Longitarsus tmetopterus Jacobson

Konstantinov, Alexander S., 2005, New species of Middle Asian Longitarsus Latreille with discussion of their subgeneric placement (Coleoptera: Chrysomelidae), Zootaxa 1056, pp. 19-42 : 30-31

Longitarsus tmetopterus Jacobson

( Figs. 14 , 54–66)

Longitarsus tmetopterus Jacobson, 1893: 246 (Type locality: Kazakhstan, Jany­Kurgan: Fluss San­

sar. Lectotype Ψ ZMAS, designated here).

L. zeravshanicus Palij, 1970:13 (Type locality: Tadzhikistan, Leninabad (1.IV.1955, ♂). Type mate­

rial lost). Lopatin, 1977: 220 (synonymy).

L. nurataicus Palij, 1970:13 (Type locality: Kzakhstan, Nuratau. Lectotype ♂ USNM, designated

here). Lopatin, 1977: 220 (synonymy).

Comments: Longitarsus tmetopterus was described from male and female collected near the town of Janykurgan, west of Karatau mountain ridge on 29.III.1892 by D. Glasunov. A single female with the following labels was found in the collection of Zoological Institute (ZMAS): 1) Turkestan, Sansar, Glasunov, 1892; 2) L. tmetopterus typ. Jac. G. Jacobson det.; 3) k. G. Jacobsona. This specimen is designated here as a lectotype in order to have a unique bearer of this name and the standard for its application. A lectotype label is added: Lectotype Longitarsus tmetopterus Jacobson , des. A. Konstantinov 2005. The second specimen mentioned in the original description was not found. In addition, 5 females collected by K. Anger, on 21.III.1900 at Kalanmor, Zakaspiiskoi oblasti, and identified as L. tmetopterus by G. Jacobson are present in Lopatin collection and ZMAS.

Proposing L. nurataicus, Palij (1970) did not designate the holotype and depository of his collection is unknown. It is highly possible that it was lost. For L. zeravshanicus , the holotype was indicated. In addition to the locality of the holotype, Palij (1970) sites the following locations: 1) env. of Samarkand (Bulungur), 31.I.1963 (Ψɗ), 6.III.1963 (2 Ψ), 27.III. 1963 (Ψɗ); 2) env. of Nura­Tau, Karakum, 11.V.1945 (3ɗ). Palij (1970) indicated the following locations for L. nurataicus : river valleys of Chu, Syr­Dar’ya, and Zeravshan, piedmont of Turkestanskii, Zeravshanskii, Kirgizskii, Nuratau mountain ridges. He noted that L. nurataicus was collected in hundreds at the last location in April and May of 1965 and 1966. The specimens collected by Palij in 1965 and 1966 near Nuratau, obviously known to him at the time when he described the species, are available in the USNM collection. A male with the following label is designated as the lectotype: khr. Nuratau, 45 km NW. Dzhizaka, 8.IV.1966 (USNM). Five specimens with the same labels are designated as paralectotypes (3 USNM, 2 ZMAS). Four more specimens with the same labels except for 7.IV.1966 are also designated as paralectotypes (3 USNM, 1 ZMAS). Lectotype designation is done in order to have a unique bearer of this name and the standard for its application.

Lopatin (1977) synonymized both L. nurataicus and L. zeravshanicus with L. tmetopterus . The female genitalia of the lectotypes of L. tmetopterus ( Figs. 64–66 ) and L. nurataicus ( Figs. 60–63 ) are nearly identical which leaves no doubt in their synonymy. Also type localities of both species are part of the vast territory of Karakum ­ Kyzylkum deserts. As for L. zeravshanicus , there are no specimens that can be unambiguously attributed to this name. Specimens identified as topotypes of L. zeravshanicus in the Lopatin collection originate from Nuratau, and most likely are L. nurataicus (sensu Palij). The holotype of L. zeravshanicus was collected near Leninabad (northern Tadzhikistan) along the valley of Syr­Dar’ya river, not isolated from the main range of L. tmetopterus .

The female genitalia of the lectotype of L. tmetopterus and specimens determined by Jacobson as L. tmetopterus (Anger collecting) ( Figs. 54–59) exhibit some relatively minor differences in the shape of the vaginal palpi and spermathecal pump, but they are not significant enough to warrant recognition of separate species.

Figures

FIGURES 12 – 16. Median lobe of aedeagus (ventral and lateral views). 12 ­ L. sengloki. 13 ­ L. sogdianus. 14 ­ L. tmetopterus. 15 ­ L. truncatellus. 16 ­ L. turkomanorum.

FIGURES 64 – 66. Female abdomen and genitalia of L. tmetopterus (Lectotype). 64 ­ spermatheca. 65 ­ tignum and internal abdominal sclerites. 66 ­ vaginal palpi.

FIGURES 60 – 63. Female abdomen and genitalia of L. tmetopterus (L. nurataicus sensu Palij). 60 ­ spermatheca. 61 ­ vaginal palpi. 62 ­ abdominal tergites. 63 ­ tignum and internal abdominal sclerites.