Belisana liupanshui Wang, S. Li & Yao, 2024

Wang, Bing, Li, Jinglin, Li, Shuqiang & Yao, Zhiyuan, 2024, Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China, ZooKeys 1216, pp. 265-302 : 265-302

publication ID	https://doi.org/ 10.3897/zookeys.1216.132561
publication LSID	lsid:zoobank.org:pub:0E173F72-36D3-4B97-BC43-F096067A3D1E
DOI	https://doi.org/10.5281/zenodo.13992754
persistent identifier	https://treatment.plazi.org/id/4C5D21CD-86E7-43FA-B974-0235DB601B2B
taxon LSID	lsid:zoobank.org:act:4C5D21CD-86E7-43FA-B974-0235DB601B2B
treatment provided by	ZooKeys by Pensoft
scientific name	Belisana liupanshui Wang, S. Li & Yao
status	sp. nov.

Treatment

Belisana liupanshui Wang, S. Li & Yao sp. nov.

Figs 4 View Figure 4 , 5 View Figure 5 , 22 C, D View Figure 22 , 24 C, D View Figure 24

Type material.

Holotype: China • ♂; Guizhou, Liupanshui, Pan County, Biyun Cave ; 25 ° 46.527 ' N, 104 ° 38.278 ' E; alt. 1468 m; 13 Apr. 2007; J. Liu & Y. Lin leg.; IZCAS - Ar 45181 GoogleMaps . Paratypes: China • 5 ♂; same data as for holotype; IZCAS - Ar 45182 –86 GoogleMaps • 7 ♀; same data as for holotype; IZCAS - Ar 45187 –93 GoogleMaps .

Diagnosis.

The new species resembles B. jiuxiang Zhang, Li & Yao, 2024 ( Zhang et al. 2024 b: 261, figs 4 A – D, 5 A – H, 18 C, D, 20 C, D) by having similar male chelicerae (distal apophyses directed towards frontally, but tips pointing inwards; Fig. 5 D View Figure 5 ) and epigyne (epigynal pockets on median part of epigynal plate, epigynal plate posteriorly straight; Figs 5 A View Figure 5 , 24 C View Figure 24 ), but can be distinguished by procursus with prolatero-subdistal and subdistal membranous processes (arrows 1, 3 in Figs 4 C View Figure 4 , 22 C View Figure 22 vs absent) and nearly rectangle dorso-subdistal membranous process (arrow 4 in Figs 4 C View Figure 4 , 22 C View Figure 22 vs angular), by bulbal apophysis with angular subdistal apophysis (arrow in Fig. 5 C View Figure 5 vs absent), and by vulval pore plates anteriorly narrow and posteriorly wide (pp in Figs 5 B View Figure 5 , 24 D View Figure 24 vs nearly quadrilateral).

Description.

Male (holotype): Total length 1.71 (1.79 with clypeus), prosoma 0.63 long, 0.67 wide, opisthosoma 1.08 long, 0.81 wide. Leg I: – (4.04, 0.29, 4.50, 5.96, –), leg II: 11.08 (2.80, 0.30, 2.97, 3.92, 1.09), leg III: 7.98 (2.22, 0.25, 1.98, 2.72, 0.81), leg IV: 10.20 (3.00, 0.26, 2.63, 3.45, 0.86); tibia I L / d: 63. Eye interdistances and diameters: PME – PME 0.09, PME 0.08, PME – ALE 0.02. Sternum width / length: 0.51 / 0.44. Habitus as in Fig. 5 E, F View Figure 5 . Dorsal shield of prosoma yellowish, with brownish radiating marks; clypeus brownish; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (pa in Fig. 5 D View Figure 5 ) and pair of curved distal apophyses (distance between tips: 0.17; da in Fig. 5 D View Figure 5 ). Palp as in Fig. 4 A, B View Figure 4 ; trochanter with ventral apophysis (as long as wide, arrow 1 in Fig. 4 B View Figure 4 ); femur with retrolatero-proximal protrusion (arrow 2 in Fig. 4 B View Figure 4 ); procursus simple proximally but complex distally, with prolatero-subdistal membranous process (arrow 1 in Figs 4 C View Figure 4 , 22 C View Figure 22 ), sclerotized distal apophysis (arrow 2 in Figs 4 C View Figure 4 , 22 C View Figure 22 ), subdistal membranous process (arrow 3 in Figs 4 C View Figure 4 , 22 C View Figure 22 ), dorso-subdistal membranous process (arrow 4 in Figs 4 C View Figure 4 , 22 C View Figure 22 ), and retrolateral membranous flap (f in Figs 4 D View Figure 4 , 22 D View Figure 22 ); bulb with hooked apophysis bearing angular subdistal apophysis (arrow in Fig. 5 C View Figure 5 ) and simple embolus (e in Fig. 5 C View Figure 5 ). Retrolateral trichobothria on tibia I at 4 % proximally; legs with short vertical setae on metatarsi.

Female (paratype, IZCAS - Ar 45187 ): Similar to male, habitus as in Fig. 5 G, H View Figure 5 . Total length 2.53 (2.65 with clypeus), prosoma 0.67 long, 0.75 wide, opisthosoma 1.86 long, 1.80 wide; tibia I: 4.05; tibia I L / d: 51. Eye interdistances and diameters: PME – PME 0.11, PME 0.07, PME – ALE 0.02. Sternum width / length: 0.52 / 0.46. Epigyne simple and flat, posteriorly straight, with pair of median pockets 0.20 apart (ep in Figs 5 A View Figure 5 , 24 C View Figure 24 ). Vulva with ridge-shaped anterior arch bearing pair of angular lateral sclerites (arrow in Figs 5 B View Figure 5 , 24 D View Figure 24 ), and pair of anteriorly narrow and posteriorly wide pore plates (3 × longer than wide, pp in Figs 5 B View Figure 5 , 24 D View Figure 24 ).

Variation.

Tibia I in five male paratypes ( IZCAS - Ar 45182 –86): 4.50, 4.55, 4.62, 4.65, 4.70. Tibia I in the other six female paratypes ( IZCAS - Ar 45188 –93): 3.56–3.85.

Habitat.

The species was found in the dark zone inside the cave.

Distribution.

China (Guizhou, type locality; Fig. 1 View Figure 1 ).

Etymology.

The specific name refers to the type locality; noun in apposition.