Clubiona filifera Dankittipakul

Dankittipakul, Pakawin & Singtripop, Tippawan, 2008, Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand, Zootaxa 1747, pp. 34-60 : 41-42

Clubiona filifera Dankittipakul sp. n.

( Figs 20–21 , 48–51 )

Type material. Holotype: ɗ, Khao Yai National Park, Khao Yai [Nakhorn Ratchasima Province, Northeastern Thailand], forests surrounding park headquarters, 800–1000 m, 15.ix.2006, P. Dankittipakul ( MHNG).

Paratypes: 1 ɗ, same data as holotype ( TNHM).

Etymology. The specific epithet filifera means bearing thread, which is derived from Latin fero (= to bear, carry) and filum (= thread); it refers to the very long, apically coiled filiform embolus on the male palp.

Diagnosis. Clubiona filifera sp. n. possesses certain characters associated with the japonica -group and resembles the species described below in having a simple, unbranched retrolateral tibial apophysis ( Fig. 51 ) and by the sinuate sperm duct on the tegulum forming a double loop ( Figs 49–50 ). This new species is also recognized by the absence of conductor, the filiform, apically coiled embolus and by the spiniform tegular apophysis ( Fig. 51 ). Clubiona filifera sp. n. is described as differing from the typical form of the Southeast Asian japonica -group in having different opisthosomal color patterns ( Fig. 48 ) and being larger in size. Clubiona filifera sp. n. is very similar to C. cirrosa Ono, 1989 occurring in the Ryukyus, Japan in having strongly coiled embolus situated apically but it is distinguishable from the latter species by the shape of tegular apophysis.

Description. Male (holotype): Total length 6.2; prosoma 3.0 long, 2.0 wide; opisthosoma 3.2 long, 1.7 wide.

Prosoma ( Fig. 48 ). Long-oval, narrowed behind ocular area, in profile strongly raised, highest just in front fovea; integument smooth, covered with numerous fine hairs. Dorsal shield of prosoma yellow, pars cephalica darker in ocular area, without color markings. Chelicerae protruding, dark brown. Labium and endites yellowish brown. Sternum yellow.

Eyes. Eyes with black rings. Eye sizes and interdistances: AME 0.16, ALE 0.11, PME 0.13, PLE 0.13, AME–AME 0.08, AME–ALE 0.03, PME–PME 0.30, PME–PLE 0.16; MOQ: 0.25 long, 0.43 anterior width, 0.46 posterior width.

Legs. Legs yellow; anterior tibiae, metatarsi and tarsi slightly darker; without conspicuous dark annulations. Leg formula 4213.

Leg measurements:

I II III IV

Femora 2.00 2.45 1.90 2.82

Patellae + tibiae 2.93 3.20 2.48 3.70

Metatarsi 1.50 1.80 1.43 2.95

Tarsi 0.70 0.90 0.60 0.80

Total 7.13 8.35 6.41 10.27

Opisthosoma ( Fig. 48 ). Elongate-oval, furnished with a thick tuft of hairs anteriorly; dorsum with cardiac region pale brown, starting at the front and running along middle for a third of the length; posteriorly dark reddish brown coloring is formed into chevron-like bands; venter pale. Spinnerets white.

Male palp ( Figs 20–21 , 49–51 ). Retrolateral tibial apophysis small, digitiform, apically hooked in ventral view. Bulb with distinctly broad membranous prolateral portion, occupying most of the apical part; tegulum with sinuate sperm duct forming a double loop. Long and filiform embolus distally coiled, resting on apical portion of the tegulum; embolic base originating on baso-prolaterally of the bulb. Tegular apophysis spiniform, situated on apico-retrolateral side near the embolic tip.

Female: unknown.

Natural history. The holotype was obtained from mixed deciduous forests behind a park headquarters. Distribution. Known only from the type locality, Khao Yai National Park, northeastern Thailand ( Fig. 69 ).

Figures

FIGURE 69. Records of Clubiona species treated in the present study: * C. filicata; C. suthepica sp. n.; C. melanosticta; C. octoginta sp. n.; C. campylacantha sp. n.; Ο C. filifera sp. n.; C. abnormis sp. n.

FIGURES 20 – 23. Clubiona filifera sp. n., male holotype (20, 21). 20 male palp, dorsal; 21 ditto, retrolateral. Clubiona suthepica sp. n., male holotype (22, 23). 22 male palp, dorsal; 23 ditto, retrolateral. Scale bars = 0.1 mm (20 – 23).

FIGURES 48 – 51. Clubiona filifera sp. n., male holotype. 48 habitus, dorsal; 49 male palp, ventral; 50 ditto, proventral; 51 ditto, retrolateral. Scale bars = 1.0 mm (48); 0.1 mm (49 – 51).