Stenus atratulus Erichson, 1839

Serri, Sayeh & Frisch, Johannes, 2016, Species diversity, chorology, and biogeography of the Steninae MacLeay, 1825 of Iran, with comparative notes on Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae), Deutsche Entomologische Zeitschrift 63 (1), pp. 17-44 : 25

Stenus atratulus Erichson, 1839 Fig. 18 , Suppl. material 1

Chorology.

The West Palaearctic Stenus atratulus is widely distributed throughout Europe, the Mediterranean including the Maghreb countries Algeria and Tunesia, and Turkey eastwards to Iran, where it is most abundant in the northwest (Fig. 18 ), from where it dispersed across the South Caspian mountains as far east as the Koppeh Dag at about 58°E in North Khorasan, the easternmost confirmed locality of the species. Interestingly, Stenus atratulus was not found in the Zagros Mountains south of Hamedan Province, but it occurs in the dryer Qohrud Range, where we recorded it at Shir Kuh and in the mountains of Kerman (Fig. 18 ).

Biogeographical characterization.

Due to its Mediterranean distribution centre, we classify Stenus atratulus as a Holomediterranean species following Puthz (2012a: 78).

Figures

Figures 13 - 18. Distribution of Steninae in Iran. 13. Stenus guttula species group: ○ Stenus erythrocnemus, ● Stenus maculiger; 14. Stenus hopffgarteni species group: ○ Stenus fuscicornis; 15. Stenus humilis species group: ○ Stenus callidus; 16. Stenus impressus and Stenus incanus species groups: ○ Stenus heinzianus, △ Stenus taurus; 17. Stenus latifrons and Stenus melanarius species groups: ▲ Stenus latifrons, ○ Stenus incrassatus, □ Stenus morio, ● Stenus peripherus; 18. Stenus melanopus species group: ○ Stenus atratulus, ● Stenus piscator.