Alysson pertheesi Gorski 1852

Nemkov, Pavel G., 2014, The digger wasps of the genus Alysson Panzer (Hymenoptera: Crabronidae: Bembicinae) of Russia and adjacent territories, with a key to species and new synonymies, Zootaxa 3838 (3), pp. 276-286 : 280-281

Alysson pertheesi Gorski 1852

Alysson pertheesi Gorski 1852: 178 , ♀. Syntypes: ♀♀, Lithuania, Vilnius (ZMHU).

Alyson festivus Mocsáry, 1879: 129 , ♀ (as festivum, incorrect original termination). Holotype or syntypes: ♀, Hungary, Budapest (TMB). Synonymized with Alysson pertheesi by Handlirsch, 1888: 250.

Alysson pertheesi: Handlirsch 1888: 251 ; de Dalla Torre 1897: 564; Kokujev 1906: 216; Tsuneki 1965: 25, 1968: 5, 1974: 361; Bohart & Menke 1976: 458; Kolesnikov 1977: 318; Pulawski 1978: 212; Kazenas 1980: 85; Mityaev et al. 1980: 119; Tsuneki 1982: 18; Nemkov 1986:106; Wu & Zhou 1987: 214; Tsuneki 1991: 199; Nemkov Kazenas, Budrys & Antropov 1995: 455; Shlyakhtenok & Skibinska 2002: 34; Nemkov 2005: 158; Hua 2006: 277; Terayama 2006: 2, 4; Nemkov 2007a: 74, 2007b: 997, 2008: 24, 2009: 127; Baghirov 2011: 141; Nemkov 2012b: 445.

Alysson fuscatus var. jaroslavensis Kokujev 1906: 216 , ♀. Holotype: ♀, Russia, Yaroslavskaya Oblast, Yaroslavl (ZIN), examined. New synonymy.

Alysson harbinensis Tsuneki 1967: 7 , ♀. Holotype: ♀, China, Heilongjiang, Harbin (NNML). New synonymy.

Alysson harbinensis: Tsuneki 1968: 5 ; Bohart & Menke 1976: 458; Wu & Zhou 1987: 214; 1996: 136; Hua 2006: 277.

Alysson pertheesi forma typica Tsuneki and Itami, 1973: 13. Not available (Article 16). Alysson pertheesi forma nigra Tsuneki and Itami, 1973: 13, ♀. Syntypes: ♀ ♀, Japan, Niigata Prefecture, Shidaihama (Itami coll., K. Tsuneki coll.). Not available (Article 16).

Alysson spinosus jaroslavensis: Bohart & Menke 1976: 458 ; Shlyakhtenok & Skibinska 2002: 34. Alysson jaroslavensis: Pulawski 1978: 212 , 213; Pankevich et al. 1989: 95; Blagoveshchenskaya 1994: 91; Voblenko et al. 1996: 15.

Material examined. Holotype of Alysson fuscatus var. jaroslavensis Kokujev—♀, Russia: Yaroslavskaya Oblast, Yaroslavl, 14.IX 1896 (Kokujev) ( ZIN). Russia: 1♂, Leningradskaya Oblast, Tolmachevo, 13.VII 1936 (Shtakelberg) ( ZIN); 2♀, 1♂, Kostromskaya Oblast, Kostroma, 24.VII 1924, 22.VII 1926, 10.VIII 1933 (Gussakovskij) ( ZIN); 1♂, Kostromskaya Oblast, Troitsk, 25.VII 1924, 22.VII 1929 (Gussakovskij) ( ZIN); 1♀, Kostromskaya Oblast, Vasilevskoe, 21.VII 1933 (Gussakovskij) ( ZIN); 1♀, 3♂, Kostromskaya Oblast, Baidarki, 19.VII, 1.VIII 1933 (Gussakovskij) ( ZIN); 1♀, Volgogradskaya Oblast, Volgograd (coll. F. Morawitz) ( ZIN); 2♀, Krasnoyarskii Krai, Malaya Minusa, 4, 7.VII 2012 (Proshchalykin, Loktionov) ( IBSS); 2♀, Krasnoyarskii Krai, 40 km NE Minusinsk, Tes, Tuba River, 7.VII 2012 (Proshchalykin, Loktionov) ( IBSS); 1♀, Krasnoyarskii Krai, Minusinsk, 9.VII 2012 (Proshchalykin, Loktionov) ( IBSS); 4♀, 2♂, Irkutskaya Oblast, Angarsk, 11.VII–9.IX 1983 (Nemkov) ( IBSS); 4♀, Irkutskaya Oblast, 15 km E Ust-Orda, Ordinsk, 2–6.VIII 1994 (Nemkov) ( IBSS); 1♀, Khabarovskii Krai, Khabarovsk, 22.VIII 1979 (Ermolenko) ( ZIN); 1♂, Khabarovskii Krai, Amurzet, 17.VI 1985 (Belokobylskij) ( ZIN); 1♀, Primorskii Krai, Yakovlevka, 23.VIII 1926 (Dyakonov, Filipev) ( ZIN); 1♀, Primorskii Krai, Olga, 27.VII 1979 (Belokobylskij) ( ZIN); 1♀, Primorskii Krai, Ussuriiskii Reserve, Kamenushka, 12.VIII 1982 (Nemkov) ( IBSS); 1♀, Primorskii Krai, Ussuriiskii Reserve, Staraya Baza, 29.VI 1991 (Lelej) ( IBSS); 1♀, Primorskii Krai, Ryazanovka, 15.VII 1992 (Lelej) ( IBSS); 2♂, Primorskii Krai, Khasan, 12.VIII 1998 (Belokobylskij) ( ZIN); 3♀, Sakhalinskaya Oblast, Sakhalin Island, 10 km E Boshnyakovo, 24.VII 2003 (Lelej, Storozhenko) ( IBSS). Mongolia: 1♂, Khövsgöl aimag, Delger-Muren River near Buren-Khan, 28–29.VI 1968 (Kozlov) ( ZIN).

Distribution. Netherlands, France, Denmark, Austria, Poland, Slovakia, Hungary, Lithuania, Belarus, Ukraine, Russia (Leningradskaya Oblast, Yaroslavskaya Oblast, Kostromskaya Oblast, Moskovskaya Oblast, Bryanskaya Oblast, Volgogradskaya Oblast, Ulyanovskaya Oblast, Altai, Krasnoyarskii Krai, Irkutskaya Oblast, Buryatiya, Amurskaya Oblast, Khabarovskii Krai, Primorskii Krai, Sakhalin, Kuril Islands: Kunashir), Mongolia, China (Heilongjiang, Taiwan), Republic of Korea, Japan (Honshu).

Justification of New Synonymy. Kokujev (1906) described var. jaroslavensis as a form of A. fuscatus (correctly A. spinosus ). Bohart and Menke (1976) treated A. fuscatus jaroslavensis as a subspecies of A. spinosus . However, it differs from the latter in having a red propodeum (black in A. spinosus ) and is similar in this and other characters to A. pertheesi . Pulawski (1978) regarded A. jaroslavensis as a valid species. The examination of the holotype of Alysson fuscatus var. jaroslavensis demonstrated that it is a junior subjective synonym of A. pertheesi .

In the original description of A. harbinensis Tsuneki (1967) indicated that the female (the male was still unknown) differs from A. pertheesi mainly in having a red metapleuron and the propodeal enclosure (enclosure all black in pertheesi ). The considerable color variation in the Eastern Asian populations of A. pertheesi was noted later by Tsuneki and Itami (1973). I was not able to examine the holotype of A. harbinensis because of the temporary closure of the collection at the Leiden Museum, but the diagnostic characters of this species are clearly within the limits of the individual variation of A. pertheesi . Specimens with a red propodeal enclosure (as in harbinensis ) occur in all parts of Russia along with the typically colored specimens (with a black enclosure), although in the western Russia the typical form prevails over the harbinensis form, while in the eastern Russia the specimens with black enclosure are found very rarely. Therefore I regard A. harbinensis as a junior subjective synonym of A. pertheesi Gorski.