Anabarhynchus darembal Ferguson

Ferguson, David J., Lambkin, Christine L. & Yeates, David K., 2014, Eight new species of Australian stiletto flies in the genus Anabarhynchus Macquart (Diptera: Therevidae) from South East Queensland, Zootaxa 3802 (4), pp. 553-582 : 558-560

Anabarhynchus darembal Ferguson sp. n.

( Figs 4 , 5 , 6 )

Type material. Holotype: Male. AUSTRALIA: Queensland: Fairy Bower, Rockhampton. Queensland, 14 September 1954; I.F.B. Common ( ANIC _29:029635) ( ANIC). Condition: Micro pinned dorsally to pith block; right hind tibia missing.

Diagnosis. Frons flat with reflective brownish grey pubescence; frontal pile black, distributed in two rows; male frons width 2.2 × ocellus; occipital macrosetae black; middle coxa posterior surface with several pale hairs; fore femur 1 pd; middle femur 0–1 pd, 2 pv; hind femur 1–2 av, 1av macroseta; all femora and tibia pale brown.

Description. Male. Body length: 8 mm. Wing length: 8 mm. ( Figs 4–6 ). Head. Frons flat, width 2.2 × ocellus width; ocellar tubercle flat; face and lower frons slightly protruding; parafacial to lower frons reflective grey; frons yellowish grey viewed anterodorsally, brown anteriorly; middle and upper frontal pile short and semi erect, in two rows, lower frons pile long and erect 3/4 length of scape. Scape length 2.8 × pedicel length; scape and pedicel grey; f1 brown with a short black seta dorsally on basal quarter; f2 & f3 brown, combined length a third length of f1. Occiput convex, brownish grey pubescence, several indistinct rows of black macrosetae, 36 on each side; postocciput to gena grey pubescence with long, pale, hair-like pile. Palp white, with whitish hair-like setae; labellum grey; prementum setae black. Thorax. Scutal chaetotaxy black (pairs): np, 4; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with thin brown dorsomedial line within broader grey vittae, margined by yellowish grey vittae.

Katepisternum and prosternal furrow without pile; pleuron grey; coxae grey with long pile admixed with black macrosetae; coxa-2 with a few pale hairs on posterior surface. Wing. Hyaline with pale grey infuscation, veins brown, stigma pale brown; costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel brown, knob grey brown. Legs. Fore femur with 1 pd; middle femur 0–1 pd, 2 pv; hind femur 1–2 av macrosetae; appressed pale pile on dorsal surfaces admixed with short black setae, ventral pile erect. All femora pale brown, with grey pubescence; all tibiae and tarsi pale brown, apically darker. Abdomen. Integument pale brown, dark brown dorsomedially; tergites 2–5 with anterior bands dark brown, reducing in breadth towards posterior, covered with appressed dark pile, posterolateral edges bright grey with pale pile; tergite 2–6 apical bands bright white; all sternites pale brown with thick cover of grey pubescence, apical bands grey white. Terminalia: Epandrium ( Fig. 6 A): twice as wide as long (measured along mid-line and between widest points when viewed dorsally), narrowing posteriorly. Sub-epandrial plate sclerotised brown laterally. Gonocoxite ( Fig. 6 B): weakly sclerotised; slightly longer than wide when viewed ventrally; several strong dark macrosetae admixed with shorter pale setae posterolaterally; small lobe-like outer gonocoxal process towards the ventromedial plane that supports dense pale pile. Gonocoxal apodeme short. Inner gonocoxal process darkly sclerotised, directed downwards and inwards, forming a broad and thin spatula-shaped apical section, pale pile on dorsal surface. Gonostylus long, darkly sclerotised and with angular projections dorsally and laterally; pale pile ventrally and along dorsal edge, directed medioventrally. Ventral lobe narrow and triangular when viewed dorsally, anterior edge tapered with sub-apical break, and connected to the gonocoxal apodeme by thin membrane. Aedeagus ( Fig. 6 C, D): distiphallus long and ventrally directed; parameral sheath darkly sclerotised, broadly triangular; the anterolateral edge raised dorsally; ventral apodeme anteriorly broad and flat with the apical margin slightly bilobed; lateral ejaculatory apodeme narrow, band-like; ejaculatory apodeme thinly sclerotised, relatively short, broad and thin, apically rounded.

Female. Unknown.

Etymology. The specific epithet is derived from the name of the original inhabitants of the land where the holotype specimen was collected, whose name is the ‘Darembal’ ( Tindale, 1974).

Comments. Known from one male collected in September from the Rockhampton area, Queensland.

Anabarhynchus darembal sp. n. keys to couplet 64 in Lyneborg (2001) and is readily separated from both A. apertus (Macquart) and A. kosciuskoensis Mann as both these species have yellow-brown inner gonocoxal processes with robust setae on the apex. In A. darembal sp. n. the inner gonocoxal process is darkly sclerotised, with an apex that is spatulate with a few, sparsely distributed, weak, pale setae.

Anabarhynchus darembal sp. n. closely resembles both A. fasciatus Macquart and A. tener Lyneborg. Anabarhynchus fasciatus has dark grey femora, middle femur without pd macrosetae and lacks pale pile on the posterior surface of the middle coxa. Anabarhynchus darembal sp. n. has pale brown femora, middle femur with 1 pd, and pile on the posterior surface of the middle coxa. While A. tener has pale occipital macrosetae, Anabarhynchus darembal sp. n. has black occipital macrosetae. Anabarhynchus darembal sp. n. appears to be most closely aligned to Lyneborg’s fasciatus species-group.

Figures

FIGURE 4. Anabarhynchus darembal sp. n., holotype male (ANIC _ 29: 029635): lateral view. Scale line = 2 mm.

FIGURE 5. Anabarhynchus darembal sp. n., holotype male, (ANIC _ 29: 029635): oblique view. Scale line = 2 mm.

FIGURE 6. Anabarhynchus darembal sp. n., holotype male terminalia (ANIC _ 29: 029635): A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view. Scale line = 0.5 mm.