Ancylis goodelliana ( Fernald, 1882 )

Gilligan, Todd, Huemer, Peter & Wiesmair, Benjamin, 2016, Different continents, same species? Resolving the taxonomy of some Holarctic Ancylis Hübner (Lepidoptera: Tortricidae), Zootaxa 4178 (3), pp. 347-370 : 361-362

Ancylis goodelliana ( Fernald, 1882)

Figs. 17–22, 65, 79

Phoxopteris goodelliana Fernald 1882 , Trans. Am. ent. Soc. 10: 69.

Diagnosis. The light gray to white costal margin and black streaks along the radius and cubitus of the forewing separate A. goodelliana from other Ancylis treated here. Some individuals of A. uncella appear similar, but the brown to reddish-brown median fascia is always expressed on the costa in A. uncella . Lighter (or worn) individuals of A. diminuatana may also appear similar, although the costa is usually not as pale in the distal half, and A. diminuatana lacks the black streak along the radius.

Redescription. Forewing. FWL Ƌ 6–9.5 mm (n=16), ♀ 6.5–8.5 mm (n=4). Forewings are brown to reddish brown and gray with white costal strigulae near the apex. The costa is white to light gray from base to near apex, and the same color is present along the dorsum. No fasciae are defined, instead a brown to reddish-brown band runs from base to apex; the dorsal margin of this band is sinuate as a result of remnants of the median fascia extending towards the tornus. Two black streaks are present: one along the cubitus to ca. one-third the distance to the termen, and one along the distal two-thirds of the radius to the apex (most evident in Fig. 22). Male genitalia. Uncus bifid to more than half its length. Valva with basal excavation nearly absent; saccular angle weakly to moderately developed with variable triangular terminal projection; neck of uniform width or slightly narrowed from sacculus to cucullus; cucullus elongate, tapering towards apex, densely setose, outer margin convex with several rows of short setae near the anal angle, apex rounded. Caulis 0.5 as long as phallus; phallus 0.7 as long as valva, with small tooth at apex (difficult to see in some preparations); vesica with ca. 40–70 deciduous lanceolate cornuti. The length of the cucullus, the width of the valval neck, and the saccular terminal projections all vary slightly between individuals, although genital variation is not as pronounced as in other species in this group. Female genitalia. As in A. uncella .

Molecular data. BIN URI: BOLD:AAA7191, BIN-sharing with A. uncella , with a minimum distance of 1.13%.

Distribution. Most specimens of A. goodelliana are from eastern North America: Nova Scotia west to Ontario, and Wisconsin south to North Carolina and Florida. Heinrich (1923) reported this species from Manitoba and Colorado, and we examined one specimen from central Alberta in the CUIC, suggesting that it also present in the West. Historical records are difficult to confirm because of confusion with A. diminuatana in many collections.

Biology. Adults are present from the end of May through mid-August. The larval host is unknown.

Remarks. Worn specimens of A. goodelliana are easily confused with A. diminuatana , and the two are often mixed in collections. A genitalic dissection will easily separate A. goodelliana from all species in the A. geminana group. Gilligan et al. (2008) incorrectly illustrated the adult and male genitalia of A. goodelliana as “ A.

diminutana .” Although A. goodelliana and A. uncella appear very similar based on COI barcode data (with a minimum distance of only 1.13%), the two species can be reliably separated by forewing pattern.