Phanaeus (Notiophanaeus) haroldi Kirsch, 1871

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 211-213

Phanaeus (Notiophanaeus) haroldi Kirsch, 1871 Plate 47A

Phanaeus haroldi Kirsch, 1871: 342 (original description. Type locality: Bogotá).

Phanaeus haroldi : Nevinson 1892: 4 (catalog, distribution, written as Phanaeus Haroldi , Kirsch); Gillet 1911a: 83 (catalog, distribution, written as Phanaeus Haroldi , Kirsch); d’ Olsoufieff 1924: 34 (characters in key); 85 (redescription), 147 (distribution); Blackwelder 1944: 209 (list of species from Latin America); Vulcano and Pereira 1967: 575 (characters in key); Vitolo 2000: 597 (characters in key); Medina et al. 2001: 140 (cited for Colombia); Noriega et al. 2009: 406 (distribution); Krajcik 2012: 204 (complete list of species); Ratcliffe et al. 2015: 197 (cited for Peru).

Phanaeus (Notiophanaeus) haroldi : Edmonds 1994: 21 (characters in key), 24 (redescription), 101 (distribution); Vaz-de-Mello 2000: 194 (cited for Brazil); Arnaud 2002a: 82 (diagnosis); Vitolo 2004: 283 (diagnosis); Carvajal et al. 2011: 320-321 (cited for Ecuador); Edmonds and Zídek 2012: 12 (characters in key); Figueroa et al. 2014: 133 (distributional records from Peru); Chamorro et al. 2018: 98 (cited for Ecuador).

Phanaeus schneblei Frey, 1963: 558 (original description); Arnaud 2002a: 82 (cited as synonym of Phanaeus haroldi Kirsch, 1871).

Type specimens.

Phanaeus haroldi Kirsch, 1871. The holotype (♀) is deposited at the SMTD (see Edmonds 1994: 24). Locality: Bogota, not examined.

Phanaeus schneblei Frey, 1963. Type material not examined.

Distribution.

Colombia, Ecuador, Peru, and Venezuela.

Records examined.

MORONA SANTIAGO: Huambi, 900 m (2 specimens CEMT); Logroño (1 specimen MQCAZ); Taisha (1 specimen MQCAZ); km 8 road to Mendez-Paute (1 specimen MQCAZ). NAPO: 4 km de San Pedro de Arajuno, 370 m (1 specimen MQCAZ); Avila Viejo Ek olm, 750 m (1 specimen MQCAZ); Cotundo Comunidad Rumiñahui, 1070 m, Kuriurcu (1 specimen MUTPL); Pano (1 specimen MQCAZ); Puerto Napo, 480 m (2 specimens MQCAZ; 1 specimen MECN); Talag Marungachi, 750 m (1 specimen MQCAZ). ORELLANA: Coca (1 specimen MQCAZ); Dayuma Campo Palanda, Pindo 14, 255 m (3 specimens MUTPL); El Coca plataforma Oso B, 250 m (1 specimen MECN); Estación Científica Yasuní PUCE, 250 m (8 specimen MQCAZ); Rodrigo Borja IAMOE (6 specimens, CEMT; 2 specimens MQCAZ; 1 specimen MECN); Ines Arango road Tiwino-río Shiripuno, 250 m (1 specimen MQCAZ); Taracoa (1 specimen MQCAZ). PASTAZA: Bosque Protector Oglán Alto, 555 m (1 specimen MUTPL); Cononaco (1 specimen MQCAZ). SUCUMBÍOS: 6 km de Dureno, Precooperativa Los Vergeles, 306 m (1 specimen MGO-UC); Campo Bermejo, 600 m (1 specimen MECN); Shusufindi Recinto la Pantera, 250 m (1 specimen CEMT). ZAMORA CHICNHIPE: km 1 road Cumbaritza-Gualaquiza (1 specimen MQCAZ); El Pangui, Eneretza, 925 m (1 specimen MQCAZ); Tunantza, Timbara (1 specimen MQCAZ).

Literature records.

LOJA: without specific locality (Noriega et al. 2009: 406). NAPO: Aliñahui Amazon Basin (Noriega et al. 2009: 406); Archidona (Noriega et al. 2009: 406); Pto Misahualli (Noriega et al. 2009: 406); 29 km E of San Pedro de Arajuno, 362 m (Noriega et al. 2009: 406); Tena, 400 m ( Edmonds 1994: 101; Noriega et al. 2009: 406); Tena, 650 m (Noriega et al. 2009: 406). ORELLANA: Payamino Research Station, 400 m (Noriega et al. 2009: 407); Loreto ( Edmonds 1994: 101; Noriega et al. 2009: 407). PASTAZA: Puyo ( Edmonds 1994: 101; Noriega et al. 2009: 407); Canelos ( Edmonds 1994: 101; Edmonds 1994: 101; Noriega et al. 2009: 407). PICHINCHA: Quito (Noriega et al. 2009: 407). SUCUMBÍOS: Lago Agrio, 250 m ( Edmonds 1994: 101; Noriega et al. 2009: 407).

Temporal data.

Collected in February, April, May, June, July, August, September, October, November, and December.

Remarks.

Inhabits the lowland evergreen forests and evergreen foothill forests of the Amazon region from 255-1070 m a.s.l. Collected with pitfall traps baited with human feces. The record for Quito is probably erroneous.