erpel D’Ange, Chatea Dames 29.viii.1998, ex horse dung, coll. M.V.L. Barclay" (4 ɗɗ, 11 ΨΨ, NHML).- "Grevenmacher, Manternach, 0 2.06.2001, leg. Dirk Rohwedder" (1 ɗ, CDR).- "Grevenmacher, East-Gonderange, Schafkot, 0 3.06.2001, leg. Dirk Rohwedder" (1 Ψ, CDR).- "Gonderange, 0 1.06.2001, leg. K. Hadulla" (1 Ψ, CKHa). Belgium: " Belgium " (1 ɗ, 1 Ψ, coll. Fry, NHML).- "Antwerpen 29.5.1898." (1 ɗ, NMP). Germany: "Mecklenbg.: Klein Schmölen, Pferdeweide an der Löcknitz, 2833/IV, 01.V.2005, Pferdekot, leg. E. Rößner" (5 ɗɗ, 5 ΨΨ, CER).- dto., nur: " 25.IV.2006 " (4 ɗɗ, CER).- "Lebus/ O. 23.5.[19]72" (1 ɗ, 1 Ψ, ZMHB).- "Lebus, Oderbruch, DDR, 21.V.1982, leg. U. Heinig" (1 Ψ, CER).- "Oderhänge bei Lebus, DDR, 28.V.1988, leg. Schröder" (1 ɗ, 1 Ψ, CER).- "Teichhorn, Volksen [bei Einbeck], 5.6.[19]55, Kuhmist, Priefert & Helmke lg." (1 ɗ, ZMHB).- "Bottendorf [wahrscheinlich: bei Obernholz], 23.6.[19]55, Lohse" (1 Ψ, ZMHB).- "Baden b. Bremen 13.5.[19]51, 53, Kerstens" (1 ɗ, CAB).- "Bremen, Baden 11/6 1950, J. D. Schroeder" (1 ɗ, CAB).- "Badener Berge. 8.IV. [18]94. Achim. Hannover, coll. Künnemann" (1 ɗ, DEI).- dto., nur " 24.IV.98 " (1 Ψ, DEI).- "Lindhorst, Kr. Stadth. Hannover, IV.1951, leg. E. Gersdorf" (1 Ψ, CJS).- "Geestgottberg, ST Altmark, Elbedeich b. Brücke, Schafkot 17.05.2006 [leg.] Malchau" (1 ɗ, 2 ΨΨ, CWM).- " Germania sept., Gatersleben (Quedlinburg), 22.6.1952. 127. [leg.] Dorn" (3 ɗɗ, 3 ΨΨ, ZMHB).- "Dornburg [bei Zerbst], JL, ST, Gommern, Weide östl. Ortsrand, 24.05.2007, Malchau" (1 ɗ, 3 ΨΨ, CWM).- "Neugattersleben, ST, BBG, Bodegrünland bei A 14, Pferd Schaf, 16.05.2007, Malchau" (1 ɗ, 2 ΨΨ, CWM).- "Breitenhagen: Sommerdeiche, 18.IV.2005, leg. E. Rößner" (2 ɗɗ, 2 ΨΨ, CER).- "Sachsen-Anhalt: NSG Lämmerberg und Vockenwinkel bei Wormsleben, 4435/IV, 29.V.1998, leg. E. Rößner" (2 ɗɗ, 4 ΨΨ, coll. CER; 3 ɗɗ, CHKa).- "Sachsen-Anhalt: Wormsleben, 4435/IV, 26.V.1999 " (2 ɗɗ, 1 Ψ, CER).- "Sachsen-Anhalt, Dabrun bei Wittenberg, 4142/3, Rinderkot, 18.05.2002 Bäse leg." (1 ɗ, 1 Ψ, CWB).- "Sachsen-Anhalt 4244/2, Premsendorf bei Jessen, an Pferdekot, 16.05.2005 K. Bäse leg." (1 ɗ, 1 Ψ, CWB).- "Beesenlaublingen, ST, BBG, A 14 Brücke, Saaledeich, 0 3.05.2007, Malchau" (1 Ψ, CWM).- "Thuringia MTB 4531/4, Numburg- West, Kelbra, 20.V. [20]05, Salzwiese, Sw B, leg. H. Sparmberg" (1 Ψ, CHS).- "Thuringia MTB 4531/4 Kelbra, Numburg, West, 27.IV.2007 Hf leg. H. Sparmberg" (2 ΨΨ, CHS).- "Thuringia, 17.V.2002, 4832/III, Haßleben, Rinderweide, trocken, Fangart: Hf, leg. H. Sparmberg" (1 Ψ, CHS).- "Kyffhäuser Schlachteberg 21.04.1989, R. Peschel" (1 Ψ, CRP).- "Bad Frankenhausen, A.06.1983, leg. E. Rößner" (1 Ψ, CER).- " BRD / Thür./ Kyffhäuser, Udersleben 250 NN, U. Hämling, 28.4.1994, leg. U. Schaffrath" (1 ɗ, 1 Ψ, CUS).- "Thuringia MTB 4632/IV, Esperstedter Ried, Hf, 17.VI. [20]05, Gebiet A, leg. H. Sparmberg" (1 ɗ, 1 Ψ, CHS).- "Rastenberg: Kapellenberg, 4834/II, HF, 08.VI.2008, leg. H. Sparmberg" (3 ɗɗ, CHS).- "Nöda: Rinderweide, Feuchtwiese, 4932/I, HF, 14.VI.2008, leg. H. Sparmberg" (1 ɗ, CHS).- "Meining[en]. (Heyd.)" (2 ɗɗ, 1 Ψ, coll. Heyden, DEI).- "Saxonia Provincia, Eilenburg, Hainichen Nord, 13.4.1953.32, [leg.] Dorn" (2 ɗɗ, 2 ΨΨ, ZMHB).- "14.5.[19]56, Naunhof [Großenhain], [leg.] Hieke" (1 ɗ, ZMHB).- "Meißen, Zadel, 24.6.[19]70" (1 Ψ, ZMHB).- "D: Saxonia cent., Köllitzsch/Elbe 29.V.2001 Grünland, Projekt Uni Jena BF" (1 ɗ, 1 Ψ, NME).- "Düben Kirsch" (1 Ψ, SMTD).- "Wehlen Märkel" (1 Ψ, SMTD).- "Grunaer Wald, Düb. Heide 3.5.[19]53, L. Dieckmann, Pferdemist" (2 ɗɗ, 2 ΨΨ, DEI).- "D, Umg. Schlüchtern, Elm, in Kuhdung, 500 NN, 11.6.1989, leg. Schaffrath" (1 Ψ, CUS).- "Lampertheim/ Rhein, S-Hessen D., 1.5.1999, leg. G. Müller" (1 Ψ, CUS).- "Darmstadt 29.4.- 30.7.1949 E. Möhn leg." (1 Ψ, SMNS).- "Hofgeismar, Hess.-Nass. 4.V. [19]30" (3 ɗɗ, 1 Ψ, SMNS).- "Wiesbaden. | Collektion Kirschbaum" (1 Ψ, NSMW).- "Frankft. Hyd." (4 ɗɗ, 2 ΨΨ, coll. Heyden, DEI).- "Rheinprovinz, Rees, 11.-13.6.[19]35, K. Ermisch." (2 ɗɗ, ZMHB).- "D., R.-Pf., UG Mainz, Nackenheim, in Pferdeäpfeln, 100 NN, 22.- 24.4.1990, leg. U. Schaffrath" (1 Ψ, CUS).- "D. Rheintal, Bacharach, Steeg Weinberge, F. Köhler 22.05.1996 " (1 ɗ, ZMHB).- "D, RP, Nahe-Glan-Gebiet, Steckweiler, Solzenberg 280 m 24.04.2001, leg. F. Köhler" (5 ɗɗ, 2 ΨΨ, CHB).- "Rheinland-Pfalz: Steckweiler (im Alsenztal), NSG Stolzenberg, 24.IV.2001, leg. Frank Köhler" (26 ɗɗ, 10 ΨΨ, CER).- "D – RP-LS, Mosel, Kobern, Bellbach-Tal 100 m, 0 9.05.2002, leg. J. Beller" (1 Ψ, CHB).- "Ger., 28.V.1978, Rh.-Pf./ Mosel, Trier, Petrisberg. Scheuern leg." (1 ɗ, CJS).- "D – Rheinl.-Pfalz/ Südpfalz, Schweighofen (SÜW) beim Bahnhof, 24.V.2006, leg. L. Schmidt" (2 ɗɗ, 4 ΨΨ, CLS).- "D/ BW/ Breisach Oberbergen, Schelinger Weide, Kaiserstuhl, 4.6.2000, leg.: Willers" (1 ɗ, ZMHB).- "Kaiserstuhl, Schelingen, 0 4.06.2000, K. Renner" (1 ɗ, CKR).- "Rheinland/Pfalz, Schweighofen, Schafdung in Falle, 19.05.2007, leg. Dirk Rohwedder" (2 ɗɗ, CDR).- "Düsseldorf, L, 5.6.1962 " (1ɗ, coll. Koch, ZMFK).- "Düsseldorf, Z, 17.6.1964 " (1 ɗ, coll. Koch, ZMFK).- "Worring[er]. Bruch, [leg.] H. Appel 18.5.1957, ex Coll. Appel" (1 ɗ, 1 Ψ, coll. CAG Wuppertal, ZMFK).- "Rh. Ufer b. Köln, [leg.] Appel 2.5.1965, ex Coll. Appel" (1 Ψ, coll. CAG Wuppertal, ZMFK).- "Bienwald, Kuhweide 500 m N Kapweyer, 49°2’4" N 8°1’20" E, 24.05.2008, leg. J. Schönfeld" (1 ɗ, 1 Ψ, CJS).- "Bienwald, Pferdeweide 1 km E Büschelberg, W der B 9, 49°1’17" N 8°11’7" E, 22.05.2008, leg. J. Schönfeld" (1 Ψ, CJS).- "Bienwald, Kuhweide 500 m W Neulauterburg, nahe der Lauter, 48°58’42" N 8°10’8" E, 23.05.2008, leg. J. Schönfeld" (1 Ψ, CJS).- "Kaiserswerth, Rheinufer (südl.), 14.5.1919, Kirsch l." (4 ɗɗ, coll. ARK, ZMFK).- "Düsseldorf, 29.5.1928, [leg.] C. Henseler" (1 ɗ, coll. ARK, ZMFK).- "Elberfeld, [leg.] Geilenk[euser]." (3 ɗɗ, coll. ARK, ZMFK).- "Niederrhein, Rees, [leg.] Horion, K., 6.1935" (1 ɗ, 1 Ψ, coll. ARK, ZMFK).- "Rees, Nrh., Juni 1935, [leg.] P. Müller, Sammlung M. Schmaus, Eing.Nr. 2, 1979" (1 ɗ, coll. ARK, ZMFK).- "Acker, Bonn, 9.5.1927, [leg.] Rüschkamp" (1 Ψ, coll. ARK, ZMFK).- "Stobeln, 7.5.1928, [leg.] C. Henseler" (1 Ψ, coll. ARK, ZMFK).- "Kastellaun, 28.6.1949, [leg.] M. Schmaus" (1 Ψ, coll. ARK, ZMFK).- "Hessen, Wetterau, 4.1935, [leg.] Schoop 09" (1 ɗ, ZMFK).- "Schelingen/Kaiserstuhl, Rheinaue, 28.4.[19]96 leg. Esser" (1 Ψ, CAB).- "Bavaria, Erlangen" (2 ɗɗ, 3 ΨΨ, ZMHB).- "Ingolst[adt]. 22.5.[19]02" (1 Ψ, SMTD).- "München, Coll. Kraatz" (1 ɗ, 1 Ψ, DEI).- "Monachium [= München], var. medius Panz. " (1 ɗ, ZSM).- "Monachium, var. affinis Sturm. " (1 ɗ, ZSM).- "Straubing 8.[19]11" (1 ɗ, ZSM).- "Geisenfeld 21.V. [19]36 | Sammlung O. Bühlmann" (1 ɗ, ZSM).- "Schleissheim 7.6.[19]04 | München H. Kulzer" (1 ɗ, 1 Ψ, ZSM).- "Hartmannshf. [= Hartmannshof], 19.6.[19]06 | München H. Kulzer" (1 ɗ, 1 Ψ, ZSM).- "München Umg., Obb." (1 ɗ, ZSM). Poland: "Posen [= Poznań], [leg.] Paulisch" (1ɗ, ZMHB).- "Ohlau [= Oława] | Coll. Letzner" (1 ɗ, DEI).- "Ost-Pr[eußen]." (1 Ψ, ZMHB).- "Silesia Kirsch" (1 ɗ, SMTD). Austria: "Neusiedlersee, 16.6.1936, Burgenland, J. Wirthumer" (1 ɗ, 1 Ψ, NMP).- " Österreich Wien | ex coll. Dr. Kallert Hamburg" (2 ɗɗ, NMP).- "Präwald [Oberösterreich, südwestlich Windischgarsten], Kr, 26.6.[19]08" (1 Ψ, ZMHB).- "Burgenland, Neusiedl" (1 ɗ, 2 ΨΨ, ZMHB).- "Neusiedler See, 25.5.1975 " (1 ɗ, ZMFK).- "Umg. Wien, [leg.] Ad. Hoffmann, ex Coll. Missionshaus SVD Steyl, Eing. Nr. 3/59" (4 ΨΨ, ZMFK).- "Neusiedler See u. Umg. A 6 [19]63 leg. Witzgall" (1 Ψ, CRP).- "Neusiedl. See u. Ug., Austria, Dr. Lechner" (1 ɗ, CAB).- "12.7.[19]73 Burgenland, Parndorf/Hutweide" (1 Ψ, CAB).- "A. NSS Apetlon 6.5.1986 leg. Hirgstetter" (1 Ψ, CRP).- "4.4.[19]74 Nähe Apetlon, Austria, Neusiedler See, Fery leg." (2 ɗɗ, 1 Ψ, CHF).- "15.5.[19]76, Pamhagen ( Austria), Neusiedler See, Pferdekot, Fery leg." (1 ɗ, CHF).- " Hungaria, Leitha-Geb., F. Blühweiss" (2 ΨΨ, ZMHB).- "Illmitz Brgld. 11.5.[19]66 Folwaczny" (5 ɗɗ, 1 Ψ, SMNS).- "Neusdl. See 25.5.1975 " (2 ΨΨ, SMNS).- "Aust. | 26960" (4 ɗɗ, 2 ΨΨ, ZMHB).- "Austria-Burgld., leg. Hebauer | Neusiedl. See, 25.5.1975 " (3 ΨΨ, CJS).- "A.i. Leitha-Geb., Ad. Hoffmann" (1 Ψ, SMTD).- " 2.VI.1928. Umgebung Apetlon am Neusiedler See" (1 Ψ, SMTD).- "Neusiedler See, leg. R. Zimmermann 1941, Schmidt" (3 ɗɗ, 2 ΨΨ, SMTD).- dto., nur " Juli 49 " (1 Ψ, SMTD).- "Bündten, coll. Stierlin" (1 ɗ, DEI).- "Villach" (1 ɗ, ZSM).- "Neusiedler See, leg. Zimmermann 1941, [coll. H.] Schmidt" (1 Ψ, ZSM).- " Austria Bgld. Parndorf, Hüdepohl 5.[19]69" (1 Ψ, ZSM).- "Neusiedler See, V.1953, [leg.] Dr. H. Lechner" (1 ɗ, CLS). Czech Republic: "Boh., Louny/Oblik 2.5.[19]62 A. Hoffner leg." (1 ɗ, 1 Ψ, NMP).- "Boh., Louny env. 30.IV. [19] 61 V. Balthasar leg." (4 ɗɗ, 5 ΨΨ, NMP).- "Závist, 21… Zeman" (1 ɗ, NMP).- "Env. de Praque, Bohême centr., Coll. Obenberger" (1 ɗ, NMP).- "Radotin 21.5.1927 " (1 ɗ, NMP).- " Bohemia, Jos. Cêrný | Pardubice 18.5.1909 | Mus. Pragensde, coll. Kracik” (1 Ψ, NMP).- "Moravia-1934, Veseli n[ad]. Mor[avon]. VI. [leg.] O. Kodym" (1 Ψ, NMP).- "Prossnitz [= Prostĕjov], [leg.] K. Kyselý” (1 ɗ, NMP). Slovakia: "Bratislava, Petržalka, [leg.] Roland" (1 Ψ, NMP).- "Slov. 3-4.[19]39, Bratislava" (1 Ψ, NMP).- "Slov. mer., Stúrovo, 28.- 30.7.1955, V. Balthasar leg.” (1 ɗ, NMP).- " Slovakia merid., Štúrovo | Parkán | [leg.] O. Kavan, 24.4.[19]49" (2 ɗɗ, 4 ΨΨ, NMP).- " Slovakia 1935. Plavec[ký] Sv[ätý] Mikuláš 27.V., [leg.] O. Kodym" (1 ɗ, NMP).- dto., nur " 5.VI. " (1 ɗ, NMP).- " Slovakia merid., Kováčovské vrchy, O. Kavan| Kováčar, 27.4.[19]49" (1 ɗ, 5 ΨΨ, NMP).- " Slovakia merid., Kováčovské vrchy, O. Kavan | Helemba, 22.4.[19]49" (1 ɗ, NMP).- "Slovakei, Pannonisches Becken, Zádiel Umg. 200 m, 0 1.07.1970 leg. J. Hadiel" (1 ɗ, CHB).- "Zadiel 1.VII. [19]70" (1 ɗ, SMNS).- " Slovakia, Štúrovo-Bet.k., 29.4.[19]54, Kr. Pospišil leg." (1 ɗ, SMTD).- " Slovaquie, Košice V.[19]37 coll. Pospišil" (1 Ψ, SMTD).- " Slowakei, Silicka planina 12.- 22.7.1993, leg. Richter" (1 Ψ, CWR). Hungary: "Hungarie c. or., Pécz, J. Pradáč leg." (1 Ψ, NMP).- "Hungarie centr., Békéscsaba env., V.1947, lgt. Škpt Havelka” (1 Ψ, NMP).- "Ujpest, Ungarn | Mus. Pragense Coll. Kracik" (1 ɗ, 1 Ψ, NMP).- " Hungaria, Esztergom, E. Bokor" (1 Ψ, ZMHB).- " Ungarn " (1 ɗ, 1 Ψ, ZMHB).- " Hungaria R. Scholz. | Coll. Döttling" (1 Ψ, SMNS).- " Ungarn, Izshak, 21.7.[19]93, leg. E. Heise" (1 ɗ, CEH).- " Ungarn, Neusiedler See, Fertöuglak, 0 4.05.1999 leg. A. Skale" (1 Ψ, CAS).- " Ungarn, Prov. Fejer, Umgebung Aba, Handfang 23.V.-7.VI.1997, leg. F. Bahr" (1 ɗ, 1 Ψ, CHB).- "Ujpest Ungarn " (1 ɗ, 2 ΨΨ, SMTD).- " Ungarn, Halbinsel Tihany, Plattensee 2.6.[19]63, leg. G. Jaeschke" (1 ɗ, 1 Ψ, DEI).- " Ungarn, Buda-hegy, E. IV.1969, leg. Fritzsche" (2 ɗɗ, 4 ΨΨ, SMTD).- " Hungaria, Cegled, 02.05.[20]02 leg. W. Richter" (2 ɗɗ, 2 ΨΨ, CWR).- "HU Mecseknadasd, Baranya Megye 09.05.[20]03 leg. H. Müller" (1 ɗ, CHM).- " Ungarn | Kollektion Dr. Fuchs" (1 ɗ, CJS; 1 Ψ, NSMW).- " Ungarn, A. via, K. Andreas" (1 ɗ, NSMW).- " Ungarn 8.10.03 " (2 ɗɗ. SMTD).- " Ungarn | Coll. Leonhard" (1 ɗ, 3 ΨΨ, DEI).- "Dunakaesz, Hungaria, ex Coll. Missionshaus SVD Steyl, Eing. Nr. 3/59" (1 ɗ, ZMFK). Switzerland: "Martigny | coll. Stierlin" (3 ɗɗ, 2 ΨΨ, DEI). France: "Reims" (1 ɗ, 1 Ψ, ZMHB).- "Bretigny s/Orge [Bretigny-sur-Orge, südlich Paris] 9/5/1963 [leg. Bause]" (10 ɗɗ, 9 ΨΨ, ZMHB).- "Toul, 6.[19]39" (2 ɗɗ, 4 ΨΨ, ZMHB).- " Frankreich, Bordeaux 4.[19]43, F. Ermisch lgt." (1 ɗ, 3 ΨΨ, ZMHB).- "St. Remy les Chevreuses, 11/5/[19]63" (2 ɗɗ, ZMHB).- "Provins (S-et M.) 22 Avr. 1946 Crottin de Cheval" (1 ɗ, SMNS).- dto., nur " 11. Mai 47 " (1 Ψ, SMNS).- " Frankreich St. Denis 6.1974" (1 ɗ, 1 Ψ, SMNS).- "Avord-Cher 18.6.1941 Folwaczny" (1 ɗ, SMNS).- " Gallia / Ardeche, Vernoux/ Valence, E.V.[19]74, Messulat" (1 Ψ, CJS).- "Etaples | Frankreich " (1 ɗ, SMTD).- "Metz 5.1916." (1 ɗ, 1 Ψ, SMTD).- "Boulogne" (1 ɗ, coll. Andrewes, NHML).- "Manche, M. St. Michel, 17.5.64 " (1 Ψ, NHML).- "S. France: (Var), La Sainte Baume. 16–28.V.1921. K. G. Blair." (3 ɗɗ, 1 Ψ, NHML).- "Environs de Bordeaux, de Vau log er " (1 ɗ, 1 Ψ, coll. Nevinson, NHML).- "Bordeaux" (1 ɗ, coll Nevinson, NHML).- "Moulins" (1 ɗ, coll. Nevinson, NHML).- "Nancy M." (1 ɗ, coll. Andrewes, NHML).- " Gallia Merid" (1 ɗ, coll. Fry, NHML).- "Gall. cent." (1 ɗ, coll. Nevinson, NHML).- " France " (1 ɗ, coll. Nevinson, NHML).- " Gallia " (1 ɗ, coll. Fry, NHML).- " Gallia Mulst" (2 ɗɗ, SMTD).- "Indre-et-Loire: Pernay, 2.v.1964, J.Rabaça legit" (2 ΨΨ, CTB).- "Jura: Arc-Senans, Les Communaux, 20.v.1956 (ex coll. Pasquier)" (1 ɗ, CTB).- "Jura: Les Arsures, 3.vi.1956 (ex coll. Pasquier)" (2 ɗɗ, 1 Ψ, CTB).- "Mayenne: Couptrain, P. Rodary legit" (1 Ψ, CTB).- "Saôneet-Loire: La Selle, iv.1950, P.Rodary legit" (1 ɗ, CTB).- "Grasse, A.M. ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 ɗ, 1 Ψ, ZMFK).- "La Salvetat (Hérault), ex Slg. R. Oberthür (Coll. Tondu) Eing.Nr. 4.1956" (1 ɗ, ZMFK).- "Lyon, ex Slg. R. Oberthür (Coll. Tondu) Eing.Nr. 4.1956" (1 ɗ, ZMFK).- "Wassy, Hte Marne, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 ɗ, ZMFK).- "Camarque, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 Ψ, ZMFK).- "Pontavert-France 27.4.1944, [leg.] Hans Pscher" (1 Ψ, NMP).- "Dpt. d’Or, Ende Mai 1898 | Riedel" (4 ɗɗ, 3 ΨΨ, NMP).- "Pyren. or., Dr. Balthasar” (1 ɗ, NMP). Great Britain: "Chesil 1864 J. Gray" (1 ɗ, SMTD).- "Hunts: Woodwalton Fen. 6.vi.1938 E. A. Duffy" (1 ɗ, NHML).- "W Pontrilas, 29.XI.1929 C. E. T." (1 ɗ, coll. C. E. Tottenham, NHML).- "Benfleet 28-iv-1930 C. E. T." (2 ɗɗ, 2 ΨΨ, coll. C. E. Tottenham, NHML).- "Bexhill 16.iv.1923 C. E. T." (2 ɗɗ, coll. C. E. Tottenham, NHML).- dto., nur " V.1923 " (2 ΨΨ, coll. C. E. Tottenham, NHML).- "Chichester 9.V.1919 C. E. T." (4 ɗɗ, 2 ΨΨ, coll. C. E. Tottenham, NHML).- "Warth Dorset or grass 6-VI-37 " (1 Ψ, NHML).- "Kent: Dartford, Marshes. 19.IV.1952. J. F. Burton, Cowdung" (1 Ψ, NHML).- "Somerset" (3 ɗɗ, coll. Andrewes, NHML).- "Cookham" (3 ɗɗ, 3 ΨΨ, NHML).- " England. On Cowdung. Totton, Hants,? Aug. 1953. SU 355127. G. R. Vardy" (1 Ψ, NHML).- " England: Dorset (09), Ballard Down, SZ0280, 16.iv.2007 ex horse dung, shaded, coll. D. J. Mann, OUMNH-2007-045" (1 ɗ, 1 Ψ, NHML).- "Power. Hampstead. 53.7" (2 ɗɗ, coll. W. A. Power, NHML).- "Power. Gravesend. 69.20" (1 ɗ, coll. W. A. Power, NHML).- "Power. Barnes. 69.18" (1 ɗ, NHML).- "Weymouth, April 21-1901." (2 ɗɗ, 2 ΨΨ, coll. Donisthorpe, NHML).- "Isle of Wight (W Brook), June 5.1899" (1 ɗ, coll. Donisthorpe, NHML).- "Rodwell, N. Weymouth, April 10-1896 " (1 Ψ, coll. Donisthorpe, NHML).- "Chingford VI.1911 " (1 ɗ, coll. H. C. Dollman, NHML).- "Etching m-6.24" (1 ɗ, coll. C. J. Saunders, NHML).- "Ryde" (1 ɗ, 4 ΨΨ, coll. D. Sharp, NHML).- "Edenbridge. June 1890 " (1 ɗ, coll. D. Sharp, NHML).- "Isle of Sheppey. G. C. C." (1 Ψ, coll. G. C. Champion, NHML).- "London distr. G. C. C." (2 ɗɗ, 3 ΨΨ, coll. G. C. Champion, NHML).- "Abbey Wood. T. W." (1 ɗ, coll. T. Wood, NHML).- "Padstow 1.V.1922 S. B." (1 ɗ, 1 Ψ, coll. C. E. Tottenham, NHML).- "Barcombe 2.4.27 flood." (1 ɗ, coll. C. J. Saunders, NHML).- dto., nur " 24.10.25 " (1 ɗ, coll. C. J. Saunders, NHML).- "Brighton 15.5. 38 L. A. Cox" (3 ɗɗ, 1 Ψ, coll. C. J. Saunders, NHML).- " Anglia " (1 ɗ, 1 Ψ, coll. Fry, NHML). Portugal: "P, Ribatejo, UG Santarem, S. de Aire, Kuh, 500 NN, 9.5.1991, leg. U. Schaffrath" (1 Ψ, CUS).- "Monforte: Herdade das Esquilas, 20.v.1997, A. Zuzarte legit" (1 ɗ, CTB).- "Resende: S. Cipriano, 22.vi.1978, J. Ramalho legit" (1 Ψ, CTB).- dto., nur " 21.vi.1979 " (5 ɗɗ, 4 ΨΨ, CTB). Azores: "Azores | Coll. Felsche, Kauf 20.1918" (2 ΨΨ, SMTD). Spain: "29.6.[19]92 (5) Navarra, Valle de Roncal, Garde, Rinderkot, Fery leg." (1 ɗ, 2 ΨΨ, CHF).- "E. Navarra, Sierra de Aralar 1000 m, 23.5.[19]96 leg. Strke" (2 ΨΨ, SMNS).- "E – Navarra, Sierra de Aralar, 700 m, S. Miguel de Aralar, 23.5.1996 Starke leg." (1 ɗ, 1 Ψ, CJS).- " Spanien, Morella, Prov. Castellon de la Plana, 1000 m, 19./ 20.VI.1992, leg. E. Rößner" (1 ɗ, CER).- "15.6.[19]90 Espana, Prov. Teruel, S. d.

Albarricin, Orihuela del Tremedal 1500 m, Schafskot, Fery leg." (1 Ψ, CHF).- "E.-Teruel, Albarracin, 15.- 17.4.2004, K. Hadulla leg." (1 Ψ, CKHa).- "2.4.[19]88 Espana, Prov. Madrid, Sierra des Guadarrama, Lozoya, Rinderkot, Fery leg." (1 ɗ, 3 ΨΨ, CHF).- "S[an]. Ildefonso" (1 Ψ, SMTD).- "Lisitan. Dyrell." (3 ɗɗ, 1 Ψ, coll. Heyden, DEI).- "Serra de Gredos, La Serrada/ Span. VI/ VII.1060, J. Vives leg." (6 ɗɗ, 3 ΨΨ, ZSM).- "Palencia, Pag[...]" (1 ɗ, ZSM).- "Sa[ntibáñez]. de Bejar, Spain, G. C. C." (1 Ψ, coll. G. C. Champion, NHML).- " Spain, Avila to El Barraco, 1400 m (30T359999 4487524) 21- 22.6.2008 " (3 ɗɗ, ex coll. A. Pizzo, NHML). Italy: " 4.5.1991 No. Italien, Mt. Giorgio bei Savona (700 m) leg: Ahrens" (2 ɗɗ, 3 ΨΨ, CDA).- "Alp. Mar. piemont., Limone, 1000–1200 m, Tetti Camila, 20-VI-1951 [leg.] G. Fagel” (1 Ψ, NMP).- " Italien 29/VI 24, K: 3411" (4 ɗɗ, 1 Ψ, NMP).- "16.6.[19]83 Toscana, Chinsi, Schafskot, Fery leg." (1 Ψ, CHF).- "Castel di Sangro, Prov. di Aquila, Paganetti" (1 Ψ, ZMHB).- "6.5.[19]90 Italia, Puglia, Umg. Laterza, S Bari, Schafskot, Fery leg." (1ɗ, CHF).- "Dr. Feige, Roma, Monte Cavo" (2 ΨΨ, ZMHB).- "20.6.[19]83 Calabria, Gambarie, Schafskot, 1000 m, Fery leg." (1 Ψ, CHF).- "27.6.[19]83 Calabria, Sila, Germano 1000 m, Rinderkot, Fery leg." (1 Ψ, CHF).- "Gargano, Ital./ IV.[19]75, leg. Niehuis" (1 ɗ, CJS).- " Insel Elba, Monte Capanne, 9.4.[19]54, Dr. David" (1 Ψ, ZMHB).- "Calabrien. Cosenza 6.6.[19]43" (1 Ψ, SMNS).- "Calabria – Asprom. Cittanova (RC) 1300 m, Zomaro, 23.VIII. [19]81 leg. F. Angelini" (1 Ψ, SMNS).- "Liguria, Testico 11.6.[19] 62 W. Liebmann" (1 ɗ, SMNS).- "Südtirol, Staben, Sonnenhang, 20.-23.5.[19]72 Rieger" (1 Ψ, SMNS).- " Italien, Lazio, Colle Albani, Lariano, C. Pacione, 9.IV.1999 lg. Ahrens, Fabrizi" (10 ɗɗ, 5 ΨΨ, SMTD).- " Italia, Lazio, Colli Albani: Vivaro 1 km E of Nemi, 5.V.1999 leg. S. Fabrizi" (2 ɗɗ, 5 ΨΨ, SMTD).- "Lombardia Italia " (7 ΨΨ, SMTD).- "Calabria, Antonimina 1905, Paganetti" (4 ɗɗ, 2 ΨΨ, DEI).- "Calabria, Aspromonte, lg. Paganetti" (1 ɗ, DEI).- "San Basilio, Murgien, Paganetti" (2 ΨΨ, DEI).- "Apulien, Mte. Gargano" (1 ɗ, SMNS).- "Monte Gargano 14.6.1907 legit M. Hilf, Coll. O. Leonhard" (1 Ψ, DEI).- "Pisa" (1 Ψ, ZSM).- "Roma" (1 Ψ, ZSM).- "Firenze: M. Giovi, 4.v.1968, S.Rocchi legit" (3 ɗɗ, 2 ΨΨ, CTB).- "Firenze: Pratolino, 2.v.1970, S. Rocchi legit" (8 ɗɗ, 11 ΨΨ, CTB).- "Firenze: Vallombrosa, Secchieta, 20.vi.1970, S. Rocchi legit" (1 Ψ, CTB).- "Ligúria: Savona, Mte. S. Giorgio, 20.iv.1965, G.Dellacasa legit" (1 ɗ, 2 ΨΨ, CTB).- "Ligúria: Savona, Mte. S. Giorgio, 17.v.1965, L.Briganti legit" (2 ɗɗ, 1 Ψ, CTB).- " Italy, Colle dell’Orso (CB) (E450310, N4607162), 1400 m 09.06.2007 " (2 ɗɗ, ex coll. A. Pizzo, NHML).- " Italy, Pantano della Zottola, Iserina (E424539, N4619338), 13.06.2007 " (2 ɗɗ, 2 ΨΨ, ex coll. A. Pizzo, NHML).- " Italy, Lazio: Monti Aurunci, Campodimele env., 824 m 41°22’30,1’’ N, 13°32’26,3’’ E, 31.V.2008 Ahrens & Fabrizi" (1 Ψ, NHML).- " Italy, Altopiano di Monte: Lago Sefro (MC), 910 m, 43°07’44,41’’ N, 12°58’01,10’’ E, 01.06.2008 " (4 ɗɗ, ex coll. A. Pizzo, NHML).- " Italy Torino (Ipla) (N400) 23.V.2007 " (2 ɗɗ, ex coll. A. Pizzo, NHML).- "Sicilien, Palermo, Ex c. Dr. Studt G." (1 ɗ, ZMHB).- "NO-Sizilien, Le Madonie, Piano Zucchi 1000 m, 23.5.2006, W. Ziegler" (4 ΨΨ, CWZ).- "I – Sizilien, Madonie, Piano Zucci 22.5.2006, Heise leg." (2 ɗɗ, 1 Ψ, CEH).- "I – Sizilien, Madonie, Piano Battaglia 23.5.2006, Heise leg." (1 ɗ, CEH).- "18.4.[19]84 Sicilia, Buccheri, Rinderkot, Fery leg." (1 ɗ, CHF).- "25.3.[19]83 Sicilia, Collesano, 400 m, Rinderkot, Fery leg." (1 Ψ, CHF).- "25.3.[19]83 Sicilia, Campofehice, 400 m, Rinderkot, Fery leg." (2 ɗɗ, CHF).- "Sicilien Messina. 25.4.[19]43" (5 ɗɗ, 8 ΨΨ, SMNS).- "Sicilien, Messina, 25.4.[19]43, W. Liebmann" (1 Ψ, DEI).- "Sicilia Italia 22.4.[19]33, W. Liebmann" (1 ɗ, DEI).- "Sicilia, Randazzo, 3.5.[19]33, W. Liebmann" (11 ɗɗ, 4 ΨΨ, DEI).- "Sizilien" (1 Ψ, SMTD).- "Sic., Palermo, Dr. Balthasar" (1 Ψ, NMP). Slovenia: "Adelsberg [= Postojna] 13.5.[19]09" (2 ɗɗ, 1 Ψ, SMTD).- "Marbg. [= Maribor], [leg.] St. Dr. Krause, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (1 Ψ, ZMFK). Croatia: "Jug., Kroatia, Istria or., Rabac, 15.–17. v. 1890 Wrase" (2 ΨΨ, CER).- "No 164 Istrien, San Pietro i.S. 4.6.1911, Schumacher-Spaney S.V." (1 ɗ, ZMHB).- "No. 53 Karst, Pared 450 m, Spaney 29.5.[19]11, Schumacher S.V." (1 ɗ, ZMHB).- "No. 123 Karst, Divača-Corgnale 450 m, 31.5.[19]11, Spaney-Schumacher S.V." (8 ɗɗ, 1 Ψ, ZMHB).- "No. 26 Karst, Občina 350 m. 28.5.[19]11, Spaney-Schumacher S.V." (1 ɗ, ZMHB).- "YU / Istrien, Rovin E.5.[19]79 leg. Heidenfelder" (3 ɗɗ, 1 Ψ, SMNS).- "Istrien, Pula Umg. 28.5.[19]74 leg. Oschütz" (1 ɗ, SMNS).- "Jugosl./ Istrien, Umag, E.3.[19]78" (1 Ψ, CJS).- "19.5.[19]37, Jugoslavia, Dalmatien | Crik-Veniza, in Dung | 599 leg. Pause" (1 ɗ, SMTD).- "Istria Noghera" (1 Ψ, SMTD).- "Meusel, Croatia | Coll. Schenkling" (1 ɗ, 1 Ψ, DEI).- " Croatia, coll. Künnemann" (1 Ψ, DEI).- "Istria Rse Ver." (1 ɗ, 1 Ψ, coll. Heyden, DEI).- "Fijume [= Rijeka]" (1 Ψ, ZSM).- "Istria." (2 ɗɗ, ZSM).- " Kroatien, Halbinsel Istrien, Rovinj, 14.5.1980, leg. F. Bahr" (1 ɗ, CLS).- "Pola, Istr., [leg.] Fr. Lang, ex Coll. Missionshaus SVD Steyl, Eing. Nr. 3/59" (1 ɗ, 3 ΨΨ, ZMFK).- "Dalm., VI.[19]37, Knin, Bal.-Fr." (1 ɗ, NMP).- "Dalmatia" (1 ɗ, 1 Ψ, NMP).- " Kroatien, Porec, 21.05.1988, leg. K. Hadulla" (2 ɗɗ, 4 ΨΨ, CKHa). Bosnia-Herzegovina: " Bosna, Korična, O. Leonhard" (1 ɗ, DEI).- " Bosnia " (1 ɗ, CJS). Montenegro: "No 451 Montenegro, Vojnik-Gebirge, Brezovi-do 1000 m, 27.6.[19]11 Schumacher S.V." (3 ɗɗ, 1 Ψ, ZMHB).- "Castelnuovo [= Herceg-Novi], [leg.] Hummler" (1 ɗ, ZMHB).- "Dalm. Castelnuovo M. Hilf 1910, Coll. O. Leonhard" (1 Ψ, DEI).- "Petrovac na Moru, Montenegro, April 1969, leg. E. Gersdorf" (1 Ψ, CJS). Serbia: "Belgrad Serbia " (1 Ψ, SMTD).- "Maglič VIII.1913, Montenegro, legit Jan Obenberger" (1 Ψ, NMP).- "Sutorina, Paganetti" (1 Ψ, NMP). Macedonia: "YU Maked., Galicica 1600 m, 20.V.1982, Krätschmer leg." (1 Ψ, CJS). Ukraine: "Olgino b. Berdjansk, Dnjepr. Gebiet, V/ VI 1937, Ukraine, leg. Nikolajew" (15 ɗɗ, 15 ΨΨ, ZMHB).- " Ukraine, Dnjeprop. obl., Umg. Berdjansk, leg. Nikolajew, 14.VI.1937 " (1 Ψ, ZMHB).- "Umg. Berdjansk, Dnjeprop. obl., 30.V.1938, leg. Gulikow" (3 ΨΨ, ZMHB).- " Ukraine, 24.4.1937, Mekortj. b. Berdjansk, Dnjepropetr. Gebiet, leg. Nikolajew" (6 ΨΨ, ZMHB).- " Ukraine, Kamienka, 35 m s. v. Rowenki [= Rovenki], VI.1918, A. Spaney" (1 ɗ, 2 ΨΨ, ZMHB).- "Süd- Russland, Falzfeinowo a. Dniepr 12.V.–7.VI.1914, W. Ramme S.G." (5 ɗɗ, 1 Ψ, ZMHB).- " Ukraine, 9.6.1935, Chomutowsk. Steppe, b. Mariupol, leg. Porischenko" (1 ɗ, ZMHB).- "Krim, Simferopol, 24.V.1942, Haltenorth S.G." (1 ɗ, 1 Ψ, ZMHB).- "Odessa" (1 ɗ, SMTD).- "Crimea" (2 ɗɗ, NHML).- "Sebastopol Krim 10.4.1903 [leg.] W. Pligiuski" (1 Ψ, NMP). Russia: "Rostov na Donu, R. m.” (1 Ψ, NMP).- "Südrussland, Sarepta | H. Rolle Berlin | ex coll. Hopffgarten Mülverstedt." (1 Ψ, NMP).- "Terskaja obl., St. Mineralni Wod., 30.IV.1928 " (13 ɗɗ, 1 Ψ, ZMHB).- "Dongebiet, 28.V.–3.VI.1918, A. Spaney" (1 Ψ, ZMHB).- "S. O. Russland, Tschernyj-Jar, Wolga, Kracht S.G." (1ɗ, ZMHB).- "Daghestan ( Rossia), Nowyi Birjusjak, leg. Worobjew, 6.6.[19]57" (1 ɗ, ZMHB).- "S.E. Russia, Volsk district. R. S. Kozlov. B. M. 1931- 39 " (2 ɗɗ, 2 ΨΨ, NHML).- "Transcaspia, Neu-Saratow" (1 Ψ, NHML).- "Kaukas. Schneider" (1 Ψ, SMTD).- "Caucasus, Alexandrofska, Ryssel S.V. / 18.4.[19]01" (3 ɗɗ, ZMHB).- "Caucasus" (1 Ψ, NHML; 1 ɗ, SMNS). Kazakhstan: "Indersk." (2 ɗɗ, 1 Ψ, coll. E. Y. Western, NHML). Romania: "Desert. Deliblát, Banat, Holtz" (4 ΨΨ, ZMHB).- " Rumänien, Donaudelta, Unirea [bei Brăila], 11.VI.1988 leg. Hornburg" (1 ɗ, 3 ΨΨ, COH).- " Roumanie, Comana Vlasca, A. L. Montandon" (1 ɗ, DEI).- "Dobroudja. Mangalia. A. L. Montandon" (1 ɗ, DEI).- "Jassy [= Iasi]" (3 ɗɗ, ZSM).- "Bukarest" (1 ɗ, ZSM). Bulgaria: "Mandra, Bulgarien, 5.5.1933, leg. Fuss" (1 ɗ, NMP).- " Bulgarien, Kitschevo, N Varna 28.V.1995, leg. H. Kalz" (1 Ψ, CHKa).- "BG, Sveti Vlas, 16.V.2005, leg. H. Müller" (3 ɗɗ, 6 ΨΨ, CHM).- " Bulgaria, Sveti Vlas, 08.05.- 19.05.[20]05 leg. W. Richter" (6 ɗɗ, 2 ΨΨ, CWR).- " Bulgarien, Umg. Sozopol, 9.6.- 20.6.1978, leg. Bringmann" (1 Ψ, CER).- "SO-Bulgarien, südl. Sosopol, Umg. Kavacite, 30. Mai 1982, leg. H. Wendt" (1 ɗ, ZMHB).- "SO-Bulgarien, südl. Burgas, Umg. Acheloi, 1./ 2.VI.1982, leg. H. Wendt" (1 Ψ, ZMHB).- " Bulgarien, U. Nessebar, 4.6.[19]67, K. Ermisch leg." (1 Ψ, ZMHB).- " Bulgarien, Primorsko, 12.6.[19]85, leg. Kriska" (1ɗ, CER).- "SO-Bulgarien: Zarewo (= Mičurin), 13.V.2002, leg. E. Rößner" (1 Ψ, CER).- "SO- Bulgarien, Strandža-Gebirge: Malko Tarnovo, 14.V.2002, leg. E. Rößner" (1ɗ, CER).- "Sofia" (1 Ψ, SMTD).- " Bulgaria, 55 km SE Burgas, Lozenec 13.V.2002, leg. L. Schmidt" (1 ɗ, 1 Ψ, CLS).- " Bulgaria, 35 km NW Burgas, 5 km W Ajtos, Trockenhang, 18.V.2002, leg. L. Schmidt" (1 ɗ, 1 Ψ, CLS).- " Bulgaria, 37 km SE Burgas, Jasna poljana, Viehweide, 17.V.2002, l. L. Schmidt" (2 ɗɗ, CLS).- "Mandra, Bulgaria, 20.5.1933, Fuss leg." (1 ɗ, 4 ΨΨ, ZMFK). Greece: " Graecia, L. Salvator." (1 ɗ, NMP).- "Ossa, Graecia " (1 ɗ, 1 Ψ, NMP).- " Greece (C. Macedonia) 258 m, Lake Kerkini env., Akitoschori N 41°16’72,5’’/E 023°10’15,3’’, 15.IV.2003 leg. O. Hillert" (1 ɗ, 1 Ψ, COH).- " Griechenland, Prionia Gebirge, Olymp, 1000 m, 27.VI.- 17.VII.1995, leg. T. Kriska" (1 ɗ, CER).- "12.4.[19] 84 N. Griech. westl. Kavalla, 300 m, Schafskot, Fery leg." (1 Ψ, CHF).- "13.4.[19]84 Griech., Larissa, Schafskot, Fery leg." (1 ɗ, CHF).- GR, Makedonien, Nomós Kavála, 10 km N Pale á Kavála, 600 m, Macchie + Kastanienwald 20.IV.1994, leg. Schawaller" (1 Ψ, SMNS).- "GR, Ossa, Oros, 500 m, Stomio, 27.5.1990, Persohn leg." (1 ɗ, SMNS).- "GR – Thessalie, Stomio 5.7.2006, Heise leg." (1 ɗ, CEH).- "GR; 5 km ö Kato Vermion, 1000 m, Baehr 1.5.[19]83" (2 ΨΨ, CJS).- "GR Vermion Oros, Kato Vermion 1300 m, Krätschmer lg. IV.1977 " (1 ɗ, CJS).- " Graecia / Thessalia, Ossa Oros/ Stomio 1000 m / 27.V.1990, Persohn leg." (1 ɗ, 1 Ψ, CJS).- "GR: 10 km w. Veria. 700 m. Baehr 30.4.[19]83" (1 ɗ, CJS).- " Greece / Monodéndri, Vikou-Gebirge 03.–06.V.1993 leg. Ringel" (1 Ψ, CHR).– "GR, Peloponnes, 1400 m, Exohori, Prov. Messinia, 2.7.2005, leg. W. Ziegler" (2 ɗɗ, 2 ΨΨ, CWZ).- " Griechenland, Peloponnesos: Araxos (30 km w Pátrai), 26.–31.V.1993, leg. E. Rößner" (1ɗ, CER).- "Balkan, Corfu. Paganetti [19]03" (1 ɗ, DEI).- "16.4.[19]84 Corfu, Roda, Pferdekot, Fery leg." (1 ɗ, 1 Ψ, CHF).- "Rhodes" (1 ɗ, NMP). Turkey: "Europ. Türkei: Istranca-dag bei Demirköy, Paß, 700 m, Heinz leg. 2.V.1973 " (1 Ψ, CJS).- "Constantinopel V.1898, W. Gottwald S.V." (1 ɗ, ZMHB).- "Kleinasien, Biga, 22.4.[19]16, Dr. Bauer S.G." (1 ɗ, ZMHB).- " Asia minor , Biledjek, v. Bodemeyer" (1ɗ, ZMHB).- " Angora. Asia min., V. M. Duchon" (4 ɗɗ, 2 ΨΨ, ZMHB).- " Türkei (Centr.), Puturgi bei Malatya, 10.VI.1993, leg. Kriska" (1 ɗ, COH).- " Turcia m., Prov. Antalya, 20 km NE Demirtas, Hochebene 1400 m, 36°31’32’’ N, 32°23’26’’ E, 28.V.2006, leg. A Kopetz" (2 ΨΨ, CAK).- " Türkei, Anatolien, Nohutlu (90 km sö von Malatya), ca. 1700 m, 5.– 9.VI.1993, leg. A. Schröder" (3 ɗɗ, 3 ΨΨ, CER).- "Ararat, Wagn[er]." (1 ɗ, ZMHB).- "Tauria, Part. | 26959" (1 ɗ, ZMHB).- "Kleinasien, Anatolien | Kollektion Dr. Fuchs" (1 Ψ, CJS).- " O. Türkei 20.–26.VI. [19]75, 30– 40 km ö Bingöl, leg. D. Berhauer" (1 Ψ, CJS).- "Kleinasien, Anatolien | Kollektion Dr. Fuchs" (3 ɗɗ, 3 ΨΨ, NSMW).- "v. Bodemeyer, Klein-Asien, Eski-Chehir [= Eskişehir] | Kollektion Dr. Fuchs" (1 ɗ, NSMW).- " Asia minor , Burna, v. Bodemeyer | Kollektion Dr. Fuchs" (1 Ψ, NSMW).- " Asia minor , Sultan Dagh, v. Bodemeyer" (1 ɗ, NMP).- "Klein-Asien, Eski-Chehir, v. Bodemeyer" (2 ɗɗ, NMP).- "Eski Chehir, Coll. Purkynĕ." (1 ɗ, NMP).- " Asia minor , Ak-Chehir, v. Bodemeyer” (1 ɗ, 1 Ψ, NMP).- "v. Bodemeyer, Anatolia, Asia minor ” (1 ɗ, NMP).- " Anatolia, Asie Mineure, Coll. Obenberger” (3 ɗɗ, 3 ΨΨ, NMP).- "v. Bodemeyer, Klein Asien, Bulghar Maaden” (1 ɗ, 1 Ψ, NMP). Georgia: "Borshom Caucasus, O. Schneider" (1 Ψ, coll. Heyden, DEI). Iran: "N. Iran 2.VI. [19]77, 40 km s Sari | Elb., leg. D. Bernhauer" (1 ɗ, CJS).- " Persia Astrabad [= Gorgan]" (4 ɗɗ, 4 ΨΨ, SMTD).- "N-Iran, Prov. Teheran, C-Elburz, Damavand Mt. Gosfand Sara, Larijan, 35° 52’ N 52° 07’ E, 2300 m, 29.05.2008, leg. A. Skale (11a)" (4 ɗɗ, 3 ΨΨ, CAS).- "Elbrus Geb., Persien, Rttr." (2 ɗɗ, NMP).- " Persien, Luristan, v. Bodemeyer" (2 ɗɗ, NMP). Azerbaijan: "Lenkoran, Dr. Martin, ex Slg. R. Oberthür (Coll. Tondu) Eing. Nr. 4.1956" (2 ΨΨ, ZMFK).

Not localised: "L. Chamboy, 8.8.38 " (1ɗ, ZMHB).- "Transcaspien | Coll. Döttling" (1 Ψ, SMNS).- "Wan 22.4.12 " (1 ɗ, 1 Ψ, ZSM).- "Kasp. Meer-Geb., Petrowsk, 1897 Korb" (1 ɗ, ZSM).

Redescription. Body length: 8–13 mm, body oval, weakly convex, dull; head and pronotum black with dark green metallic sheen, rarely cupreous or intensively metallic green; elytra yellow with rows of asymmetric dark spots sometimes arranged in stripes; ventral face, pygidium and legs black with very weak metallic sheen, rarely cupreous; dorsal pilosity short and light.

Head. Antenna brown, clavus black; anterior edge of clypeus reflexed, with transverse wrinkles, head coarsely punctate to granulate between frontal carina and vertex.

Pronotum. Lateral margins behind anterior angles straight or convex, anterior angles weakly rounded, posterior angles sinuate, basis with complete and distinct marginal line (Fig. 2AJ); punctation coarse, dense, very dense and granulate beside anterior angles, on midline and beside posterior angles with narrow smooth impunctate sections.

Elytra. Yellow, base narrowly dark, sutural interval widely or completely dark with green sheen, epipleura completely light; spots black without or with only very weak green sheen, asymmetric (Fig. 2I, 2J), base always without spots, spots mostly confluent (Figs. 2K–M), sometimes condensed into symmetric stripes (Figs. 2N, 2 O), or covering the intervals almost entirely (Fig. 2 O); intervals chagreened, punctation even, granulate; striae shallow, weakly punctate, 7th stria weakly curved.

Pygidium. Base with marginal line, dull, punctation fine, moderately dense and simple, superficial.

Aedeagus. Figs. 3A, 3B, 3Q–AB; lamella copulatrix: 3AG–AK, 3AP–AT (see Table 1 ).

Remarks. We searched for the collection J. G. Kugelann (1753–1815) to find the type specimens of his Copris medius . According to Horn et al. (1990: 218) his beetle collection was passed to Hoffmann at Warsaw. The collection Kugelann is currently preserved at the Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw. According to information of the curator, Mrs. D. Mierzwa (personal communication, 2008), there is no specimen in the collection under either the name ' Copris [ Onthophagus ] medius ' or O. vacca . Consequently it must be assumed that the type material was lost or destroyed. For an unambiguous definition of O. medius and in order to differenciate between the two European species of the O. vacca complex, we designate here a neotype for O. medius . We chose for neotype a male from Poland (Lyck = Ełk).

Male, major form: Reflexed portion of anterior edge of clypeus bluntly truncate, rarely weakly sinuated (Fig. 2Q); frontal carina absent, impunctate on this spot; carina of vertex with large lamella with 1st sides basally parallel, apically with a sharply pointed horn being almost rectagular erect and with s-shape arch (lateral view) (Fig. 2AF–AG); Pronotum on anterior declivity smooth, medially with a deep impression, marginal line of base beside posterior angles complete (Fig. 2AJ).

Male, minor form: Anterior edge of clypeus sinuate (Fig. 2S); frontal carina weakly indicated to slightly elevate, curved; carina of vertex with a small lamella being slightly elevated at middle; pronotum anteriorly with rugose punctures, medially with a pair of small, smooth convexities, marginal line of base beside posterior angles complete.

Female, major form: Anterior edge of clypeus weakly sinuate; frontal carina distinct, elevate, weakly curved; carina of vertex low, as wide as entire posterior edge of head, each end blunt and weakly elevated (Figs. 2X, 2AC); pronotum at anterior marginal declivity widely smooth and only sparsely punctate, medially with a slightly anteriorly produced and medially slightly incised convexity, beside it on each side with a weak convexity, marginal line of base beside posterior angles complete (Fig. 2AJ).

Female, minor form: Anterior edge of clypeus weakly sinuate; frontal carina short, elevate; carina of vertex wide and low, without or only with slightly indicated elevated ends (Figs. 2 Y, 2AE); pronotum anteriorly with rugose punctures, medially with a slightly transeverse smooth convexity, basal marginal line complete.