Crassabwa Lugo-Ortiz & McCafferty 1996b

Kluge, Nikita J., Gattolliat, Jean-Luc, Salles, Frederico F. & Novikova, Eugenia A., 2018, Revision of the Afrotropical genus Susua Lugo-Ortiz & McCafferty 1998 (Ephemeroptera: Baetidae: Protopatellata), Zootaxa 4434 (1), pp. 1-28 : 26

Genus Crassabwa Lugo-Ortiz & McCafferty 1996b

Hierarchical name: Crassabwa /g2 (sine Susua ).

Type species Centroptilum flavum Crass 1947 .

Diagnosis. Among characters of Crassabwa reported in the previous paper ( Kluge et al. 2017), autapomorphies of this taxon are: (1) long costal projection of hind wing [ Kluge et al. 2017: Crassabwa (21)]; (2) peculiar structure and pigmentation of tergalii [ Kluge et al. 2017: Crassabwa (25), (26)]; (3) peculiar form of larval last abdominal tergum [ Kluge et al. 2017: Crassabwa (28)]; (4) peculiarity in form of unistyligers of male imago [ Kluge et al. 2017: Crassabwa (31)].

Other characters separating Crassabwa from Susua , are the following: larval coloration [ Kluge et al. 2017: Crassabwa (1)]; thickened 2nd segment of maxillary palp [ Kluge et al. 2017: Crassabwa (8)]; form of postsubalar sclerite [ Kluge et al. 2017: Crassabwa (11)]; shape of fore wing [ Kluge et al. 2017: Crassabwa (20)]; large denticles on larval abdomen [ Kluge et al. 2017: Crassabwa (22)]; retention of ability to make rhythmical movements by tergalii of all I–VII pairs [ Kluge et al. 2017: Crassabwa (27)]; large denticles on larval caudalii [ Kluge et al. 2017: Crassabwa (29)].

Species composition. Three species are attributed to Crassabwa for certain: C. flava (Crass 1947) , C. ludmilae Kluge et al. 2017 and C. ameliae Kluge et al. 2017 ( Kluge et al. 2017).

Species-specific characters. In a previous paper ( Kluge et al. 2017), we reported that besides other characters, larva of Crassabwa flava differs from C. ludmilae and C. ameliae by the presence of long, pointed scales on abdominal terga ( Kluge et al. 2017: Figs 20–21 ). It should be added, that in C. flava such long and pointed scales are present, besides certain areas of abdominal terga, also on certain areas of head, pronotum and mesonotum ( Figs 111–112 ); on pronotum and mesonotum these scales occupy mainly pigmented areas between sigilla. In C. ludmilae and C. ameliae such long and pointed scales are absent, and the same areas of pronotum and mesonotum are covered by short, oval scales ( Figs 113–114 ). In all three species of Crassabwa fore protoptera, as well as thoracic pleura and sterna bear only short, blunt scales (as in Figs 33 , 90 ).

Figures

FIGURES 17–27. Larva of Susua sigiensis. 17, base of tibia of fore leg, anterior view (setae on posterior side shown by interrupted lines); 18, apex of femur and base of tibia of middle leg, anterior view (setae on posterior side shown by interrupted lines); 19, claw; 20–26, tergalii I–VII; 27, paraproct.

FIGURES 111–114. Exuviae of larval pronotum of Crassabwa and its scales. 111–112, C. flava; 113–114, C. ludmilae.

FIGURES 28–33. Exuviae of left half of pronotum and mesonotum of last instar larva of Susua sigiensis. 28, general view; 29–31, bunches of spatulate setae on places shown by arrows; 32, scales on pronotum; 33, scales on posterior part of fore protopteron.

FIGURES 87–90. Exuviae of left half of pronotum and mesonotum of last instar larva of Susua niandanensis from Zambia. 87, general view; 88, scales on pronotum; 89, scales on mesonotum; 90, scales on posterior part of fore protopteron.