Iodocephalopsis eberhardtii (Gagnep.) Bunwong & H.Rob., Proc. Biol. Soc. Washington 122(3): 358. 2009.

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 74

Iodocephalopsis eberhardtii (Gagnep.) Bunwong & H.Rob., Proc. Biol. Soc. Washington 122(3): 358. 2009.

Iodocephalus eberhardtii Gagnep., Notul. Syst. (Paris) 4: 18. 1920.

Camchaya eberhardtii (Gagnep.) Kitam., Acta Phytotax. Geobot. 23: 71. 1968.

Iodocephalopsis eberhardtii Type. Vietnam, Annam, Lang-bian, Eberhardt 1711 (holotype: P!). Fig. 9C .

Iodocephalus glandulosus Kerr, Kew Bull.: 326. 1935.

Iodocephalopsis eberhardtii Type: Thailand, Chiang Mai, Doi Sutep, A.F.G. Kerr 789 (holotype: BM!).

Description.

Perennial herbs, 0.5-1 m tall. Stems erect, terete, inconspicuously ribbed, puberulous with T-shaped hairs and glands. Leaves alternate, obovate-lanceolate, 4-10 by 1-4 cm, margin serrate or entire, apex acute, base attenuate, chartaceous; both surfaces puberulous with whip-shaped hairs, cylindrical hairs and capitate glands; lateral veins 5-8-paired; petioles up to 3 mm long. Capitulescences terminal or axillary, corymbose or solitary. Capitula campanulate, 8-10 mm long, pedunculate. Receptacle convex, 2.5-4 mm in diam., glabrous. Involucre campanulate, in 3-4 series, 6-7 mm long. Phyllaries imbricate, dull green, margin piliferous, outer surface arachnoid, glandular; the outer and the middle ones ovate, apex acute to acuminate; the inner ones ovate-lanceolate, apex acute to acuminate. Florets 15-25; corollas funnelform, purple or white, puberulous, glands capitate; corolla tubes 2.5-4 mm long; corolla lobes 2-2.5 mm long. Anthers ca. 2 mm long, apical appendage acute, base rounded. Styles purple, 3-5 mm long, branches 1-2 mm long. Achenes turbinate, 3-5 mm long, glandular, 7-10-ribbed. Pappus absent.

Distribution.

Thailand: Chiang Mai, Chiang Rai, Nan, Phitsanulok, Loei, Khon Kaen, Nakhon Ratchasima, Phetchaburi, Nakhon Nayok. Laos, Myanmar.

Specimens examined.

Thailand, Chiang Mai, Doi Sutep Pui national park, 18°48.39'N, 98°54.90'E, 8 Oct 2007, S. Bunwong 335 (KKU, US); Doi Sutep Pui national park, 4 Oct 1986, Y. Paisooksantivatana 1857-86 (BK); Doi Sutep Pui national park, 14 Oct 1979, T. Shimizu, H. Toyokuni, H. Koyama, T. Yahama & T. Santisuk T-18670 (L); Loei, Phu Reau, 23 Dec 1982, H. Koyama, H. Terao, Th. Wongprasert T-31570 (BKF, L); Phu Luang, 24 Dec 1982, H. Koyama, H. Terao, Th. Wongprasert T-30837 (BKF); Phu Kra Dung, 18 Jan 1991, Din 6 (P); Phu Kra Dung,1 Nov 1984, G. Murata, C. Phengklai, S. Mitsuta, T. Yahara, H. Nagamasu & N. Nantasan T-42599 (L); Phu Kra Dung, 31 Oct 1984, S. Mitsuta, H. Nagamasu, T. Yahara & N. Nantasan T-42260 (BKF); Khon Kaen, Phu Khiew, 7 Nov 1984, G. Murata, K. Iwatsuki, C. Phengklai & C. Charenphol T41669 (BKF); Nakhon Nayok, Khao Yai, 12 Aug 1974, J.F. Maxwell 74-808 (BK).

Diagnostic characters.

Iodocephalopsis eberhardtii is different from Camchaya spp. by having 3-colporate pollen, no pappus and phyllaries without marginal spines.

Ecology.

Edge of evergreen and pine-oak forests, alt. 700-1700 m; flowering August to December.

Vernacular name.

Muang Cha Rad (ม่วงจรัส).

Figures

Figure 9. Morphology of Vernonieae in Thailand 5. A-B Gymnanthemum extensum C Iodocephalopsis eberhardtii D Koyamasia calcarea E-F Koyamasia curtisii G Kurziella gymnoclada H Monosis parishii I Monosis volkameriifolia