Cyclorhipidion achlys Smith, Beaver, Pham & Cognato, 2022

Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong & Cognato, Anthony I., 2022, New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae), Zootaxa 5209 (1), pp. 1-33 : 8-9

Cyclorhipidion achlys Smith, Beaver, Pham & Cognato sp. nov.

( Figs 21–24 )

http://zoobank.org/ urn:lsid:zoobank.org:act:17DCDCE9-DF6C-4B6F-8B94-F57908A9DE74

Type material. Holotype, female: VIETNAM, Lao Cai Prov., Hoang Lien N.P., 22.35N, 103.77E, 1500m, 17.v.2019, VN159, SM Smith, AI Cognato, ex 3–5 cm branch, SAX 397 [DNA voucher], MSUC_ARC_314124 ( MSUC).

GenBank. CAD: OP607183 and COI: OP617754 .

Diagnosis. This species is distinguished by the short, steep declivity that is ~1/5 of total elytral length, armed with large distinct tubercles on interstriae 1 and 3, interstriae 2 minutely granulate on basal third; moderate body length; posterolateral margins rounded; declivital interstriae 1 setae in three confused rows, interstriae 2 setae uniseriate; declivity strongly shagreened; strial and interstrial punctures small, shallow, strongly confused and indistinguishable.

Cyclorhipidion achlys shares a similar declivital setal pattern and size as C. pelliculosum ( Eichhoff, 1878) , but can be distinguished by the strial punctures which are large, distinct and seriate.

Similar species. Cyclorhipidion pelliculosum

Description (female). 3.4 mm long (n = 1); 3.09 × as long as wide. Body dark reddish brown, antennae and legs brown. Head: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface subshining, impunctate, alutaceous, finely rugose. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrow, triangular, deeply impressed. Antennal scape short and thick, as long as club. Pedicel narrower than scape, as long as funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club approximately circular and flat, type 3; segment 1 corneous, transverse on anterior face, occupying about 2/5 of club; segment 2 narrow, corneous; segments 1‒2 present on posterior face. Pronotum 1.09 × as long as wide. In dorsal view subquadrate, sides convex, type 3, narrowly rounded anteriorly; anterior margin without serrations. In lateral view elongate with disc much longer than anterior slope, type 7, disc flat, summit at apical 2/5. Anterior slope shagreened, with densely spaced, fine, narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc dull, alutaceous, densely, finely punctate, finely setose, setae recumbent, hair-like. Lateral margins obliquely costate. Base transverse, posterior angles broadly rounded. Scutellum moderate, broad, linguiform, flush with elytra, flat, shining. Elytra: 2.0 × as long as wide, 1.83 × as long as pronotum. Base transverse, edge oblique, humeral angles rounded, parallel-sided in basal 9/10, then strongly rounded to apex. Disc flat, shining, striae and interstriae moderately setose, setae long, erect, hair-like, striae not impressed, punctures minute, confused, spaced by 4‒6 diameters of a puncture; interstriae not impressed, minutely punctate, punctures equal to strial punctures, strongly confused, separated by 3‒6 diameters of a puncture. Declivity occupying 1/5 of elytra, very steeply rounded, declivital face convex, strongly shagreened, distinctly separated from smooth, shining disc; striae 1‒3 equidistant, strial and interstrial punctures small, shallow, strongly confused, indistinguishable, strial punctures as large as those of disc; interstriae 1 and 2 flat, interstriae 3 convex, interstriae 1 bearing three rows of confused setae on declivital face, interstriae 2 bearing one row of uniseriate setae, setae long, semi-erect hair-like; interstriae 1 and 3 armed with a row of moderate to large distinct tubercles, interstriae 2 minutely granulate on basal third. Posterolateral margin rounded, granulate, extending to interstriae 7. Legs: procoxae contiguous. Protibiae semi-circular with evenly rounded outer edge, broadest at apical 1/3; posterior face smooth; apical 1/3 of outer margin with eight moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae broad, flattened; outer margin evenly rounded each with 14, and 12, small to moderate socketed denticles, respectively.

Distribution. Vietnam.

Host plants. Unknown.

Etymology. G. achlys = mist in eyes before death. Achlys is the Greek Goddess of death-mist, and the personification of misery and suffering. This species represents the numerous nearly indistinguishable pseudocryptic Cyclorhipidion species that dwell in the misty mountain tops of Southeast Asia and perpetuate misery among taxonomists. A noun in apposition.

Figures

FIGURES 21‒36. Lateral, dorsal, posterior oblique and anterior view of Cyclorhipidion achlys holotype, 3.4 mm (21–24), Cyclorhipidion conidentatus holotype, 3.1 mm (25–28), Cyclorhipidion gladigerum holotype, 2.6 mm (29–32), Cyclorhipidion lapilliferum holotype, 3.4 mm (33–36).