Diamesa steinboecki Goetghebuer, 1933

Makarchenko, Eugenyi A., Semenchenko, Alexander A. & Palatov, Dmitry M., 2022, Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes, Zootaxa 5125 (5), pp. 483-512 : 494

Diamesa steinboecki Goetghebuer

( Figs. 26–29 , 34–36 , 73 , 77 )

Diamesa steinboecki Goetghebuer, 1933: 73, 1939: 16 ; Pagast 1947: 487; Wülker 1959, figs. 2–4; Pankratova 1970: 83; Serra-Tosio 1971: 182, 1974: 168; Schmid 1993: 41; Langton & Visser 2003: 47; Ashe & O’Connor 2009: 285; Rossaro & Lencioni 2015: 65; Montagna et al. 2016: 329.

Material examined. 1 adult male, TAJIKISTAN: Murgab District, Gorno-Badakhshan Autonomous Region, Pamir Mountains , Sasykteke River , ( Sauksay River basin), altitude 3815 m above sea level, 25.VIII.1987, N 39.288347, E 72.370182, leg. D. Shcherbakov ; 2 adult males, 2 mature pupae (male and female), SWITZERLAND: Valais Canton, Swiss Alps, Muttbach Stream , 16.IX.2018, altitude 2500 m above sea level, leg. B. Lods-Crozet ; 1 adult male, FRANCE: cold spring, French Alps, Massif of Pelvoux, Lautaret Pass , 9.IX.1966, leg. B. Serra-Tosio ; 1 mature pupa (male), 2 mature pupae (females), 2 larvae, French Alps, Massif of Aiguilles-Rouges, Balm Stream , 8.IX.2021, altitude 1862 m above sea level, N 45.97933, E 6.840494, leg. P. Clévenot.

Remarks. Despite the rather widespread of D. steinboecki in the mountains of Europe, a detailed description of the adult male, the variability of brachypteric and macropteric forms at different altitude above sea level, was made only for the French Alps (Serra-Tosio 1971, 1973, 1974). Therefore, we found it expedient to give a brief re-description of males based on our material from the Pamir Mountains, where the species was found for the first time, and the Swiss Alps, as well as to add some data to the description of Serra-Tosio (1971, 1974) from the French Alps. Data of Serra-Tosio (1974) in our description are highlighted in bold.

Figures

FIGURES 26–29. Males of Diamesa steinboecki Goetghebuer from Pamir Mountains (26), Swiss Alps (27–28) and French Alps (29). 26, gonocoxite and gonostylus; 27, 29, hypopygium in dorsal view; 28, transverse sternapodeme, phallapodeme and aedeagal lobe. Designations are the same as in Figures 1–8.

FIGURES 30–36. Wings of Diamesa dragani sp. nov. (30), D. kownackii sp. nov. (31), D. maisaraensis sp. nov. (32), D. marinskiyi sp. nov. (33) and D. steinboecki Goetghebuer from Swiss Alps (34), French Alps (35) and Pamir Mountains (36).

FIGURES 68–74. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (69–70), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205–210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of Aiguilles–Rouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov).

FIGURES 75–78. Type localities of Diamesa moubayedi sp. nov. (75), Diamesa zagrosica sp. nov. (76), D. steinboecki Goetghebuer (77) and D. praecipua Saether et Willassen (78). 75, glacial karstic spring, Sannyn Mount, alt. 2000 m a. sea l., Central Lebanon (photo by J. Moubayed); 76, Dare Daei River, Zagros Mountains, alt. 2244 m a. sea l., Lorestan Province, Iran (photo by D.M. Palatov); 77, Muttbach Stream, Valais Canton, Swiss Alps (photo by Jean–Louis Lods); 78, Rishi Ganga River, right tributary of the Alaknanda River near town of Badrinath, alt. 3128 m a. s. l., Himalayas, India (photo by D.M. Palatov).