Hipposideros Gray 1831

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Chiroptera - Family Hipposideridae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press, pp. 365-379 : 367

Hipposideros Gray 1831

Hipposideros Gray 1831 , Zool. Misc., Vol. 1: 37.

Type Species: Vespertilio speoris Schneider 1800

Synonyms: Chrysonycteris Gray 1866 ; Cyclorhina Peters 1871 ; Gloionycteris Gray 1866 ; Phyllorhina Bonaparte 1837 ; Ptychorhina Peters 1871 ; Speorifera Gray 1866 ; Syndesmotis Peters 1871 ; Syndesmotus Waterhouse 1902 ; Thyreorhina Peters 1871 .

Species and subspecies: 67 species with 76 subspecies:

Species Hipposideros abae J. A. Allen 1917

Species Hipposideros armiger Hodgson 1835

Subspecies Hipposideros armiger subsp. armiger Hodgson 1835

Subspecies Hipposideros armiger subsp. fujianensis Zhen 1987

Subspecies Hipposideros armiger subsp. terasensis Kishida 1924

Subspecies Hipposideros armiger subsp. tranninhensis Bourret 1942

Species Hipposideros ater Templeton 1848

Subspecies Hipposideros ater subsp. ater Templeton 1848

Subspecies Hipposideros ater subsp. amboinensis Peters 1871

Subspecies Hipposideros ater subsp. antricola Peters 1861

Subspecies Hipposideros ater subsp. aruensis Gray 1858

Subspecies Hipposideros ater subsp. gilberti Johnson 1959

Subspecies Hipposideros ater subsp. nicobarulae Miller 1902

Subspecies Hipposideros ater subsp. saevus K. Andersen 1918

Species Hipposideros beatus K. Andersen 1906

Subspecies Hipposideros beatus subsp. beatus K. Andersen 1906

Subspecies Hipposideros beatus subsp. maximus Verschuren 1957

Species Hipposideros bicolor Temminck 1834

Subspecies Hipposideros bicolor subsp. bicolor Temminck 1834

Subspecies Hipposideros bicolor subsp. atrox K. Andersen 1918

Subspecies Hipposideros bicolor subsp. erigens Lawrence 1939

Subspecies Hipposideros bicolor subsp. hilli Kitchener 1996

Subspecies Hipposideros bicolor subsp. major K. Andersen 1918

Subspecies Hipposideros bicolor subsp. selatan Kitchener 1996

Subspecies Hipposideros bicolor subsp. tanimbarensis Kitchener 1996

Species Hipposideros breviceps Tate 1941

Species Hipposideros caffer Sundevall 1846

Subspecies Hipposideros caffer subsp. caffer Sundevall 1846

Subspecies Hipposideros caffer subsp. angolensis Seabra 1898

Subspecies Hipposideros caffer subsp. nanus J. A. Allen 1917

Subspecies Hipposideros caffer subsp. tephrus Cabrera 1906

Species Hipposideros calcaratus Dobson 1877

Subspecies Hipposideros calcaratus subsp. calcaratus Dobson 1877

Subspecies Hipposideros calcaratus subsp. cupidus K. Andersen 1918

Species Hipposideros camerunensis Eisentraut 1956

Species Hipposideros cervinus Gould 1854

Subspecies Hipposideros cervinus subsp. cervinus Gould 1854

Subspecies Hipposideros cervinus subsp. batchianus Matschie 1901

Subspecies Hipposideros cervinus subsp. labuanensis Tomes 1859

Subspecies Hipposideros cervinus subsp. misoriensis Peters 1906

Species Hipposideros cineraceus Blyth 1853

Subspecies Hipposideros cineraceus subsp. cineraceus Blyth 1853

Subspecies Hipposideros cineraceus subsp. wrighti Taylor 1934

Species Hipposideros commersoni (E. Geoffroy 1813)

Species Hipposideros coronatus Peters 1871

Species Hipposideros corynophyllus Hill 1985

Species Hipposideros coxi Shelford 1901

Species Hipposideros crumeniferus (Lesueur and Petit 1807)

Species Hipposideros curtus G. M. Allen 1921

Species Hipposideros cyclops Temminck 1853

Species Hipposideros demissus K. Andersen 1909

Species Hipposideros diadema (E. Geoffroy 1813)

Subspecies Hipposideros diadema subsp. diadema E. Geoffroy 1813

Subspecies Hipposideros diadema subsp. ceramensis Laurie and Hill 1954

Subspecies Hipposideros diadema subsp. custos K. Andersen 1918

Subspecies Hipposideros diadema subsp. enganus K. Andersen 1907

Subspecies Hipposideros diadema subsp. euotis K. Andersen 1905

Subspecies Hipposideros diadema subsp. griseus Meyen 1883

Subspecies Hipposideros diadema subsp. masoni Dobson 1872

Subspecies Hipposideros diadema subsp. mirandus Thomas 1914

Subspecies Hipposideros diadema subsp. natunensis Chasen 1940

Subspecies Hipposideros diadema subsp. nicobarensis Dobson 1871

Subspecies Hipposideros diadema subsp. nobilis Horsfield 1823

Subspecies Hipposideros diadema subsp. oceanitis K. Andersen 1905

Subspecies Hipposideros diadema subsp. reginae Troughton 1937

Subspecies Hipposideros diadema subsp. speculator K. Andersen 1918

Subspecies Hipposideros diadema subsp. trobrius Troughton 1937

Species Hipposideros dinops K. Andersen 1905

Species Hipposideros doriae Peters 1871

Species Hipposideros durgadasi Khajuria 1970

Species Hipposideros dyacorum Thomas 1902

Species Hipposideros edwardshilli Flannery and Colgan 1993

Species Hipposideros fuliginosus Temminck 1853

Species Hipposideros fulvus Gray 1838

Subspecies Hipposideros fulvus subsp. fulvus Gray 1838

Subspecies Hipposideros fulvus subsp. pallidus K. Andersen 1918

Species Hipposideros galeritus Cantor 1846

Subspecies Hipposideros galeritus subsp. galeritus Cantor 1846

Subspecies Hipposideros galeritus subsp. brachyotis Dobson 1874

Subspecies Hipposideros galeritus subsp. insolens Lyon 1911

Subspecies Hipposideros galeritus subsp. longicauda Peters 1861

Species Hipposideros gigas Wagner 1845

Species Hipposideros grandis G. M. Allen 1936

Species Hipposideros halophyllus Hill and Yenbutra 1984

Species Hipposideros hypophyllus Kock and Bhat 1994

Species Hipposideros inexpectatus Laurie and Hill 1954

Species Hipposideros inornatus (McKean 1970)

Species Hipposideros jonesi Hayman 1947

Species Hipposideros lamottei Brosset 1985

Species Hipposideros lankadiva Kelaart 1850

Species Hipposideros larvatus (Horsfield 1823)

Subspecies Hipposideros larvatus subsp. larvatus Horsfield 1823

Subspecies Hipposideros larvatus subsp. barbensis Miller 1900

Subspecies Hipposideros larvatus subsp. leptophyllus Dobson 1874

Subspecies Hipposideros larvatus subsp. neglectus Sody 1936

Subspecies Hipposideros larvatus subsp. poutensis Allen 1906

Species Hipposideros lekaguli Thonglongya and Hill 1974

Species Hipposideros lylei Thomas 1913

Species Hipposideros macrobullatus Tate 1941

Species Hipposideros madurae Kitchener and Maryanto 1993

Subspecies Hipposideros madurae subsp. madurae Kitchener and Maryanto 1993

Subspecies Hipposideros madurae subsp. jenningsi Kitchener and Maryanto 1993

Species Hipposideros maggietaylorae Smith and Hill 1981

Subspecies Hipposideros maggietaylorae subsp. maggietaylorae Smith and Hill 1981

Subspecies Hipposideros maggietaylorae subsp. erroris Smith and Hill 1981

Species Hipposideros marisae Aellen 1954

Species Hipposideros megalotis Heuglin 1862

Species Hipposideros muscinus Thomas and Doria 1886

Species Hipposideros nequam K. Andersen 1918

Species Hipposideros obscurus Peters 1861

Species Hipposideros orbiculus Francis, Kock, and Habersetzer 1999

Species Hipposideros papua Thomas and Doria 1886

Species Hipposideros pelingensis Shamel 1940

Species Hipposideros pomona K. Andersen 1918

Subspecies Hipposideros pomona subsp. pomona K. Andersen 1918

Subspecies Hipposideros pomona subsp. gentiles K. Andersen 1918

Subspecies Hipposideros pomona subsp. sinensis K. Andersen 1918

Species Hipposideros pratti Thomas 1891

Species Hipposideros pygmaeus Waterhouse 1843

Species Hipposideros ridleyi Robinson and Kloss 1911

Species Hipposideros rotalis Francis, Kock, and Habersetzer 1999

Species Hipposideros ruber Noack 1893

Subspecies Hipposideros ruber subsp. ruber Noack 1893

Subspecies Hipposideros ruber subsp. guineensis K. Andersen 1906

Species Hipposideros scutinares Robinson, Jenkins, Francis, and Fulford 2003

Species Hipposideros semoni Matschie 1903

Species Hipposideros sorenseni Kitchener and Maryanto 1993

Species Hipposideros speoris (Schneider 1800)

Species Hipposideros stenotis Thomas 1913

Species Hipposideros sumbae Oei 1960

Subspecies Hipposideros sumbae subsp. sumbae Oei 1960

Subspecies Hipposideros sumbae subsp. rotiensis Kitchener and Maryanto 1993

Subspecies Hipposideros sumbae subsp. sumbawae Kitchener and Maryanto 1993

Species Hipposideros thomensis Bocage 1891

Species Hipposideros turpis Bangs 1901

Subspecies Hipposideros turpis subsp. turpis Bangs 1901

Subspecies Hipposideros turpis subsp. alongensis Bourret 1942

Subspecies Hipposideros turpis subsp. pendleburyi Chasen 1936

Species Hipposideros vittatus Peters 1852

Species Hipposideros wollastoni Thomas 1913

Subspecies Hipposideros wollastoni subsp. wollastoni Thomas 1913

Subspecies Hipposideros wollastoni subsp. fasensis Flannery and Colgan 1993

Subspecies Hipposideros wollastoni subsp. parnabyi Flannery and Colgan 1993

Discussion: Revised by Hill (1963 b). The genus is apparently paraphyletic, but alternative phylogenies (e.g., those of Bogdanowicz and Owen [1998] and Hand and Kirsch [1998]) disagree about genus and species relationships. Accordingly, I have retained the traditional contents of Hipposideros pending a thorough revision. Species groups follow Koopman (1994) with some modifications.