Prionocerus coeruleipennis Perty, 1831

Geiser, Michael, 2010, Studies on Prionoceridae (Coleoptera: Cleroidea). II. A revision of the genus Prionocerus Perty, 1831, Zootaxa 2328, pp. 1-48 : 6-22

publication ID

https://doi.org/ 10.5281/zenodo.193097

DOI

https://doi.org/10.5281/zenodo.6210927

persistent identifier

https://treatment.plazi.org/id/724F87F2-9058-5028-BB8D-F950E87D87EB

treatment provided by

Plazi

scientific name

Prionocerus coeruleipennis Perty, 1831
status

 

Prionocerus coeruleipennis Perty, 1831

Prionocerus coeruleipennis Perty, 1831: 33 .

Prionocerus fuscipennis Lewis, 1879: 464 . Synonymised by Champion, 1919: 328. Prionocerus forticornis Schaufuss, 1887: 126 . Synonymised by Bourgeois, 1890: 175. Prionocerus brevicornis Schaufuss, 1887: 126 . Synonymised by Bourgeois, 1890: 175. Prionocerus coeruleipennis var. diversicollis: Pic, 1920b: 12 . n. syn.

Type material examined: P. coeruleipennis : Holotype Ƥ "I., Java., Epiphyta collaris Haan, caeruleipennis Perty"; ZSMU, in good condition.

P. f u s c i p e n n i s: Holotype Ƥ "Yokohama, Pryer, Japan, G. Lewis, 1910–320"; " Prionocerus fuscipennis Lewis "; " Type H. T."; " Prionocerus coeruleipennis Perty, Det. G. C. C. "; NHML, in good condition.

P. f o r t i c o r n i s: Syntype 3 " Prionocerus forticornis m. Philippin[es]", the back side of the label says " Prionocerus coeruleipennis Philipp Ƥ "; MNHB, in good condition.

P. brevicornis : Syntype Ƥ " Prionocerus brevicornis m. S.Celebes"; MNHB, in good condition.

P. coeruleipennis var. diversicollis : Holotype Ƥ "Manau Riang, Palembang" [ Indonesia: Sumatra]; MNHP. The holotype is in bad condition, having been damaged by dermestid larvae, lacking its entire abdomen and parts of the legs.

Other material examined: PAKISTAN (?): "Murree, coll. Staudinger" 1 3 ( SMTD).

INDIA: " India, 58 60" 2 Ƥ ( NHML); „Indes, Boys“ 1 Ƥ ( MHNG). SIKKIM: "Sikkim, Harmand 1886" 1 Ƥ ( MNHP).

ANDAMAN ISLANDS: "Andaman Islands, Roepstorff, Fry Coll., 1905.100" 2 Ƥ (NHML).

JAPAN (?): " Nippon Moyen, Env. de Tokio et Alpes de Nikko, J. Harmand 1901" 1 Ƥ (MNHP); " Japonia, Shikoku" 1 Ƥ (MTMB).

CHINA: " China " 1 Ƥ (NHMB), 1 3 (NHML). HAINAN: Hong Mo Leng, Fan Ta Chuen, 1. V. 1929, Lingnan University 5th. Hainan Is. Expedition 1929 1 Ƥ (NHML); Hung Mo Tung valley, on trail Fan Ta Chuen to Poh Shaang, 31. VII.–1. VIII 1929, Lingnan University 5th. Hainan Is. Expedition 1929 1 Ƥ (NHML); Hung Mo Tung valley, Fan Ta Chuen, 23. VII., Lingnan University 5th. Hainan Is. Expedition 1929 1 Ƥ (NHML); 15 km Dongxing, 1,5 km W Jianfeng, 70 m, 25. I. 1996, Jäch leg. 1 Ƥ (NHMW); "Hainan Is., S. China, Apr. 28 & 29, 1932, William K. Hoffmann" 1 3 (BIMH); Ch'ung mei, 15 mi. SE. of Naam-fung, Lin-kao District, 24.–26. VIII. 1932, F. K. To leg. 1 Ƥ (BIMH); Nai-suen, 21 mi. S.E. of Naam-fung, Lin-kao Dist., 31. VIII. 1932, F. K. To leg. 2 3 2 Ƥ (BIMH). YUNNAN: "Yunnan mérid., Vallée du Nam-Ti ou Pei-Ki- Ho entre Lao-Kay et Mong-Tsé, Dr. Gervais 1904" 1 Ƥ (MNHP); Xishuangbanna, 37 km NW Jinghong, Guo Men Shan (NNNR), 22°17.913' N / 100°38.850' E, 1080 m, KF blüh. Castanopsis, 26. V. 2008, A. Weigel leg. 1 3 (NKME); Xishuangbanna, 26 km W Jinghong, vi. Meng Song (NNNR), 22°04.65' N / 100°33.98' E, 1400–1800 m, at flowers of Castanopsis trees, 30. V. 2008, A. Weigel leg. 1 Ƥ (NKME). FUJIAN: Ziyungdonshan, NW slopes, 25°46' N / 117°20' E, 900–1100 m, 21.–22. VI. 2007, Jaroslav Turna leg. 1 Ƥ (CMG).

MYANMAR: " Burma " 1 3 (MNHP); " Burma, coll. Staudinger" 1 Ƥ (SMTD); "Burmah, Bowring, 63 47*" 3 Ƥ (NHML); "Burham, H. E. Andrewes Bequest, B. M. 1922–221" 1 Ƥ (NHML); "Birmah, Fry Coll., 1905.100" 1 Ƥ (NHML); " Burma (Süd-Ost), 5. XI. 1989, leg. Sab. Steinke" 1 Ƥ (SMNS); "Shwegoo-Myo, Fea, Ottobre 1885" 1 3 (MNHB); " Birmania, Shweego-Myo, Fea, Ottobre 1885" 1 3 (MCGD); "Rangoon, Birmania, Fea, 1887" 1 Ƥ (MCGD); "Rangoon, 96–198" 1 Ƥ (NHML); "Rangoon, Archbald Coll., B. M. 1926–391." 1 Ƥ (NHML); "Palon (Pegu), L. Fea, VIII. IX, 87" 1 Ƥ (MCGD); "Maulmein, H. E. Andrewes Bequest, B. M. 1922–221" 1 3 (NHML); "Maymyo, V. 10, H. L. Andrewes" 1 3 2 Ƥ (NHML); "Maymyo, VI. 10, H. L. Andrewes" 2 3 2 Ƥ (NHML); "Maymyo, VI. 14, H. L. Andrewes" 2 3 (NHML); Mon State, Kyaikto, 1.–5. VI. 2003, M. Klícha leg. 1 Ƥ (CMG); Mon State, Kyaikto-Pauktaw, 3.–6. VI. 2003, M. Klícha leg. 1 3 (CMG); "Shan States, Manders, Sharp Coll., 1905–313" 2 3 1 Ƥ (NHML); E Shan State, Kengtung (Kyaingtong), 14.–15. VI. 1997, J. Rejsek leg. 1 Ƥ (SMNS); S. Shan States, Pekkong, 900 m, 11. X. 1934, Malaise leg. 1 Ƥ (NMRS); S. Shan States, Road 40 km E of Taunggyi, 25. IX. – 13. X. 1934, Malaise leg. 1 Ƥ (NRMS); "Tenasserim, 44 24" 2 Ƥ (NHML); "Tenasserim, coll. Helfer" 14 3 10 Ƥ (MNPR); Tenasserim, Mekane, 90 km E of Moulmein, 200 m, 2.–8. XI. 1934, Malaise leg. 1 Ƥ (NRMS).

LAOS: "Annam-Laos, India, coll. W. Wittmer" 1 3 4 Ƥ (NHMB); "Annam-Laos, India " 1 3 3 Ƥ (NHMB); " Annam Laos " 16 3 17 Ƥ (MTMB); " Laos, Tonkin, coll. Staudinger" 2 Ƥ (SMTD); "Giranville, 3.– 24. XI. 1919, R. V. de Salvaza" 1 Ƥ (MNHP); "Keng Chap, près de Buenoco? [illegible], Laos " 1 Ƥ (MNHP); "Song Van, Ht. Mekong" 1 Ƥ (MNHP); Phongsaly prov., Phongsaly env., 21°41–2' N, 102°06–8' E, ~ 1500 m, 28. V.–20. VI. 2003, M. Brancucci leg. 1 3 3 Ƥ (NHMB); Phongsaly prov., Phongsaly env., 21°41–2' N, 102°06–8' E, ~ 1500 m, 28. V.–20. VI. 2003, V. Kubán leg. 1 3 (NHMB); Phongsaly prov., Phongsaly env., 21°41–2' N, 102°06–8' E, ~ 1500 m, 28. V. – 30. VI. 2003, P. Pacholátko leg. 1 3 3 Ƥ (NHMB); Phongsaly prov., Phongsaly env., 21°41' N, 102°06' E, ~ 1500 m, 6.–17. V. 2004, M. Brancucci leg. 2 Ƥ (NHMB); Phongsaly prov., Phongsaly env., 21°41' N, 102°06' E, ~ 1500 m, 6.–17. V. 2004, P. Pacholátko leg. 1 Ƥ (NHMB); Phongsaly prov., Phongsaly env., 21°41' N, 102°06–8' E, ~ 1500 m, 6.–17. V. 2004, V. Kubán leg. 2 3 (NHMB); Phongsaly prov., Ban Sano Mai, 21°21' N, 102°03' E, ~ 1150 m, 19.–26. V. 2004, P. Pacholátko leg. 2 Ƥ (NHMB); Luang Namtha Prov., Luang Namtha env., 21°02–04' N / 101°22–24' E, an Sträuchern und niedriger Vegetation, 31. VII. 2006, M. Geiser leg. 1 Ƥ (CMG); Luang Namtha Distr., Luang Namtha env., 2.– 9. V. 2007, M. Pejcha leg. 2 Ƥ (CMG); Louang Namtha env., 21°00.3' N 101°24.6' E, 31. V. 1997, E. Jendek & O. Šauša leg. 1 3 (NHMB); 20 km NW Louang Namtha, 21°09.2' N / 101° 18.7' E, alt. 900 +/– 100 m, 24.– 30. V. 1997, E. Jendek & O. Šauša leg. 2 Ƥ (MTMB); Louang Namtha prov., Namtha ⊣ Muang Sing, 21°09' N 101°19' E, 900–1200 m, 5.–31. V. 1997, Vit Kubán leg. 3 3 (NHMB); Oudomxay Prov., Natheng Village, Nam Veun stream, 29. IV. 2005, N. Jönsson, T. Malm & B. Viklund leg. 1 Ƥ (NRMS); S-Udomxai prov., Pak Beng, 450 m, 19°53'37'' N, 101°07'51'' E, 18.–27. V. 2001, Jiři Kolibač leg. 1 Ƥ (NHMB); Louang Prabang prov., Ban Song Cha, 20°33' N 102°14' E, 1200 m, 24. IV.–16. V. 1999, Vit Kubán leg. 1 Ƥ (NHMB); Luangphrabang, Thong Khan, 19°35' N 101°58' E, ~ 750 m, 11.–21. V. 2002, Vit Kubán leg. 2 Ƥ (NHMB); "Moung Sing, NW of Luang Prabang, 650 m, 6.–10. VI. 1960, Bamboo, S. Quate & L. Quate Collectors" 1 Ƥ (BIMH); Luang Prabang prov., Muang Phou Khoune, 19°26.38' N / 102°25.6' E, 1370 m, flying around in town (daytime), 20. VI. 2009, M. Geiser & D. Hauck leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 4 3 1 Ƥ (NHMB), 1 3 1 Ƥ (CMG); Louang Prabang prov., Ban Kiukacham, 19°33.56' N / 102°13,87' E, 1400 m, flying in village, 20. VI. 2009, M. Geiser & D. Hauck leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 3 (NHMB); Sayaboury Prov., Sayaboury, 17. V. 1965, Native Collector leg. 1 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 30. V. 1965, Native Collector leg. 4 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 14. VII. 1965, Native Collector leg. 7 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 9. VIII. 1965, Native Collector leg. 2 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 6. XII. 1965, Native Collector leg. 2 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 14. XII. 1965, Native Collector leg. 1 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 25. III. 1966, Native Collector leg. 3 3 5 Ƥ (BIMH), 1 3 1 Ƥ (CMG); Sayaboury Prov., Sayaboury, 13. IV. 1966, Native Collector leg. 5 3 9 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 21. IV. 1966, Native Collector leg. 2 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 13. IV. 1966, Native Collector leg. 1 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 20. V. 1966, Native Collector leg. 2 Ƥ (BIMH); Vientiane prov., Phou Khao Khouay N. P., 18°27' N / 102°49' E, 2005, P. Kresl leg. 3 3 3 Ƥ (NMPC); Vientiane Municipal., Phou Khao Khouay NBCA, Ban Van Hua env., 800–1000 m, 5.–20. V. 2005, M. Pejcha leg. 7 3 2 Ƥ (CMG); "Ban Van Heue, 20 km E of Phou-kow-kuei, 15.–31. V. 1965, Native Collector" 1 3 14 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 15.–31. V. 1965, Native Collector leg. 3 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 15. VII. 1965, Native Collector leg. 2 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 30. IX. 1965, Native Collector leg. 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 31. VII. 1965, Native Collector leg. 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 16. III. 1966, Native Collector leg. 2 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 29. III. 1966, Native Collector leg. 1 3 (BIMH); Vientiane Prov., Ban Van Eue, 15. IV. 1966, Native Collector leg. 2 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 29. IV. 1966, Native Collector leg. 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 15. V. 1966, Native Collector leg. 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 31. V. 1966, Native Collector leg. 1 3 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 15. VI. 1966, Native Collector leg. 3 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 30. VII. 1966, Native Collector leg. 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 15. XI. 1966, Native Collector leg. 1 Ƥ (BIMH); Vientiane Prov., Ban Van Eue, 1.–15. IX. 1967, Native Collector leg. 2 Ƥ (BIMH); Houa Phan Prov., Vieng Xai, am Fenster eines Gebäudes, 11. VIII. 2004, M. Geiser leg. 1 Ƥ (CMG); Houa Phan prov., ca. 3 km NE Sam Neua, 1030 m, 20°25.45' N / 104°04.50' E, village, visiting flowers of Sambucus sp., 2. VI. 2009, Michael Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 1 Ƥ (NHMB); Hua Phan prov., ~ 20°12' N / 104°01' E, Phu Phan Mt. 1500–1900 m, 17. V.–3. VI. 2007, M. Brancucci leg. 1 3 (NHMB); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1340–1870 m, 20°12–13.5' N / 103°59.5– 104 °01' E, 10. V.–9. VI. 2009, C. Holzschuh & local coll. leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 4 Ƥ (NHMB); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1340–1870 m, 20°12–13.5' N / 103°59.5– 104 °01' E, 2.–9. VI. 2009, M. Brancucci leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 1 3 3 Ƥ (NHMB); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1300–1900 m, 20°11–13' N / 103°59– 104 °01' E, flying around in village (daytime), 10.–17. VI. 2009, Michael Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 3 3 3 Ƥ (NHMB); Houa Phan prov., Muang Sop Bao, ca. 300 m, 20°42.39' N / 104°23.66' E, flying around in town, 6. VI. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 3 1 Ƥ (NHMB); Houa Phan prov., Ban Pahang, 1370 m, 20°43.35' N / 104°29.22' E, flying around in village, 7. VI. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 3 1 Ƥ (NHMB); Houa Phan prov., Muang Et env., Ban Muang Van, 360 m, 20°49' N / 104°03' E, flying around in village (daytime), 3.–6. VI. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 1 3 5 Ƥ (NHMB); Xieng Khouang prov., Phonsavan town ⊣ Phou Phadeng Mt., 1100–1200 m, 19°26' N / 103°13' E, flying around in town (daytime), 9. V. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 Ƥ (NHMB); Xieng Khouang prov., Phonsavan town ⊣ Phou Phadeng Mt., 1100–1200 m, 19°26' N / 103°13' E, flying around in town (daytime), 18.–24. VI. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 1 Ƥ (NHMB); "Rég. de Xieng-Khouang, Trannhinh, 1200 A 1200 M., R. Miéville 1920" 1 Ƥ (MNHP); Xieng Khouang prov., Ban Na Lam ⊣ Phou Sane Mt., 1300–1500 m, 19°37–8' N / 103°20' E, village, visiting flowers of Sambucus sp., 27.–30. V. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 4 3 1 Ƥ (NHMB), 1 3 1 Ƥ (CMG); Xieng Khouang prov., Muang Khoun, 1070 m, 19°19.91' N / 103°21.99' E, flying around in town (daytime), 22. VI. 2009, M. Geiser & D. Hauck leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 3 1 Ƥ (NHMB); Xieng Khouang prov., Ban Thaviang, 400 m, 19°02' N / 103°23' E, flying around in village (daytime), 21.–22. VI. 2009, M. Geiser & D. Hauck leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 3 2 Ƥ (NHMB); "Tonpheng, 14. IX. 1965, Native Collector" 1 Ƥ (BIMH); "Tonpheng, 20. IV. 1966, Native Collector" 1 Ƥ (BIMH); "Sedone Prov., Pakse, 23. V. 1965, P. D. Ashlock Collector" 1 3 (BIMH); "Sedone Prov., Pakse, 5. VII. 1965, Native Collector" 1 Ƥ (BIMH); "Sedone Prov., Pakse, 3. VIII. 1965, Native Collector" 1 Ƥ (BIMH); " Ile de Khong, 6. II. 1965, Native Collector" 1 Ƥ (BIMH).

CAMBODIA: " Cambodja, Mouhot, Fry Coll., 1905.100" 1 Ƥ (NHML); " Cambodge, A. Pavie 1886" 2 Ƥ (MNHP); " Cambodge, A. Pavie 1893" 1 3 9 Ƥ (MNHP); " Cambodge, Harmand 1875" 1 3 3 Ƥ (MNHP); " Cambodge, E. Le Moult 1916" 1 Ƥ (MNHP); " Cambodge " 1 3 1 Ƥ (MNHP).

VIETNAM: " Vietnam bor., Sevčik, 7. 81" 1 3 (NHMB); "Song-Chay, 1908, Indo Chine, coll. Dussault" 1 3 2 Ƥ (NHMB); "Rivière Noire, 1909, Indo Chine, Coll. Dussault" 1 Ƥ (NHMB); " Tonkin, Langue 1887" 3 Ƥ (MNHP); " Tonkin, Pethellaz.-45–97." 1 3 (MNHP); " Tonkin, Collection Le Moult" 1 Ƥ (NMPC); " Tonkin " 3 3 (MNHP); " Tonkin, Coll. E. Reitter, B. M. 1934-449." 1 Ƥ (NHML); "Haut-Tonkin, Roget.–111– 96" 2 Ƥ (MNHP); " Tonkin, région de Tien-Yen, E. de la baie d'Alorez, région littorale, Lieut. Poimeur 1908" 1 3 (MNHP); " Tonkin, Than-Moi, Juni-Juli, H. Fruhstorfer" 1 3 (MNHB); " Tonkin, Montes Mauson, Apr. Mai, 2–3000´, H. Fruhstorfer" 1 3 1 Ƥ (MNHB); "Lac Tho, Tonkin, de Cooman, coll. W. Wittmer" 1 3 (NHMB); " Tonkin, Cho-Moi, Roget 111–96" 1 Ƥ (MNHP); " Tonkin (Monkay), Brocars 1894" 1 3 2 Ƥ (MNHP); " Tonkin, Dongvan, 6 / 5 1917, R. Vitalis de Salvaza" 1 Ƥ (MNHP); " Tonkin, Phuly, V. 1917, R. Vitalis de Salvaza" 1 Ƥ (MNHP); " Tonkin, Bao-Ha, 1923, H. Stevens, Sladen-Godman Trust Exped., B. M. 1924–329" 1 Ƥ (NHML); " Tonkin, Bao-Ha, 7. XII. 1923, H. Stevens, Sladen-Godman Trust Exped., B. M. 1924–329" 1 Ƥ (NHML); " Tonkin, Région d'Hanoi, Su-Yut près Cho-Bo, Louis Duport 1909" 1 Ƥ (MNHP); "P. Hanoi, 56, lgt. Dr Syrûček" 1 Ƥ (NMPC); "Hanoi" 1 Ƥ (MNHP); "Hanoi, 3201" 1 Ƥ (NHML); "Lang-Son et Cao Bang, Girard 1894" 1 3 8 Ƥ (MNHP); „Ban-danh Lang-son, E. Autran, 1898“ 3 Ƥ (MHNG); "Lang Son ( Tonkin)" 2 Ƥ (MNHP); 52 km SW of Lang Son, 21°35' N 106°30' E, 370 m, 27. IV.–6. V. 1996, Pacholátko & Dembický leg. 14 3 4 Ƥ (NHMB); "Hoo-Binh, Tonkin " 1 Ƥ (MNHP); " Tonkin, Hoabinh. Aug. 1918, R. V. de Salvaza" 1 Ƥ (MNHP); Ha son binh prov., Hoa binh, 4.–7. VI. 1986, J. Rybniček leg., Freiwilliger Museumsverein Basel 1987. 1 Ƥ (NHMB); Hoa Binh, coll. W. Wittmer 5 3 1 Ƥ (NHMB); Ha Son Binh prov., Than Loc, singling, No. 112, 27. X. 1986, Mészáros, Oláh & Vásárhelyi leg. 1 3 (MTMB); "Central Tonkin, Chiem-Hoa, Aug. Sept., H. Fruhstorfer" 1 3 (MNHB). "Central Tonkin, Chiem-Hoa, Aug. Sept., H. Fruhstorfer" 5 3 2 Ƥ (MNHB); " Tonkin Centr., Région de Chim-Hoa et de Tuyen-Quan, A. Weiss 1901, Été" 3 Ƥ (MNHP); " Tonkin, Prov. de Tuyen-Quan, haute rivière claire, A. Weiss 1901, Été" 1 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, Print Été" 3 3 4 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, mars-avril" 1 3 4 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, avril 1901, case I" 1 3 7 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, avril, case II" 1 3 2 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, avril-juin" 5 3 15 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, juillet" 1 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, juill.-sept." 1 3 3 Ƥ (MNHP); " Tonkin Centr., Env. de Tuyen-Quan, A. Weiss 1901, oct.-déc." 1 Ƥ (MNHP); " Tonkin, Ha-Giang, Cap e Bonifacy 1903" 1 3 3 Ƥ (MNHP); " Tonkin, Vallées de la haute riv. claire, entre Hagiang et Vinh-Tuy, principalement a Bac-Quang, J. de Retz 1908" 7 Ƥ (MNHP); "Hagiang-Pa-kha, 1901–06, Indo Chine, coll. Dussault" 3 3 7 Ƥ (NHMB); "Pa-kha, 1914, Indo Chine, coll. Dussault" 1 3 1 Ƥ (NHMB); "M es du Ht Song Chai, Rabier 258–95" 3 3 3 Ƥ (MNHP); Son La prov., Moc Chau, singling, No. 93, 26. X. 1986, Mészáros, Oláh & Vásárhelyi leg. 1 Ƥ (MTMB); Vinh Phu Prov., Tam Dao, 29.–31. V. 1997, Y. Arita et al. leg., Exot. Insect Coll., Nos. 22155–22157. 2 3 1 Ƥ (KMNH); Vinh Phu Prov., Tam Dao, 29. V.–3. VI. 1997, Native leg., Exot. Insect Coll., No. 19907. 1 Ƥ (KMNH); N Vietnam, 75 km NW Hanoi, Tam Dao, 16.–23. V. 1991, J. Strnad leg. 2 Ƥ (NHMW); Vinh phu prov., Tam Dao, 27. V.–2. VI. 1986, J. Rybniček leg., Freiwilliger Museumsverein Basel 1987. 5 3 2 Ƥ (NHMB); Vinh Phu prov., Tam-Dao, 20. V. 1989, J. Strnad leg. 2 3 (NHMB); Vinh Phu prov., Tam Dao, 12.–24. V. 1989, Jan Strnad leg. 1 3 (NHMB); Vinh Phu prov., Tam Dao, 20.–28. VII. 1990, Jan Strnad leg. 1 3 (NHMB); Vinh Phu prov., Tam Dao, 6.–9. V. 1990, L. Dembický leg. 1 3 (NHMB); Vinh Phu prov., Tam Dao, V. 1990, J. Picka leg. 1 Ƥ (NHMB); 70 km NW of Hanoi, Tam Dao, 21°27' N 105°39' E, 900–1200 m, 1.–8. VI. 1996, Dembický & Pacholátko leg. 1 3 1 Ƥ (NHMB); 70 km NW of Hanoi, Tam Dao, 21°27' N 105°39' E, 900– 1200 m, 1.–8. VI. 1996, Pacholátko & Dembický leg. 2 3 (NHMB); Yen Bai ⊣ Tam Dao, 17. V. 1990, Vit Kubán leg. 1 Ƥ (NHMB); Yen Bai, Hoang Lien Son distr. 16.–17. V. 1990, J. Horák leg. 1 3 1 Ƥ (NHMB); Hoang lien son prov., Yen Bai, V. 1990, J. Picka leg. 1 Ƥ (NHMB); "Yen-Bay, 1907, Indo Chine, coll. Dussault" 1 Ƥ (NHMB); Yen Bai, Hoang Lien Son distr. 16.–17. V. 1990, J. Horák leg. 1 3 1 Ƥ (NHMB); Hoang lien son prov., Yen Bai, V. 1990, J. Picka leg. 1 Ƥ (NHMB); " Tonkin Centr., Env. de Yen-Bai, A. Weiss 1901" 3 Ƥ (MNHP); " Tonkin, Lao-Kay, Dr Chevalier 1902" 1 3 (MNHP); " Tonkin, env. immédiats. de Lao- Kay, Cap ne Sauvez 1908" 1 3 1 Ƥ (MNHP); "Chapa, prov. de Laokay. Ht-Tonkin" 1 Ƥ (MNHP); " Tonkin, Chapa, mai 1916, R. Vitalis de Salvaza" 1 Ƥ (MNHP); "Chapa" 1 Ƥ (MNHP); Hoang lien son prov., Sa Pa, VI. 1990, Duang Tat Tu leg. 1 3 (NHMB); Sa Pa, Lao Cai prov., 250 km SW Hanoi, Sapa vill., env. Hoang Lien Son Nat. Res., 1250 m NN, 31. V.–3. VI. 1998, A. Napolov leg. 1 3 (NKME); "Frontière Chine-Tonkin, Région de Lao-Kay et Ho-Kheou, Ch. Dupont 1900" 1 3 15 Ƥ (MNHP); "Bao-Lac ( Tonkin)" 1 3 (MNHP), 2 3 (NMPC); "Bao-Lac ( Tonkin), coll. W. Wittmer" 1 3 1 Ƥ (NHMB); Bac Kan Prov., Ba Be National Park, 300 m, 30. IV. 2006, M. Wakabayashi leg. 1 Ƥ (KMNH); Bac Kan Prov., Ba Be National Park, 650–700 m, 12. IV. 2006, M. Wakabayashi leg. 1 Ƥ (KMNH); Trung Trang prov., Cat Ba, 200 m, 20°48' N / 107°00' E, at light, No. 171, 16. V. 1987, Matskási, Oláh & Topál leg. 2 3 (MTMB); Quang ning prov., Ha long, 29. V.–1. VI. 1985, J. Picka leg. 1 3 (NHMB); N Vietnam, Cuc-phuong, 25. 5. 1986, J. Macek leg. 1 Ƥ (NMPC); Ninh Binh Prov., Cuc Phuong, 27. IV. 1996, Y. Okushima leg. 1 3 1 Ƥ (KMNH) 1 3 (CMG); Ninh Binh Prov., Cuc Phuong, 24.–29. IV. 1998, Y. Arita leg. 1 Ƥ (KMNH); 100 km S from Hanoi, Cuc-Phuong nat. park, 2.–12. V.

1991, E. Jendek leg. 3 3 2 Ƥ (NHMB); Cuc-Phuong, 2.–11. V. 1991, Jan Strnad leg. 1 3 2 Ƥ (NHMB); " Annam " 1 Ƥ (NMPC); " Annam, Thua-Luu, 0–800 m, 8. 8. 1938, Björkegren" 3 Ƥ (NRMS); " Annam, B'sré (80 km NNW fr. Djiring), 1000 m, 7.–38, Björkegren" 1 Ƥ (NRMS); " Annam, Quang-Tri, M. Maunier 1922" 1 3 (MNHP); "Quantri, Annam, coll. Staudinger" 1 3 2 Ƥ (SMTD); " Annam, Phuc-Son, Nov. Dez., H. Fruhstorfer" 1 Ƥ (MNHB); "Cochinch., Morica" 1 Ƥ (MNHP); "Cochinchine, Amiral Vignes 1898" 18 3 24 Ƥ (MNHP); "Cochinchine, Harmand 1872" 2 3 2 Ƥ (MNHP); „Cochinchine, P. de Borre, Coll. D. Ach. 1. 90“ 1 3 5 Ƥ (MHNG); "Cochinchine, Tay Ninh, Fouquet, 1904" 5 3 3 Ƥ (MNHP); "Cochinchine, Bienhoa, 12. 1902, Cap. Fouquet" 3 3 2 Ƥ (MNHP); "Cochinchine, Env. de Saigon, Simard 1902" 1 3 (MNHP); "Saigon, H. Swale, 1913–117" 1 Ƥ (NHML); "Saigon" 1 3 3 Ƥ (CMG), 1 3 (NHMW); „Saigon, P. de Borre“ 1 Ƥ (MHNG); "Cochinchine, Cap Saint-Jacques, Cap ne Modest 1909" 1 Ƥ (MNHP); "Cap St. Jacques, Coll. A. Bonhoure 1909" 1 Ƥ (MNHP); "Cap St. Jacques (Cap'. Modest), Coll. A. Bonhoure 1909, Novembre" 1 Ƥ (MNHP); "Cochinchine, Env de Baria, Cap ne Modest 1907" 1 3 (MNHP); S-Vietnam, Nam Cat Tien Nat.Park, 1.–15. V. 1994, Pacholatko & Dembicky leg. 2 3 1 Ƥ (NHMW).

THAILAND: „ Siam “ 1 Ƥ (MHNG), 1 Ƥ (MNHP); " Siam, Bocourt 327–02" 9 3 8 Ƥ (MNHP); " Siam, Bowring, 63 47*" 1 3 7 Ƥ (NHML); " Siam, S. S. Flower, 98.217" 1 3 (NHML); " Siam, [locality written in Thai], 16. Mar. 1943, Lot 918, To: Czecho, List: 068–05, Lot: 918/56" 1 Ƥ (NHMB); " Siam, [locality written in Thai], 7. Oct. 1959, Lot 609, To: Czecho, List: 068–05, Lot: 609/56" 1 Ƥ (NHMB); Chiang Mai, 29. IV. 1985, Sab. Steinke leg. 1 Ƥ (SMNS); Chiang Mai, 9. V. 1985, Sab. Steinke leg. 1 3 (SMNS); Chiang Mai Pref., Doi Suthep, 1.–5. VI. 1986, H. Hirasawa leg., Exot. Insect Coll., Nos. 9877–9890. 6 3 8 Ƥ (KMNH); Chiang Mai, 18.49° N, 99.00° E, Doi Suthep to Doi Pui, 19.–23. IV. 1991, L. Dembický leg. 1 Ƥ (NHMB); Doi Pui, 17. V. 1985, Sab. Steinke leg. 1 3 (SMNS); Chiang Mai, NW Mae Taeng, 25. IV. 2003, R. Grimm leg. 1 3 (SMNS); Maetaeng, Chiang Mai, 21. VII. 1991. 2 3 (NHMB); N-Thailand, Pai, 22.–29. V. 1996, A. Kudrna jr. leg. 1 3 (CMG); Mae Hong Son prov., Soppong env., 600 m, 28. V.–2. VI. 1999, M. Riha leg. 1 3 1 Ƥ (NHMB); Namuang, 21.–26. V. 1993, Pacholatko & Dembicky leg. 1 Ƥ (NHMB); " Siam, Tak, 20. Aug. 1961, Lot 2783, To: Czecho, List: 068–05, Lot: 2783/81" 1 3 1 Ƥ (NHMB); Medaw, Burmese frontier, 4. XI. 1934, Malaise leg. 1 Ƥ (NRMS); Nan Pha Khab, 11.–15. V. 1993, Pacholatko & Dembicky leg. 1 Ƥ (NHMB); Sangkhlaburi, 8.–12. IV. 1991, Jan Farkač leg. 1 Ƥ (NHMB); "Bangkok, Larnaudie 349–64" 2 Ƥ (MNHP); Bangkok, IX. 1919, F. J. Godfrey leg., 1920–244. 1 Ƥ (NHML); "Ransont, 1871, Bankok" 2 3 2 Ƥ (NHMW); Koh Samui, 10. I. 1988, Madl leg. 1 Ƥ (NHMW); Ko Samui isl., Coral Cave, 23. IV. 1991, F. Hiess leg., Sammlung A. Kopetz 1 Ƥ (NKME); "Prachuab khiri khan, May 5. 1961, Lot 2747, To: Czecho, List: 068–05, Lot: 2747/56" 1 Ƥ (NHMB); Chumphon prov., Pha To env. 9° 48' / 98° 47°, 27. III.–14. IV. 1996, P. Prudek leg. 1 Ƥ (NHMB); Betong, Gunung Cang dun vill., Yala dist., 25. III.–22. IV. 1993, J. Strnad leg. 1 3 (NHMB); "Siamese Malay States, Jalor: Biserat, 11. VII. 1901, Coll. N. Annandale and H. C. Robinson, No. 181, 1903–281" 1 Ƥ (NHML); "Setul, Siam, H. N. Ridley, 1910–264." 1 Ƥ (NHML).

MALAYSIA: " Malacca " 1 Ƥ (MNHB); " Malacka, Franklin Müller" 1 3 1 Ƥ (DEIM); " Malakka, Coll. Franklin" 2 Ƥ (DEIM); "Malay, Casteln, Fry Coll., 1905.100" 2 Ƥ (NHML); "Fed. Malay States, 1909, C. J. Brooks, B. M. 1931–570" 2 3 3 Ƥ (NHML); " Malacca, Pascoe Coll., 93–60" 1 Ƥ (NHML); "Borneo North, Whitehead, Fry Coll., 1905.100" 1 Ƥ (NHML). PENANG: "P. Penang, 600–800 m, II. 89, Loria et Fea" 1 3 1 Ƥ (MCGD); "P. Pinang, Batu Ferringghi", 28. VII. 1975, Kling leg. 1 3 2 Ƥ (NHML); "Penang, Bowring 63 47*" 5 3 5 Ƥ (NHML); "Penang, 96–85" 1 Ƥ (NHML); "Penang, H. N. Ridley, 1903–229" 1 Ƥ (NHML); Bukit Bakar Kapor, 24. XI. 1961, H. T. Pagden leg., Pres. by Com. Inst. Ent., B. M. 1962–1, 1 Ƥ (NHML). PERAK: " Malakka, Perak" 4 Ƥ (MNHB); " Malacca, Perak, Jachan V." 1 Ƥ (MNHB); "Perak, Sharp Coll. 1905–313" 1 3 3 Ƥ (NHML); "Taiping, Malay States, W. B. Orme, 1911–131" 1 Ƥ (NHML); Taiping, 1972, Liew leg. 1 Ƥ (MHNG); Maxwell’s Hill, 6. I. 1974, Pfanner leg. 1 Ƥ (MHNG); E Taiping, 500–800 m, V.-VI. 1978, H. Knorr leg. 2 Ƥ (SMNS); " Indien, Rwela Rangsar, Perak" 1 Ƥ (MNHB); "Tapah, Perak, G. B. Cerruti 1900" 1 Ƥ (MNHP). PAHANG: "Pahang, Bingham, 1903–183" 1 Ƥ (NHML); Fraser's Hill, 3500 ft. 7. VI. 1941, H. M. Pendlebury, F.M.S. Museums, Ex F.M.S. Museum, B. M. 1955–354 1 Ƥ (NHML). NEGERI SEMBILAN: "Negri Sembilan F. M. S., Port Dickson, IX. 1941, Ex F.M.S. Museum, B. M. 1955–354" 1 3 (NHML). SELANGOR: Batu Cave prés Kuala Lumpur, 24. VII. 1969, R. Pilet leg. 1 Ƥ (MHNG). SARAWAK: "Borneo, Sarawak, 1865–66, Coll. G. Doria" 4 Ƥ (MCGD), 1 Ƥ (MNHP); "Borneo, Xántus, Sarawak " 2 Ƥ (MTMB); " 1907–1909, C. J. Brooks, B. M. 1936–681" 1 3 3 Ƥ (NHML); "1909, C. J. Brooks, B. M. 1931–570" 1 3 (NHML); "Puak, Sarawak, G. E. Bryant 2. V. 14, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Puak, Sarawak, G. E. Bryant 2. V. 14, G. Bryant Coll. 1919–147" 2 Ƥ (NHML); "Lundu, Sarawak, G. E. Bryant, 7. I. 14, G. Bryant Coll. 1919–147" 1 Ƥ (NHML), "Quop, W. Sarawak, II.–III. 1914, G. E. Bryant, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Quop, W. Sarawak, IV. 1914, G. E. Bryant, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Kuching, Borneo, Andrewes Bequest., B. M. 1922–221" 3 3 (NHML); "Kuching, J.E.A. Lewis, 1910–116" 2 3 2 Ƥ (NHML); "Matang Rd, 3 1/ 2 miles, 8. 5. '09, J. E. A. Lewis. 1910–116." 1 3 1 Ƥ (NHML); "Matang, 6. 11. 14, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Mt. Matang, W. Sarawak, G. E. Bryant, I. 1914, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "S. Tengah, 10. 5. 0 9, Sarawak, J.E.A. Lewis, 1910–116." 1 Ƥ (NHML). SABAH: "Borneo, Trusan, Aug. 1900 " 1 3 (MNHP); "Sandakan, C. V. Creagh., 96–197" 1 Ƥ (NHML); "Sandakan, Brit. N. Borneo, W. B. Pryer, 1914-484" 1 3 1 Ƥ (NHML). BANGGI (Sabah): "Banguey" 1 3 (MNHP); "Banguey b. Borneo" 1 3 (ZSMU); "Banguey b. Borneo, coll. Staudinger" 16 3 10 Ƥ (SMTD); "Bangkei, H. Kühn, 1885, coll. Staudinger" 1 Ƥ (SMTD).

SINGAPORE: "Sing., Wallace" 1 Ƥ (MNHP); " Singapore " 3 Ƥ (MNHB); "C. J. Saunders, B. M. 1933– 227" 1 3 1 Ƥ (NHML); "H. N. Ridley, 1900–82 " 1 Ƥ (NHML); "H. N. Ridley, 1900–116" 1 Ƥ (NHML); "H. N. Ridley, 1901–77 " 1 Ƥ (NHML); "H. N. Ridley, 1904-47 " 1 3 (NHML); "H. N. Ridley, 1904–160" 1 Ƥ (NHML); "H. N. Ridley, 1907–51 " 1 Ƥ (NHML); "H. N. Ridley, 1909–194" 5 3 2 Ƥ (NHML); "H. N. Ridley, 1910–284" 1 Ƥ (NHML); "G. Lewis, 1915–38 " 1 3 (NHML); " Singapore, 1962, B. C. Chubman" 1 Ƥ (NHML); "Botanic Gdns. Singapore, 94–132." 1 Ƥ (NHML).

BRUNEI: " Brunei, Borneo" 1 3 (MNHP), 1 Ƥ (MTMB).

INDONESIA: "Sunda Ins., Excell. v. Sundt G." 1 Ƥ (MNHB). SUMATRA: „Sumatra“ 1 Ƥ (MHNG); "Sumatra, coll. Hayek" 1 3 (ZSMU); "Sumatra, coll. Staudinger" 2 Ƥ (SMTD); "Sumatra, Coll. Kraatz" 1 3 4 Ƥ (DEIM); "Sumatra, Dr. Will" 2 Ƥ (MNHB); "Sumatra, Dr. Will, coll. Hauser" 1 Ƥ (MNHB); "Sumatra, Dehy" 1 Ƥ (MNHP); Aceh, 16 km S Kutacane, 350 m, 18. VIII. 1992 1 Ƥ (NHMB); "Pajakumbo" 1 Ƥ (MNHP); Bukit Tinggi, 23. II. 1954, A. H. G. Alston, B. M. 1954-414 1 Ƥ (NHML); Bukittinggi env., 14.–16. II. 1991, Bocák & Bocákova leg. 1 Ƥ (NHMB); "Deli, S. Wiene, F. R. Mason, Ex Coll. F. Mason, Brit. Mus. 1922–173" 1 Ƥ (NHML); "Deli '01" 1 Ƥ (DEIM); "N. O. Sumatra, Deli, L. Martin S." 1 3 6 Ƥ (MNHB); "E Coast Sumatra, Doerian Moelan, Brindjei, Lt. R. Coughtrie, 1915–184" 1 Ƥ (NHML); "Medan S O.K, L. Fulmek, Tabakopslagplaats" 1 Ƥ (NHMW); N-Sumatra, D. Toba, Samosir, Tuk-Tuk, 3. II. 1990, Schillhammer leg. 1 Ƥ (NHMW); N-Sumatra, Gunung Sibayak/Brastagi, N Toba See, 1500–2000 m, 20. II. 1990, Schödl leg. 1 Ƥ (NHMW); "N. O. Sumatra, Tebing-tinggi, Dr. Schultheiss, Coll. Schultheiss" 3 3 4 Ƥ (DEIM); "Palembang, Sumatra 1900, Jos. Schmitz, Coll. Kraatz" 1 Ƥ (DEIM); "Sumatra, Beskera" 1 Ƥ (MNHB); "Balighe, X. 90 – III. 91, E. Modigliani, coll. W. Wittmer" 1 Ƥ (NHMB); "Medan S. O.K., L. Fulmek, coll. W. Wittmer" 1 Ƥ (NHMB); "Indragiri, Sumatra, V. Meckel 99" 1 3 1 Ƥ (NHMB); Utara, Berastagi, 1300 m, 20. X. 1991, W. Barries leg. 1 Ƥ (NHMB); "Lampung, Buxton, Fry Coll., 1905.100" 3 Ƥ (NHML). RIAU: Penyengat Island, 7.–11. VII. 1991, B. Baláza leg. 1 Ƥ (MTMB). NIAS: "Is. Nias, 1897–98, U. Raap" 3 Ƥ (MCGD); " Ile Nias" 1 3 (MNHP); "Nias, Coll. Kraatz" 4 Ƥ (DEIM). BATU: "Isole Batu, 1896– 97, H. Raap" 1 Ƥ (MCGD). MENTAWAI: S-Siberut, w Muarasiberut, Toteburu, 16. II. 1991, Schillhammer leg. 1 3 (NHMW); Siberut, Madobak, 18. II. 1991, Schödl leg. 1 3 1 Ƥ (NHMW); Siberut, Madobak, W Muarasiberut, 19. II. 1991, Schödl leg. 1 Ƥ (NHMW); Sipora, 20. X. 1924, H. H. Karny 1 Ƥ (NHML); "Sipora, Sereina, V.-VI. 94, Modigliani" 3 Ƥ (MCGD). ENGGANO: "Engano, Kifa-juc., Modigliani, V. 1891 " 1 Ƥ (MCGD); "Engano, Bua-Bua, Modigliani, V. VI. 1891 " 1 Ƥ (MCGD). KALIMANTAN: "S. O. Borneo, Grabowsky S. V." 1 Ƥ (MNHB); "Bornéo, Lohaban, R. Oberthür 1898" 2 Ƥ (MNHP); "Pontianak, Bruegel, 2. 1907." 1 Ƥ (ZSMU); "Ponteanak, Borneo, VII. 07 " 12 3 6 Ƥ (BIMH), 1 3 1 Ƥ (CMG); "Borneo, Pontianak, F. Muir, VII. 07 " 3 3 1 Ƥ (BIMH); "Borneo, Pontianak, F. Muir, Collection of W. M. Giffard" 1 3 (BIMH); "Borneo, Pontianak, Dr. Will" 2 Ƥ (MNHB); "W. Borneo, Telok Ayer, F. Muir" 1 Ƥ (BIMH); "W. Borneo, Telok Ayer, F. Muir, VII. 07 " 1 3 (BIMH); "Martapura, S. E. Borneo, Doherty, 1891, Sharp Coll., 1905–313." 1 Ƥ (NHML); "Sanga Sanga, Moorjawa, E. Borneo, H. D. Jensen. 1906-45." 2 Ƥ (NHML); "Sampit, Borneo, S= Rupert" 1 3 1 Ƥ (MNHB); "Maratoea Island, O. Borneo, Mjöberg" 1 Ƥ (NRMS). JAVA: " India or., coll. E. Friv., Jawa" 1 3 2 Ƥ (MTMB); "Java" 1 3 1 Ƥ (DEIM), 4 Ƥ (MNHB), 2 Ƥ (NHMB), 1 Ƥ (MHNG); "Java, Dr. Will" 1 3 1 Ƥ (MNHB); "Java, ancienne coll." 1 3 (MHNG); „Java, P. de Borre“ 2 3 3 Ƥ (MHNG); "Java, coll. Staudinger" 1 Ƥ (SMTD); "Java, Nyman" 1 3 (NRMS); "Java, Mellb." 1 Ƥ (NRMS); "Java, J. D. Pasteur 268–94" 1 3 1 Ƥ (MNHP);: "Java, 48 92" 1 Ƥ (NHML); "Java, Bowring 63 47*" 1 3 1 Ƥ (NHML); "Java, Horsfield, 60– 15, 1 3 (NHML); "Java, Coll. Kraatz" 1 3 10 Ƥ (DEIM); "Fruhstorfer, Java, Coll. Kraatz" 1 3 (DEIM); "Pfeiffer" 1 Ƥ (NHMW); "Warsberg, 868" 2 Ƥ (NHMW); "Java: Pelabuan Ratu" 1 Ƥ (DEIM); "Linga, Röttger, Hist.-Coll. ( Coleoptera ), Nr. 32614" 1 Ƥ (MNHB); "Java, Linga, Hist.-Coll. ( Coleoptera ), Nr. 32614" 1 3 2 Ƥ (MNHB); "Preanger, Java, Ankauf Muche" 1 Ƥ (SMTD); "Coll. F. C. Drescher, Java, Preanger, N. O. I. Bandoeng, III. 1936, 900 m, coll. W. Wittmer" 1 Ƥ (NHMB); "Coll. F. C. Drescher, Java, Preanger, N. O. I. Bandoeng, 8. V. 1936, 750 m, coll. W. Wittmer" 1 Ƥ (NHMB); "V. Java, Bandoeng, 700 m, 2/ XII. 1943, Kurt Landberg" 1 Ƥ (NRMS); "C. O. Nandjoeng-Djaja, G. Simpal, 600 Mr., Preanger, Java, II. 1934, leg. C. P. J. de Haas, coll. W. Wittmer" 1 Ƥ (NHMB); "Buitenzorg, +/– 1000 m, Febr. 90. I. Z. Kannegieter" 1 3 (MNHP); "Java, Buitenz., Kember" 2 Ƥ (NRMS); "Buitenzorg, 25. XII. 1921, Karny" 1 Ƥ (DEIM); "B'zorg, III. 32 " 1 Ƥ (NHMB); "Buitenzorg, M. Fleischer S. V. 98 " 1 Ƥ (MNHB); "Java occident., Pengalengan 4000, 1893, H. Fruhstorfer" 1 Ƥ (MNHP); "Boeloe Lawang, Res. Pasoeroean (Holz)" 1 3 (MNHP), 1 Ƥ (NMPC); "Gaharan, Res. Besoeni, E. Java, coll. W. Wittmer" 1 Ƥ (NHMB); "Silosanen, E. Java, coll. W. Wittmer" 1 Ƥ (NHMB); "Soekaboemi, Java, leg. Dr. F. Weber" 1 Ƥ (NHMB); Batjang, Mt. Gedeh, 1100 m, IX.-X. 1919, R. Ratter cond. 1 Ƥ (NHMB). SUMBAWA: " Insel Sumbawa, coll. W. Staudinger" 2 Ƥ (SMTD); Tambora, 600 m, 9. XI. 1941, Rolf Blomberg leg. 2 Ƥ (NRMS); "B. Aroe Hassa, Sambawa, 2–5000', Doherty IX. X., Brit. Mus. 1923–320 2 Ƥ (NHML); Batudulang, 30 km S of Sumb. Besar, 1000 m, 10. II. 1994, Bolm leg. 1 Ƥ (SMNS). KOMODO: " Flores I., Komodo I., J. K. de Jong, VII. 1937, coll. W. Wittmer" 1 Ƥ (NHMB). FLORES: "Sunda Ex. Rensch, Ende – Flores, 15. VI. 1927 " 1 Ƥ (MNHB). SUMBA: Lindiwatju, 29. IX. 1949, Dr. Bühler & Dr. Sutter leg. 1 Ƥ (NHMB). ADONARA: "Andonare, Laboenarang, 2-4000', Doherty XI., Brit Mus. 1923–320" 1 Ƥ (NHML). ROTE: Rote, IX. 1935, C. Bühler & Meyer leg. 2 Ƥ (NHMB). TIMOR (west): Soe, XII. 1951, Handschin leg. 1 3 1 Ƥ (NHMB); "Gunung Leo, Ned. Timor, 2-4000', Doherty XI. XII., Brit. Mus. 1923–320" 2 Ƥ (NHML). SULAWESI: "I. Celebes" 1 Ƥ (NHMP); "Celebes, 55 22" 1 3 (NHML); "Celebes, 57 101" 1 Ƥ (NHML); "Célébes, 2037–77, de la Savinière" 1 Ƥ (MNHP); "S. Celebes, 250 m, 3. 1941" 1 Ƥ (NHMB); "Drs. Sarasin, S. Celebes, Makassar" 1 Ƥ (NHMB); "Drs. Sarasin, C. Celebes, Palu-Tal" 1 Ƥ (NHMB); "Drs. Sarasin, N. Celebes, Tondano" 1 Ƥ (NHMB); "Drs. Sarasin, S. O. Celebes, Ussu" 1 3 (NHMB); "Drs. Sarasin, Luwu-Djaladja" 1 Ƥ (NHMB); "Tondano, Celebes, coll. Staudinger" 7 3 7 Ƥ (SMTD); Kandari, III. 1874, O. Beccari leg. 2 Ƥ (MCGD); Kandari, VII. 1874, O. Beccari leg. 1 3 1 Ƥ (MCGD); "Nord-Celebes, Toli-Toli, Nov.-Dez. 1895, H. Fruhstorfer" 1 Ƥ (MNHP); Sulawesi Utara, Dumoga-Bone N. P., "Fog 9, Mixed crops", 1. III. 1985, R. Ent. Soc. London, Project Wallace 3 3 3 Ƥ (NHML); Sulawesi Utara, Dumoga-Bone N. P., Lowland forest 200– 300 m, on plants, I. 1985, R. Ent. Soc. London, Project Wallace 1 Ƥ (NHML); Sulawesi Utara, Dumoga-Bone N. P., Base camp area, ca. 190 m, XII. 1985, R. Ent. Soc. London, Project Wallace 1 Ƥ (NHML); Sulawesi Utara, Dumoga-Bone N. P., Base camp area, ca. 190 m, at light, X. 1985, R. Ent. Soc. London, Project Wallace 1 Ƥ (NHML); Sulawesi Utara, Dumoga-Bone N. P., Base camp area, ca. 190 m, at light, XI. 1985, R. Ent. Soc. London, Project Wallace 1 3 (NHML); Sulawesi Utara, Dumoga-Bone N. P., Plot A, ca. 200 m, Lowland forest, Flight intercept trap, VII. 1985, R. Ent. Soc. London, Project Wallace 1 3 (NHML); Tokala Atas, 20. II. 1980, M. J. D. Brendell leg., B.M. 1980–280. 1 Ƥ (NHML). BUTON: "Ins. Buton" 1 Ƥ (MNHB). TANIMBAR: "Jandema, Tenimber, Doherty, VI. VII., Brit. Mus. 1923–320" 1 3 4 Ƥ (NHML); "Larat, F. Muir, B. M. 1914-413" 2 Ƥ (NHML); "Larat, Malay, 12–07, Coll. F. Muir" 2 3 2 Ƥ (BIMH); "Larat Ins. (Tanimbar I.)" 1 3 2 Ƥ (MNHB). BACAN: "Ins. Batjan" 1 Ƥ (MNHP); "Batchian, Bowring, 63 47*" 1 Ƥ (NHML). BURU: "Buru, X. 1875, O. Beccari" 2 Ƥ (MCGD). AMBON: " Amboina, Malay , IV–08, Coll. F. Muir" 1 3 (BIMH); " Amboina, F. Muir, V –08" 1 3 (BIMH); " Amboina, F. Muir " 2 3 2 Ƥ (BIMH); "Amboyna, Bowring, 63 47*" 2 Ƥ (NHML); " Amboina, 1873 , O. Beccari" 1 Ƥ (MCGD); " Amboina , XII.

1874, O. Beccari" 1 3 2 Ƥ (MCGD); " Amboina, Coll. Kraatz " 1 Ƥ (DEIM); "Dr. Doleschal, 1859, Amboina " 1 Ƥ (NHMW); " Amboina , v. Martens, Hist.-Coll. ( Coleoptera ), Nr. 40761" 1 Ƥ (MNHB); " Amboina , v. Rosenberg" 1 Ƥ (MNHB); " Ambon, IV. 1941, Sei Waitoenoe, E. Lundquist, coll. W. Wittmer" 1 3 (NHMB); Wasi, 300–600 m, 6. II. 1989, Schillhammer leg. 1 Ƥ (NHMW); " Ambon I., Waai, 26. XII. 1961, A. M. R. Wagner Collector" 1 Ƥ (BIMH); "Waai, P. Ambon, 18. VII. 1959, leg. A. M. R. Wegner" 1 Ƥ (NHMB). CERAM: "Ceram, coll. W. Wittmer" 1 Ƥ (NHMB); "Ceram" 1 Ƥ (NHMB); "Illo, C. Ribbe, 1884, coll. Staudinger" 1 Ƥ (SMTD); Umg. Wahai, 12. II. 1989, Schödl leg. 1 3 1 Ƥ (NHMW); "Kisoci, Ceram" 1 3 1 Ƥ (MNHB); "Ceram, Illo, C. Ribbe 1884" 2 Ƥ (MNHB); Solea, 12 km SE Wahai, 17. I.–6. II. 1997, J. Horák leg. 1 Ƥ (NHMB); Solea, 12 km SE Wahai, 17. I.–6. II. 1997, S. Bilý leg. 2 Ƥ (NHMB). GORONG: "Gorom- Ins." 1 Ƥ (MNHB). WATUBELA: "Tevor Ins." 3 Ƥ (MNHB). TERNATE: " Molucche, Ternate, Coll. Bruijn, 1875" 1 Ƥ (MCGD). HALMAHERA: " Moluques, Gilolo, Raffray & Maindron 1878" 2 Ƥ (MNHP); "Halmahera, an Andai?, Beccari, 1875" 1 Ƥ (MCGD). ARU: "Isole Aru, Wokan, O. Beccari 1873" 4 Ƥ (MCGD); "Aru Insel, Wamma Dobbo, C. Ribbe 1883" 1 Ƥ (MNHB). MISOOL: "Mysol, 61 124" 2 Ƥ (NHML). WAIGEO: "Waigiou, 62 7 " 1 Ƥ (NHML).

TIMOR-LESTE: "Dili, Port. Timor, 2500', Doherty IV. V. Brit. Mus. 1923–320" 1 Ƥ (NHML).

PHILIPPINES: "Philipp" 1 Ƥ (NHMB); " Philippines " 5 3 4 Ƥ (NMBE); „ Philippines, W. Parsons“ 3 Ƥ (MHNG); "Cum 40." 1 Ƥ (NHMW); "Philipp, coll. Staudinger" 10 Ƥ (SMTD); "Ins. Philip., coll. Staudinger" 1 Ƥ (SMTD); "Phil Isla " 1 3 1 Ƥ (NHML); "Phil. Islds, 479" 1 Ƥ (NHML); "Phiillip Islands, Fry Coll., 1905.100" 1 Ƥ (NHML); " Philippine Is., E. M. Ledyard., B. M. 1925-491" 4 Ƥ (NHML); " Philippine Islands, Sharp Coll. 1905–313" 1 3 (NHML). LUZON: "Luzon, Jagor, Hist.-Coll. ( Coleoptera ), Nr. 39320" 1 Ƥ (MNHB); "Luzon, coll. W. Wittmer" 1 3 (NHMB); "Drasche, Luzon 1876" 1 Ƥ (NHMW); Prov. Aurora, Baler, 14. VI. 82, J. P. Schauenberg leg. 1 Ƥ (MHNG); "Paeta, Luzon, coll. Staudinger" 1 3 1 Ƥ (SMTD); Macobod, VII. 1915, coll. W. Schultze 1 3 1 Ƥ (SMTD); "Baguio, Luzon, coll. Staudinger" 1 3 (SMTD); "Rizal Prov., coll. W. Schultze" 1 Ƥ (SMTD); "Rizal, Montalban, III. 1915, coll. W. Schultze" 1 3 (SMTD); "Manila, Philipp., coll. Staudinger" 1 Ƥ (SMTD); "Luzon, P. I., Manila, coll. W. Schultze" 2 3 1 Ƥ (SMTD); "Manila, v. Möllendf., coll. Staudinger" 1 3 (SMTD); "Flach, Manila" 1 Ƥ (NHMW); "Manila, 1903–352." 1 Ƥ (NHML); "Manila, 92–20." 1 3 3 Ƥ (NHML); "Mt. Makiling, Laguna, P. I., IX. 4. 1930, F. C. Hadden Collection" 1 Ƥ (NHML); "Mt. Makiling, Laguna, P. I., VI. 26. 30, Collected by F. C. Hadden" 1 Ƥ (BIMH); "Mt. Makiling, Laguna, P. I., VII. 24. 30, F. C. Hadden Collection" 1 Ƥ (BIMH); "Mt. Makiling, Laguna, P. I., 9–16–30, Collected by F. C. Hadden" 2 Ƥ (BIMH); "Los Banos, Laguna, P. I., XI. 21. 30 " 1 Ƥ (BIMH); "Los Banos, Philippine Ids., 1916–1917, F. X. Williams Collector" 1 Ƥ (BIMH); "Los Banos, Philippine Ids., VII– 17–1921, F. X. Williams Collector" 1 Ƥ (BIMH); "Los Banos, Philippine Ids., XI–1921, F. X. Williams Collector" 5 3 2 Ƥ (BIMH); "Camarines Sur, Mt. Iriga, 500–600 m, 22. IV. 62, H. M. Torrevillas Collector" 2 3 1 Ƥ (BIMH); "Camarines Sur, Mt. Iriga, 500–600 m, 23. IV. 62, H. M. Torrevillas Collector" 1 Ƥ (BIMH); "Camarines Sur, Mt. Iriga, 500–600 m, 24. IV. 62, H. M. Torrevillas Collector" 3 3 (BIMH); "Camarines Sur, Mt. Iriga, 500–600 m, 27. IV. 62, H. M. Torrevillas Collector" 1 3 (BIMH); "Camarines Sur: Mt. Isarog, 20 km E of Naga, 500–600 m, 16. IV. 1963, H. M. Torrevillas, Light Trap" 1 Ƥ (BIMH); "Camarines Sur: Mt. Isarog, 750–850 m, 13.–15. V. 1963, H. M. Torrevillas Collector" 1 Ƥ (BIMH); "Camarines Sur: Mt. Isarog, 800–850 m, 25. IV. 1963, H. M. Torrevillas, Light Trap" 1 Ƥ (BIMH). MINDORO: "Mansalay, E. Taylor, coll. W. Schultze" 1 3 (SMTD); 28 km S Calapan, Balete, 27.–29. XI. 1992, Schillhammer leg. 2 3 1 Ƥ (NHMW); 28 km S Calapan, Balete, 27.–29. XI. 1992, Jäch leg. 1 Ƥ (NHMW); E Puerta Galera, Sabang, XI. 1992, Jäch leg. 1 3 2 Ƥ (NHMW); E Puerta G., Sabang, ex. sterc., 23. XI. 1992, Schillhammer leg. 1 Ƥ (NHMW). SAMAR: Catbalogan, 24. IV. 1915, Boettcher leg., coll. W. Wittmer 1 Ƥ (NHMB); "Catbalogan, Samar" 1 Ƥ (DEIM); "Catbalogan, coll. W. Schultze" 1 Ƥ (SMTD); "Osmena, Samar, P. I., VII–25. 45, K. S. Hagen Collector" 1 Ƥ (BIMH). MASBATE: "Aroroy, leg. Böttcher" 3 Ƥ (MNHB). PANAY: "Antique Culasi, coll. W. Schultze" 1 Ƥ (SMTD). NEGROS: Mt. Kanlaon env., 16. II. 1994, Graindl & Seyfert leg. 1 Ƥ (NHMW). CEBU: Cebu-City, 24. II. 1991. 1 Ƥ (SMNS). LEYTE: Visca, N Baybay, IV.-IX. 1991, Untersuchungsbelege, A. Engler leg. 1 3 1 Ƥ (SMNS); "Tacloban, coll. W. Schultze" 1 Ƥ (SMTD); "Palo, Leyte, 15 Jan 57 " 1 Ƥ (BIMH); "Palo, Leyte, 3. III. 57, WU7" 1 Ƥ (BIMH); "Palo, Leyte, 5. III. 57, WU7" 1 Ƥ (BIMH); "Palo, Leyte, 24. III. 57, WU7" 1 Ƥ (BIMH); "Palo, Leyte, P. I., 12. XI. 57, WU7" 1 Ƥ (BIMH); "Palo, Leyte, P. I., 2. XII. 57, WU7" 1 Ƥ (BIMH); "Palo, Leyte, P. I., 15. III. 58, WU7" 1 Ƥ (BIMH); "Burauen bei Leyte, leg. Böttcher" 1 3 1 Ƥ (MNHB). PANAON: "Panaon, 4. XII. 1915 " 2 3 (MNHB). MINDANAO: "Mindanao, P. I., coll. Staudinger" 1 Ƥ (SMTD); Mindanao, ... [illegible] 1 Ƥ (MNHP); "Mindanao, Zamboanga, coll. W. Schultze" 1 Ƥ (DEIM); "Davao, Mindanao, Dr. Platen" 1 Ƥ (MNHP); "Momungan b. Lanao, leg. Böttcher" 2 Ƥ (MNHB); Bugo, Misamis Or. B. M. 1937–107 1 Ƥ (NHML); Mt. Malindang, 800–1300 m, 8°16.07' N / 123° 35.76' E, 10.–13. I. 2007, Bolm leg. 1 3 1 Ƥ (CMB). PALAWAN: Port Barton, 150 m, 14.–18. XII. 1990, Bolm leg. 1 Ƥ (NHMB); "N. Palawan, Binaluan, Nov.-Dez. 1913, leg. G. Boettcher" 1 3 (MNHB). BALABAC: "Pasig, Balaba, Mar. 4. 1957, Yoshio Kondo Collector" 1 Ƥ (BIMH).

PAPUA NEW GUINEA: "Stephansort, Astrolabe Bai, Biró 1899" 1 3 (MTMB).

PALAU: "Palau-Inseln, Semper" 2 Ƥ (MNHB).

MICRONESIA: "W. Caroline Is., Fais I., Apr. 28, 1954, J. W. Beardsley Collector" 1 Ƥ (BIMH).

CAMEROON (?): "S. O. Kamerun, Lolodorf, L. Conradt 95, Coll. Kraatz" 1 Ƥ (DEIM).

UGANDA: " Uganda, Roscoe, Sharp Coll. 1905–313" 1 Ƥ (NHML).

TANZANIA: "D. O. Afrika, Vosseler G." 2 Ƥ (MNHB); Afr. or. all. 1907, coll. W. Wittmer" 2 Ƥ (NHMB); "Dt. O.-Afrika, coll. W. Wittmer" 1 Ƥ (NHMB); "D. O. Africa, Dr. Holtz, Sachsenwald, 13. 9. 03 " 2 Ƥ (MNHB); " Deutsch-Ostafrika, Coll. Kraatz" 1 Ƥ (DEIM); "D. O. Afrika, coll. Staudinger" 2 Ƥ (SMTD); "Kondoa, Bloyet 1885" 1 Ƥ (MNHP); "D. O. Afr., Unt. Umuba X. 15, leg. Methner" 1 Ƥ (MNHB); "Morogoro, "D. O.A., XII. 0 6, leg. Holz" 4 Ƥ (MNHB); "D. O. Africa, Morogoro, II. 14, leg. Methner" 1 Ƥ (MHHB); "D. O.-Afrika, Uluguru-Berge, XI.-XII. 98, Götze S." 1 3 1 Ƥ (MNHB); "Unguru, D. O. Afrika, P. Ringler, Halle, coll. Staudinger" 1 Ƥ (SMTD); "Usambara, Nguelo" 1 3 1 Ƥ (NHMB); "Usambara, Neu Bethel, V– VI–1903 " 1 Ƥ (MNHB); "Usambara Neu Bethel, 21. I. 1905 " 1 Ƥ (MNHB); "Usambara, Neu Bethel, 7. 3. 1905 " 1 Ƥ (MNHB); "Kwai, West-Us., D. O. A., Dez. 14, leg. Methner" 1 3 (MNHB); "D. Ost- Afrika, Usambara, Kwai, P. Weise S." 1 3 5 Ƥ (MNHB); "Usamb., Kwai, P. Weise S." 2 Ƥ (MNHB); "D. O. Africa, Usambara, P. Weise S." 1 Ƥ (MNHB); "Usambara, Mlusi Miss., 200 m, 15. 12. 91, Conradt S." 1 Ƥ (MNHB); "Kwamkussu, W. Usambara, 1640 m, II. 12 " 1 Ƥ (MNHB); "Usambara, Derema, Conradt S." 4 Ƥ (MNHB); " O. Usambara, F. Fischer leg." 1 Ƥ (MNHB); "W. Usambara, J. Buchwald S. " 2 Ƥ (MNHB); "D. O.-Afrika, W. Usambara, A. Borgert S." 1 Ƥ (MNHB); "Usambara, Neu Bethel, Jul. Moser" 2 Ƥ (NHMW); Usambara Mts. Lushoto, 6.–11. V. 1999, A. Kudrna jr. leg. 1 Ƥ (CMG); Usambara Mts., Irente near Lushoto, 1450 m, 22. II. 2008, Bellmann leg. 1 Ƥ (CMG); Usambara Mts., Irente near Lushoto, 1450 m, 22. II. 2008, Bellmann leg. 1 Ƥ (CAB); "Usambara, Coll. Kraatz" 1 3 2 Ƥ (DEIM); "Usambara, D. O.A., Coll. v. Bennigsen" 1 Ƥ (DEIM); "Usambara, Nguelo, Coll. Kraatz" 1 Ƥ (DEIM); "Afr. Or., Usambara, Fry Coll., 1905.100" 1 3 2 Ƥ (NHML); " Zanzibar, Usambara, Fry Coll., 1905–100" 1 3 (NHML); "Usambara, E. Africa, 1893 94, 95 239" 1 3 1 Ƥ (NHML); "Usambara, E. Africa, 4. 4. 94, 95 239" 1 Ƥ (NHML); "Usambara, Tanga, Reimer S." 1 Ƥ (ZSMU); "Amani par, Tonga, Afr. or., 1907, coll. W. Wittmer" 2 Ƥ (NHMB); "D. O. Afrika, Amani, X.-XII. 0 3, Vosseler S.G." 1 3 2 Ƥ (MNHB); "D. O. Afrika, Amani, 1.- 4. II. 0 6, Vosseler S.G." 1 3 2 Ƥ (MNHB); "D. O. Afrika, Amani, 1.–20. II. 0 6, Vosseler S.G." 1 Ƥ (MNHB); "D. O. Afr. Mombo, 8. 0 4, Vosseler G." 1 Ƥ (MNHB); "Daressalam" 2 Ƥ (MNHB); "Daressalam, D. O.Africa" 1 3 (ZSMU); "Daressal., D. O. Afr., coll. Staudinger" 1 Ƥ (SMTD); "Daressalam, D. O. Africa, Dr. Krauss, coll. Staudinger" 1 Ƥ (SMTD); "D. O. Afrika, Daressalam, Pangani u, Hinterld., Regner S. G." 2 Ƥ (MNHB); "Tanga" 1 Ƥ (MNHB); "Tanga, XII. 02 " 1 Ƥ (MNHB); "D. O. Afrika, Tanga, Vosseler S. G." 1 Ƥ (MNHB); "D. Ost-Afrika, Tanga, Umgegend, III. IV. 93, P. Neumann S." 1 Ƥ (MNHB); "D. Ost-Afrika, Korogwe, Mitte V. 93, O. Neumann S." 1 3 1 Ƥ (MNHB); "D. O. Afrika, Mikesse Safari, Tendaguru" 1 Ƥ (MNHB); "S. O. Victoria, Nyansa-See, G. A. Fischer" 1 Ƥ (MNHB); "Mittleres Rufiji, Schuster" 1 3 (MNHB); "Mkomasi- Steppe, V. 03 " 1 3 (MNHB); "Papyrus-Sumpf, süd-ö. Kilimandjaro, 20.–21. I. 0 6, D. O. Afr. coll. Dr. Chr. Schröder" 1 3 (MNHB); " Tanzania or., Kibiti, 9. 12. 1993, lgt. Snizek" 1 Ƥ (NHMB); "Pande. Tanga. 2. 8. 1952, Frl. Menzi" 1 Ƥ (NHMB); Lushoto, J. Plava leg. 1 Ƥ (NHMB); Utete–(Rufiji r.), 28. XII. 1976, J. Moravec leg. 1 3 (NHMB); "Bondei, Coll. v. Bennigsen" 1 Ƥ (DEIM); "Mpapua, Coll. v. Bennigsen" 1 Ƥ (DEIM); "Mva, Coll. v. Bennigsen" 1 Ƥ (DEIM); "Pangani Steppe, Coll. Bennigsen" 1 Ƥ (DEIM); "Ukami, D. O. Afrika, coll. Staudinger" 6 3 8 Ƥ (SMTD); "Morogoro, D. O. Afr., coll. Staudinger" 1 Ƥ (SMTD); "Tanganika, D. O. A., coll. Staudinger" 2 3 (SMTD); "Nguelo, D. O. Afr., coll. Staudinger" 1 3 1 Ƥ (SMTD); "Tanga, D. O. Afr., coll. Staudinger" 1 Ƥ (SMTD); "Tanga, coll. Staudinger" 1 Ƥ (SMTD); "G. E. Africa, Rd. to Kilossa, Usagara Dist., Dec. 22–26, 1910, 1500–2500 ft., S. A. Neave, 1911–177" 4 Ƥ (NHML); Ruvu, base camp, 6°47' S / 38°31' E, X. 1992, WRB Hynd leg., BMNH(E) 1998–129. 1 Ƥ (NHML); „ Zanzibar, Raffray, 299, coll. D. Ach. 1. 90“ 2 Ƥ (MHNG). "Höhnel 85, Ost Afrika, Höhnel Sansibar " 1 Ƥ (NHMW); " Zanzibar, Raffray" 2 Ƥ (MCGD); "A. Raffray, Zanzibar " 1 3 (MNHP).

KENYA: "Brit. O. Afr., Lindblom" 1 Ƥ (NRMS); "B.' Ost-Afrika" 1 Ƥ (MNHP); "Ngong, van Someren, 551, Pres. by Com. Inst. Ent., B. M. 1968–26" 1 3 (NHML); "Mazeras, Kenia, Br. E. Africa, El. 1500 ft., XI– 17–1921, R. Dummer, Frank R. Mason Collection, Ex coll. F. R. Mason, Brit. Mus. 1923–141" 1 3 (NHML); "Brit. E. Africa, Nairobi, 1919–103, E. A. Elliott Coll." 1 Ƥ (NHML); "van Someren, Rabai, August 1930, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 3 3 (NHML); "van Someren, Rabai 8–37, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 1 3 (NHML); "Jeffery, Kilifi, V. 44, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 1 Ƥ (NHML); "G. W. Jeffery, Kilifi, I. 45, Imp. Inst. Ent., coll.no. /0829, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 1 Ƥ (NHML); "G. W. Jeffery, Kilifi, I. 45, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 4 Ƥ (NHML); "Kilifi, 3. 45, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 3 Ƥ (NHML); "van Someren, Kilifi 6.46, Imp. Inst. Ent., coll.no. /0829, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 1 Ƥ (NHML); "van Someren, Kilifi (J.), Kenya, 7. 48, V. G. L. van Someren Collection. Brit. Mus. 1959-468." 4 Ƥ (NHML); "Dr. van Someren, Sokoke, 1 24, V. G. L. van Someren Collection, Brit. Mus. 1959-468." 1 Ƥ (NHML); "Africa or., Ikutha" 1 3 1 Ƥ (MNHB); "Br. O. Afrika, Taru– Mombasa, F. Thomas S." 1 Ƥ (MNHB); "Brit. O.-Afrika, Takaungu, Thomas S." 1 Ƥ (MNHB); Malindi, Gede Forest, V. 1990, Werner leg. 1 3 (NHMW).

UNCLEAR OR IMPRECISE LOCALITIES: „Indes orient., A. Naville, 601.94“ 1 Ƥ (MHNG); „[illegible], Ach. Geilenkeuser, 1893“ 1 Ƥ (MHNG); "Parr. p." 1 3 (NHMW); "Acc. No. 2412, Govt. Lab. Coll., Collected by W. Schultze, Coll. W. Schultze" 1 Ƥ (SMTD); "Indo-China" 2 3 2 Ƥ (MTMB); " India or." 1 Ƥ (MTMB); "Borneo, Xanthus" 1 Ƥ (MTMB); "Nord-Borneo, Pagel S." 2 Ƥ (MNHB); "Borneo, Dr. Will" 1 Ƥ (MNHB); "W. Borneo, Dr. Will" 1 3 (MNHB); "A. d. Sammlung Dr. Chr. Schröder's, Tareh ... (illegible), 1100 m " 1 3 (MNHB); "A. d. Sammlung Dr. Chr. Schröder's, Tareh ... (illegible), 1600 m " 1 3 1 Ƥ (MNHB); "A. d. Sammlung Dr. Chr. Schröder's, Tareh ... (illegible), 2000 m " 1 3 1 Ƥ (MNHB); "Haut Tonkin et Bas-Yunnan, entre Han-Hao, Muong Hun (près Lao-Kay) et Ban- Kam-Coun, Lieut. Lesourt 1905" 1 3 1 Ƥ (MNHP); "Cochinchine et Sumatra, Beauvais 1902" 1 Ƥ (MNHP); "Afrique au sud de l'Equateur?" 1 Ƥ (MNHP); "482 41" 1 Ƥ (MNHP); "2044 41" 1 Ƥ (MNHP); "847 85" 2 Ƥ (MNHP); "6965 85" 1 Ƥ (MNHP); "613 91" 1 Ƥ (MNHP); "614 91" 1 Ƥ (MNHP); " India Or." 1 Ƥ (NMPC); " Ind. Or., Helfer" 3 3 1 Ƥ (NMPC); "Neuguinea, coll. L. Salvator" 1 Ƥ (NMPC); "E. Ind., 43 43" 2 3 (NHML). "E. Indies., ex Standen., G. C. Champion, B. M. 1964–540" 1 Ƥ (NHML); "G. Lewis, 1915–38 " 1 Ƥ (NHML); "Balch, 62 30" 1 Ƥ (NHML); "Borneo, Bowring, 63-47*" 1 Ƥ (NHML); "Sharp Coll. 1905–313." 1 Ƥ (NHML); "Borneo, ... [illegible], German Mission, Fry Coll., 1905.100" 1 Ƥ (NHML); "Borneo, Fry Coll., 1905.100" 1 Ƥ (NHML); "N. Borneo, Exp. 1912, Mohari, 2662" 1 3 (DEIM); "Borneo, Pascoe Coll., 93–60." 1 Ƥ (NHML); "Massai, 87.32" 1 Ƥ (NHML); "Thurdy" 1 3 (NHML); "Thurdy, Andrewes Bequest, B. M. 1922–221" 1 3 3 Ƥ (NHML); "Thurdy, H. Swale, 1913–117" 1 3 (NHML); "Shuegyin, H. E. Andrewes Bequest, B. M. 1922–221" 1 3 (NHML); "Ost-Afrika, Fruhstorfer" 2 3 1 Ƥ (MTMB); "Africa" 1 Ƥ (MTMB); "Ostafrika, Coll. Kraatz" 1 3 (DEIM); "Ost-Afrika, Fruhstorfer, Coll. Kraatz" 2 Ƥ (DEIM); "Tansea lama, 17– 2–20 " 2 3 (DEIM).

WRONGLY LABELLED: "Austral." 1 Ƥ (MNHB); „ Tanger, Autran“ 1 Ƥ (MHNG); "Cafreria" 1 Ƥ (MNHP); "madag." 1 Ƥ (MNHP); " Mexico, Temescaltepec, H. E. Hinton, R. L. Usinger collectors, Hinton coll., B. M. 1939–583 1 3 (NHML); " Uruguay, 18.36" 1 3 (NHML).

NO LOCALITY: 1 Ƥ (NHML); 2 Ƥ (NHMW); „coll. Melly“ 2 Ƥ (MHNG); "149" 1 Ƥ (MNHP); "Patria?, coll. Staudinger" 1 Ƥ (SMTD).

Measurements 3 (n = 482): TBL 9.3–12.8 mm, L-h 7.7–10.9 mm

HL 1.5–1.9 mm, PL 1.4–2.0 mm, EL 6.3–8.5 mm

Measurements Ƥ (n = 1065): TBL 9.7–13.2 mm, L-h 8.3–11.1 mm

HL 1.4–2.1 mm, PL 1.5–2.0 mm, EL 6.8–9.1 mm

Differential diagnosis: Easily distinguished from P. b i c o l o r and P. viridiflavus by elytral coloration, from P. paiensis by entirely black femora and dark antennae, from P. wittmeri by stouter, broader habitus, from P. opacipennis , P. championi and P. opacipennis usually by more shining, bright blue coloration (but not always), and by the less rugose and slightly less dense elytral puncture. The shape of the male aedeagus (paramera and apex of median lobe) clearly distinguishes P. coeruleipennis from all other species.

Redescription: Habitus as in figs. 31 (3) and 32 (Ƥ). Body metallic blue, sometimes greenish blue or metallic dark green. Pronotum bright reddish orange. Antennae almost black, with the terminal segment (sometimes also part of segments 9–10) brownish orange, the three basal segments often partly reddish. Coloration of maxillary palpi to a variable extent reddish, sometimes almost entirely reddish orange, sometimes more infuscate to almost entirely black. Tarsi black or bluish black, claws orange brown.

Head behind the eyes about half as wide as middle part of pronotum. Vertex shining, very sparsely and very finely punctate and with some small black hairs. Frons between eyes shining, sparsely punctate, narrower than the length of the first antennal joint in males, about as broad in females; in front of the eyes distinctly depressed and slightly rugose. Clypeus trapezoidal, about half as long as wide at the base and slightly convex, with some very fine punctures. Labrum about as long as wide, rather flat and more coarsely punctate than the head, with some long blackish hairs.

Male antennae reaching the first quarter of the elytra in length, until slightly after the humeral callus. First three segments subfiliform, segments 4–5 widened and flattened, 6–10 of subtriangular shape, very strongly widened. Last segment not conspicuously widened, but large and strongly emarginate. First segment long, second very short, only about half as long, third long, almost like the first, the next three of about the same length, then slightly decreasing in length, but increasing in width, the last segment longest, almost 1.5 times as long as the first. Female antennae slightly shorter and smaller, only scarcely reaching the humeral callus; subapical segments a bit shorter and stouter, but otherwise everything of similar shape than in males.

Pronotum about as long as wide or slightly wider, maximal length: maximal width 1: 1–1: 1.1; subhexagonal (widest in the middle), base almost straight, all angles bluntly rounded; with two oblique impressions in the basal half, at both sides of the disc, finely bordered, the bordering becoming very vague in the middle of the apical and front margins, often completely reduced in the middle of the front margin; shining, without any microsculpture; regularly, but very sparsely covered with some inconspicuous, suberect black hairs and with some long black setae along the apical half of the outer margin.

Elytra about 2.4–2.5 times as long as wide in males, about 2.1–2.3 times in females, widest slightly behind the middle; humeral area convex, flattened towards the apex; finely, densely and regularly punctate over their whole surface, punctures slightly shallower towards the apex; regularly covered with fine, brownish, short recumbent hairs, and bearing some few, long, erect, pale hairs, partly arranged into rows; usually with two or three indistinct, hardly visible granular rows. Margins distinctly crenulate, strongly in humeral area, getting slightly weaker towards the apex, the crenules all bearing blackish setae. Scutellum about twice as wide as long, all angles bluntly rounded; punctures fissure-like, very fine (much finer than on the elytra), covered with fine regular pubescence like the elytra, but without long erect hairs.

Femora metallic, shining, with fine punctures and rather long, greyish pubescence. Tibiae not metallic, densely covered with brownish setae. Tarsi also not metallic, with blackish or brownish setae.

Abdomen with rather long, but not very dense, greyish, recumbent pubescence and some longer black, suberect setae; with rather sparse, shallow punctures and somewhat rugose texture.

Male: Last abdominal sternite small, much narrower than the other segments, subtriangular in shape, sides strongly converging towards the rounded apex (fig. 22). Basal margin rather shallowly emarginate. Apical margin deeply incised, forming an approximately oval opening hole for the aedeagus, together with the slightly emarginate last tergite.

Aedeagus (figs. 1, 8, 15): Process of phallobase pointing backwards. Paramera flat, almost straight, slightly arcuate, their tip with a small dorsal tooth and some very small teeth or granulae along the ventral edge; apical part with long, brownish hairs. Seen in dorsal view, they are broad, straight and almost parallel, with only a narrow gap between them. Median lobe with base curved ventrally, central part almost straight and slightly narrowed towards the tip. Apex (in lateral view) dorsally with a strong tooth-like angle, then flat until tip; ventrally more or less continuously narrowed, rounded before the tip, not as strongly flattened as in P. b i c o l o r. Median lobe in dorsal view with elongate, oval ostium; its tip subtriangular.

Sexual dimorphism: Females have broader elytra and are usually slightly larger than males. The sexual dimorphism in the width of the frons between the eyes is not as well pronounced as in other Prionocerus , but still visible. The female antennae are usually slightly shorter and smaller, antennal joints 5–10 are slightly more dilated in males than they are in females.

Variability: The scutellum is usually metallic blue with a large orange yellow patch on the disc, this patch can be well pronounced, indistinct or even completely missing in some individuals. The elytral coloration is usually metallic blue, but can be green in some specimens. One specimen (" var. diversicollis ") has the pronotal disc infuscate, brownish (see Notes).

Distribution (fig. 43): This species is very widely distributed in large parts of the Oriental region, including South China, Wallacea and New Guinea, as well as in East Africa (see discussion).

Notes: The synonymy of P. fuscipennis Lewis with P. coeruleipennis Perty , already proposed by Champion, 1919, could be confirmed. The holotype is a large, teneral female with a somewhat brownish, coppery elytral colour, probably due to its immaturity or due to inappropriate storage after collection. There are other individuals of this kind from Borneo and Singapore at NHML, from Indonesia (Nias) at MCGD, and from Vietnam and Peninsular Malaysia at MNHB. None of these shows any significant differences from typical P. coeruleipennis .

The variety diversicollis, described by Pic (1920) was based on a strongly damaged female from Sumatra, which could be examined. From the characters still visible, it seems to be an aberrative specimen of P. coeruleipennis with greenish elytra and slightly infuscate pronotal disc. There is another specimen under " var. diversicollis " in the Pic collection, a male from Malaysia definitely belonging to P. malaysiacus . As it was not labelled as type, not mentioned in the original publication and collected at the same time as the publication came out, it cannot be regarded as a syntype of the variety diversicollis.

Although the differences between male and female antennae are not always as clear as stated by Bourgeois, his synonymy proposals of the species described by Schaufuss are certainly correct. A male syntype of P. f o r t i c o r n i s could be examined and showed no clear differences at all to typical males of P. coeruleipennis . The female syntype of P. brevicornis shows slightly smaller (shorter and a bit less broadly serrate) antennae as usual in P. coeruleipennis , but these differences can surely not be regarded as specific. Of both Schaufuss species, there was only one type found in the collection of MNHB, but as Schaufuss was known to exchange a lot of type material with other collectors, it is unknown how large the original type series was. Both type specimens have therefore to be regarded as syntypes, but a lectotype designation is not urgently needed at the moment.

NHML

Natural History Museum, Tripoli

MHNG

Museum d'Histoire Naturelle

MNHP

Princeton University

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

SuperFamily

Cleroidea

Family

Prionoceridae

Genus

Prionocerus

Loc

Prionocerus coeruleipennis Perty, 1831

Geiser, Michael 2010
2010
Loc

Prionocerus fuscipennis

Pic 1920: 12
Champion 1919: 328
Bourgeois 1890: 175
Bourgeois 1890: 175
Lewis 1879: 464
1879
Loc

Prionocerus coeruleipennis

Perty 1831: 33
1831
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