Kamyristi, Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

publication ID

https://dx.doi.org/10.3897/asp.82.e111148

publication LSID

lsid:zoobank.org:pub:43AC036E-93CC-4D79-939A-07DF54BE1A2D

persistent identifier

https://treatment.plazi.org/id/62D92992-92DA-47CE-B987-C8BD825EBBF8

taxon LSID

lsid:zoobank.org:act:62D92992-92DA-47CE-B987-C8BD825EBBF8

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Kamyristi
status

gen. nov.

3.2.1.1. Genus † Kamyristi gen. nov.

Type species.

Kamyristi exfrigore sp. nov.

Etymology.

Anagram of Taïmyrski, the Russian name of the Taimyr peninsula from where the amber pieces containing the specimens derive. Gender masculine.

Diagnosis.

As for tribe with the following additional characters: head shagreened; compound eye higher than long; occipital carina foveate (Figs 9C and F View Figure 9 ); flagellomeres cylindrical and elongate. Mesosoma less than half of metasoma length; mesoscutum reduced on anterior part of mesosoma, hiding pronotum dorsally (Figs 9B and F View Figure 9 ). Fore wing with medial cell pentagonal with 1m-cu, Cu1 and M+Cu and Cu spectral, Rs+M not aligned with M+Cu (Figs 2G, H View Figure 2 , 9B and D View Figure 9 ). Hind leg with scattered erect setae; one mesotibial and one metatibial spur.

Comments.

Two species from this genus are known, both displaying numerous similarities with the extant genus Cryptalyra from South America, e.g., the elongate smooth metasoma (Fig. 9A View Figure 9 ), the erect setae on the hind leg (Fig. 9A View Figure 9 ) or M half pigmented (Figs 2G and H View Figure 2 ). But † Kamyristi exfrigore gen. et sp. nov. displays the most surprising affinities to Cryptalyra : the ovipositor sheaths seem very reduced, only covering the basal part of the ovipositor (Fig. 9A View Figure 9 ). However, the closed marginal cell (Fig. 2G and H View Figure 2 ), the elongate flagellomeres (Fig. 9E View Figure 9 ) and the mesosoma being shagreened instead of foveate (Fig. 9C View Figure 9 ) indicate that it is not closely related to Cryptalyra . Examination of more female specimens will be necessary to confirm if reduced ovipositor sheaths are a diagnostic character for this genus. † Kamyristi gen. nov. species are the smallest megalyrids in Asia.

Due to the fore wing venation and absence of a sulcus on the vertex, this new genus resembles Dinapsini with Rs reduced. Another difference is the posterior head sculpture, distinctly shagreened, whereas it is reticulate for the extant dinapsine species. Finally, the lack of grooves behind the eyes and of a dorsal carina on the subantennal groove makes the placement in Dinapsini doubtful; they are similarly absent in † Valaa delclosi . According to Shaw (1990a), these characters are relatively homoplasious. The placement of † Kamyristi gen. nov. in the Dinapsini is not corroborated by the phylogenetic analyses (see below).