Bembidium (Cillenus), Samouelle, 1819

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 364

Subgenus Cillenus Samouelle, 1819

Cillenus Samouelle, 1819: 148. Type species: Cillenus lateralis Samouelle, 1819 by monotypy. Etymology. Unknown [masculine].

Cillenum Curtis, 1828: plate 200. Unjustified emendation of Cillenus Samouelle, 1819.

Diversity.

Two species, one along the Pacific Coast of North America, the other ( Bembidion laterale Samouelle) along the coasts of Europe and northern Africa. The three other species listed in this subgenus by Lorenz (2005: 216) belong to the subgenus Desarmatocillenus Netolitzky (see Lindroth 1980).

Identification.

Kavanaugh and Erwin (1992: 317) emended Lindroth’s (1963b) key to Bembidion to accommodate the new species.

Taxonomic Note.

This taxon is ranked as a distinct genus by many authors, with Armatocillenus Dupuis (three species in Taiwan and New Guinea), Desarmatocillenus Netolitzky (14 species in southeastern Asia and the Australian Region), Chinocillenus Netolitzky (one Chinese species), Corallicillenus Uéno (two Japanese species), and Novicillenus Uéno and Habu (two Japanese species) listed as subgenera (see Lorenz 2005: 216). Recently Sasakawa (2007) synonymized Novicillenus and Corallicillenus with Desarmatocillenus . Recent molecular data analyses conducted by Maddison and Ober (2011: 251) and Maddison (2012: 568) strongly support Cillenus as a member of Bembidion .