Dichotomius (Dichotomius) satanas angustus (Luederwaldt, 1923)

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 122

Dichotomius (Dichotomius) satanas angustus (Luederwaldt, 1923) Plate 28A

Pinotus satanas var. angustus Luederwaldt, 1923: 10 (original description. Type locality: Ecuador Sarayacu; Ost-Kordillere, Sued-Ecuador [= eastern Cordillera, southern Ecuador], 300 m; Canolos [= Canelos] Ost-Ecuador [= eastern Cordillera, Ecuador], 700 m; Macas Ost-Ecuador [= eastern Cordillera, Ecuador]; Loja-Sabanilla Ost-Ecuador [= eastern Cordillera, Ecuador]; Bannos-Mirador [= Baños-Mirador] Ost-Ecuador [= eastern Cordillera, Ecuador], 14-1600 m).

Pinotus satanas var. angustus : Luederwaldt 1929: 36 (characters in key).

Pinotus angustus : Blackwelder 1944: 206 (list of species from Latin America); Carvajal et al. 2011: 320-321 (cited for Ecuador).

Dichotomius (Dichotomius) satanas angustus : Chamorro et al. 2018: 95 (cited for Ecuador).

Type specimens.

Pinotus satanas var. angustus Luederwaldt, 1923. Two syntypes examined deposited at the MZUSP. Lectotype to be designated in a future work on this species group.

Syntype (♂): "O.ECUADOR / Canelos / F. Ohs. 23. 12. 05 [p]", "COTIPO [p, black margin, red label]", "Pinotus ♂ / angustus Lüd. / Lüd. det. 22. [hw]", "17228 [p]".

Syntype (♂): "Sud-Ecuador, / Ostcordill 3000 m. / F. Ohaus 19-x- [hw]", "17226 [p]", "COTIPO [p, black margin, red label]".

Distribution.

Colombia and Ecuador.

Records examined.

LOJA: without specific locality (1 specimen MZUSP). NAPO: Archidona (1 specimen CEMT); Cabañas San Isidro, 2 km NW de Cosanga, 2150 m (2 specimens CEMT); Cosanga Yanayacu Biost, 2150 m (3 specimens MECN); Río Hollín, 1100 m (1 specimen CEMT). PASTAZA: Canelos (1 specimen MZUSP). TUNGURAHUA: Baños El Topo, 1590 m (24 specimens CEMT). SUCUMBÍOS: La Sofía, 1800 m (2 specimens MUTPL); Sebundoy, 2200 m (2 specimens MECN). ZAMORA CHINCHIPE: Bombuscaru, Parque Nacional Podocarpus, 1150 m (2 specimens MECN); Chito Río Sangolas, 1800 m (4 specimens MUTPL); San Andres, 1850 m (4 specimens CEMT). UNDETERMINED PROVINCE: Ostcordill [= without specific locality], 3000 m (1 specimen MZUSP).

Literature records.

LOJA: Sabanilla [= El Tambo, ZAMORA CHINCHIPE] ( Luederwaldt 1923: 11). MORONA SANTIAGO: Macas: ( Luederwaldt 1923: 11). PASTAZA: Canolos [= Canelos], 700 m ( Luederwaldt 1923: 11); Sarayacu ( Luederwaldt 1923: 11). TUNGURAHUA: Bannos-Mirador [= Baños Mirador], 1400-1600 m. ( Luederwaldt 1923: 11). UNDETERMINED PROVINCE: Ost-Kordillere, Sued-Ecuador, 3000 m ( Luederwaldt 1923: 11).

Temporal data.

Collected in January, February, May, July, October, September, November, and December.

Remarks.

Inhabits the evergreen foothill forests of the Amazon region from 700-1150 m a.s.l. In the Andean region, it was recorded in the evergreen montane forests and montane cloud forests from 1600-2200 m a.s.l. Collected with pitfall traps baited with human feces.