Lasiochernes pilosus (Ellingsen, 1910)

Christophoryova, Jana, Krajcovicova, Katarina, Henderickx, Hans & Spaniel, Stanislav, 2016, A multivariate study of differentiating characters between three European species of the genus Lasiochernes Beier, 1932 (Pseudoscorpiones, Chernetidae), ZooKeys 629, pp. 51-81 : 59-61

Taxon classification Animalia Pseudoscorpiones Chernetidae

Lasiochernes pilosus (Ellingsen, 1910) Fig. 2D; Table 1

Description.

Female (7 specimens analyzed) (Table 1). Chaetotaxy of carapace: 81-96 setae, 49-63 of them situated on anterior disk, 17-25 on medial disk, posterior margin with 10-13 setae. Cheliceral galea with 6-8 short terminal rami, serrula exterior with 23-25 blades. Palps: fixed chelal finger with 44-49 and movable chelal finger with 44-49 marginal teeth; fixed chelal finger with 11-16 antiaxial and movable chelal finger with 11-15 antiaxial accessory teeth; fixed and movable chelal finger with 6-7 paraxial accessory teeth. Palpal parts without long and dense setation. Legs: tarsus IV with long tactile seta situated approximately in the middle of segment, that means 0.25-0.32 mm from the tarsal base. Chaetotaxy of tergites I–XI: 12-17 (left hemitergite 6-9 + right hemitergite 6-8): 15 (7-8 +7-8): 14-19 (8-9 + 6-10): 17-24 (7-11+ 9-13): 18-23 (9-11 + 9-12): 18-22 (8-12 + 8-11): 18-23 (8-12 + 9-11): 18-22 (9-11 + 9-11): 17-20 (8-11 + 7-10): 13-19 (6-10 + 6-9), tergite XI with 8 setae (4 + 4) and with a pair of long tactile setae. Chaetotaxy of sternites IV–XI: 8-18 (left hemisternite 4-10: right hemisternite 4-9): 17-25 (9-13 + 8-13): 19-28 (9-13 + 10-15): 19-28 (9-15 + 9-13): 17-27 (9-13 + 8-14): 18-26 (8-13 + 9-13): 17-22 (8-11 + 8-11), sternite XI with 8-14 (4-6 + 4-5) and with a pair of long tactile setae. Female spermatheca unpaired, T-shaped; anterior genital operculum with 29-44 setae and 1-2 lyrifissures, posterior operculum with 10-14 setae and 1-4 lyrifissures (Fig. 6E).

Male (5 specimens analyzed) (Fig. 2D, Table 1). Chaetotaxy of carapace: 77-89 setae, 47-57 of them on anterior disk, 18-23 on medial disk, posterior margin with 10-14 setae. Cheliceral galea with 6-7 short terminal rami, serrula exterior with 23-24 blades. Palps (Fig. 4C): fixed chelal finger with 40-50 and movable chelal finger with 41-51 marginal teeth; fixed chelal finger with 12-16 antiaxial and movable chelal finger with 13-15 antiaxial accessory teeth; fixed chelal finger with 6-7 paraxial and movable chelal finger with six paraxial accessory teeth. Palpal femur and patella with long and dense setation on their medial sides (Fig. 2D). Legs: tarsus IV with long tactile seta situated approximately in the middle of segment, that means 0.25-0.31 mm from the tarsal base. Chaetotaxy of tergites I–XI: 13-17 (left hemitergite 7-9 + right hemitergite 6-8): 14-16 (7-8 + 7-8): 15-22 (7-11 + 8-11): 18-24 (10-12 + 7-13): 19-24 (10-12 + 9-12): 18-22 (9-12 + 9-12): 16-22 (7-10 + 9-12): 17-22 (9-11 + 8-11): 15-19 (8-9 + 7-10): 10-15 (5-7 + 5-8), tergite XI with 8 setae (4 + 4) and with a pair of long tactile setae. Chaetotaxy of sternites IV–XI: 16-24 (left hemisternite 8-11 + right hemisternite 7-13): 17-26 (9-16 + 8-12): 17-31 (6-15 + 11-16): 14-30 (2-15 + 12-15): 22-29 (10-17 + 9-13): 19-27 (9-14 + 9-13): 16-22 (8-12 + 8-11), sternite XI with 8-12 (4-6 + 4-6) and with a pair of long tactile setae. Anterior genital operculum with 44-62 setae and 1-2 lyrifissures, posterior operculum with 19-26 setae and 2-6 lyrifissures (Fig. 6F).

Nymphs (Fig. 5; Table 1): The morphology of tritonymphs, deutonymphs and protonymphs is similar in most respects to that of adults (e.g. morphology of setae on body, granulation of carapace, cheliceral rallum of three blades, presence of venom apparatus in movable chelal finger (Fig. 5), presence of a pair of relatively long tactile setae on tergite XI and long tactile seta situated approximately in the middle of leg IV tarsus). Body measurements are given in Table 1.

Tritonymphs (15 specimens analyzed) (Table 1). Chaetotaxy of carapace: 71-87 setae, 43-52 of them situated on anterior disk, 17-25 on medial disk, posterior margin with 9-11 setae. Chelicera: five setae on hand, one on movable finger; galea with six short terminal rami, serrula exterior with 18-20 blades. Palps (Fig. 5A): seven trichobothria on fixed chelal finger and three on movable chelal finger; fixed chelal finger with 34-42 and movable chelal finger with 36-41 marginal teeth; fixed chelal finger with 8-11 antiaxial and movable chelal finger with 8-12 antiaxial accessory teeth; fixed chelal finger with 4-6 paraxial and movable chelal finger with 4-5 paraxial accessory teeth. Chaetotaxy of tergites I–X: 10-13 (left tergite half 5-6 + right tergite half 5-7): 10-12 (5-7 + 5-6): 10-14 (5-7 + 5-8): 11-17 (5-8 + 6-9): 12-17 (6-9 + 6-8): 11-17 (5-8 + 6-9): 13-18 (6-9 + 6-10): 12-17 (5-9 + 6-8): 11-15 (5-8 + 5-7): 9-12 (4-6 + 4-6), tergite XI with 6 setae (3 + 3) and a pair of long tactile setae. Chaetotaxy of sternites II–X: 4-12 (left hemisternite 2-6 + right hemisternite 2-6): 5-11 (2-6 + 3-6): 8-13 (4-7 + 3-7): 12-18 (5-9 + 5-10): 14-19 (7-10 + 7-10): 14-17 (7-10 + 5-9): 13-18 (6-10 + 7-10): 14-18 (7-9 + 7-9), 12-17 (6-8 + 6-9), sternite XI with 8-10 (4-5 + 4-5) and a pair of long tactile setae; sternites II with two lyrifissures.

Deutonymphs (15 specimens analyzed) (Table 1). Chaetotaxy of carapace: 44-58 setae, 28-34 of them ivesituated on anterior disk, 9-20 on medial disk, posterior margin with 6-8 setae. Chelicera: five setae on hand, one on movable finger; galea with 3-4 short terminal rami, serrula exterior with 17-19 blades. Palps (Fig. 5B): six trichobothria on fixed chelal finger and two on movable chelal finger; fixed chelal finger with 27-32 and movable chelal finger with 29-33 marginal teeth; fixed chelal finger with 5-7 antiaxial and movable chelal finger with 5-7 antiaxial accessory teeth; fixed chelal finger with 4-5 paraxial and movable chelal finger with three paraxial accessory teeth. Chaetotaxy of tergites I–X: 8-10 (left tergite half 4-5 + right tergite half 4-5): 7-10 (3-5 + 3-5): 6-10 (3-5 + 1-5): 9-10 (4-5 + 4-5): 9-10 (4-5 + 5): 7-10 (3-5 + 3-5): 9-10 (4-5 + 4-5): 9-10 (4-5 + 4-5): 8-10 (4-5 + 4-5): 4-9 (3-5 + 1-5), tergite XI with 4 setae (2 + 2) and a pair of long tactile setae. Chaetotaxy of sternites II–X: 0-1 (left hemisternite 0-0 + right hemisternite 0-1): 4-6 (2-3 + 2-3): 5-8 (2-4 + 2-4): 6-11 (3-6 + 3-5): 7-12 (4-6 + 3-6): 9-11 (4-6 + 4-6): 8-10 (5 + 3-5): 9-11 (4-6 + 4-6), 8-10 (4-5 + 3-5), sternite XI with 6-7 (3-4 + 3-4) and a pair of long tactile setae; sternites II with two lyrifissures.

Protonymphs (15 specimens analyzed) (Table 1). Chaetotaxy of carapace: 29-38 setae, 17-22 of them on anterior disk, 4-11 on medial disk, posterior margin with 6-8 setae. Chelicera: four setae on hand, none on movable finger; galea with 3-4 short terminal rami, serrula exterior with 11-14 blades. Palps (Fig. 5C): three trichobothria on fixed chelal finger and 1 trichobothrium on movable chelal finger; fixed chelal finger with 24-29 and movable chelal finger with 26-31 marginal teeth; both chelal finger without any accessory teeth. Chaetotaxy of tergites I–X: each with 6 setae (left tergite half 3 + right tergite half 3), tergite XI with 2 setae (1 + 1) and a pair of long tactile setae. Chaetotaxy of sternites II–X: 0-9 (left hemisternite 0-1 + right hemist ernite 0-9): 2 (1 + 1): 3-5 (1-3 + 1-3): 6-8 (3-5 + 3): 6-7 (3 + 3-4): 6-7 (3-4 + 3): 4-7 (3+ 1-4): 5-6 (2-3 + 3), 4-6 (2-3 + 2-3), sternite XI with 2 (1 + 1) and a pair of long tactile setae; sternites II with two lyrifissures.