Anabarhynchus wintertoni Ferguson

Ferguson, David J., Lambkin, Christine L. & Yeates, David K., 2014, Eight new species of Australian stiletto flies in the genus Anabarhynchus Macquart (Diptera: Therevidae) from South East Queensland, Zootaxa 3802 (4), pp. 553-582 : 578-581

Anabarhynchus wintertoni Ferguson sp. n.

( Figs 23 , 24 , 25 )

Type material. Holotype: Male. AUSTRALIA: Queensland: Carnarvon National Park; Mount Moffatt Section; Malaise Traps; Near Mount Moffatt, 19–23.I.1998, S. Winterton, J. & A. Skevington; (QM_T:185508) (QM). Condition: Pinned dorsally on stainless steel; end of abdomen removed for dissection post photography.

Paratypes: 4 females. AUSTRALIA: Queensland: 2♀ Carnarvon National Park: Mount Moffatt Section; Malaise; 25°03'49"S 148°01'57"E; J. Skevington, C. Lambkin; 1♀Carnarvon National Park: Mount Moffatt Section, One Mile Creek 25°03'49"S 148°01'57"E; J. Skevington, C. Lambkin, S. Evans; 6m grey Malaise. 1♀Carnarvon National Park: Mount Moffatt Section, Mount Moffat Rd; sandy creek; 25°04'01"S 148°00’50”E; J. Skevington, C. Lambkin, S. Evans; Malaise; (QM_T:185509-512) (QM).

Diagnosis. Frons flat, upper frontal pile appressed, short and black in two rows, lower frons pile erect; katepisternum with pile; prosternal furrow without pile; fore femur 5 pd 4–5 pv; middle femur 1–2 pd, 4–5 pv; hind femur 1 av macroseta; all femora yellow-brown, with the apicodorsal surface slightly darker; distiphallus posteriorly directed, tapered tube with apical end dorsally curved with a pair of large triangular processes along lateroventral edge of the basal half.

Description. Male. Body length: 8 mm. Wing: length 7 mm. ( Figs 23–25 ). Head. Frons flat, width 2.5 × ocellus width; ocellar tubercle flat; face and lower frons slightly protruding; parafacial grey, lower frons grey, midfrons beside eye with brown mark arching medially, upper frons brownish grey, short, black pile in two appressed rows either side of mid-line, lower frons pile half-length of scape. Scape length 2.8 × pedicel length; scape and pedicel grey; f1 brownish grey with a short black seta dorsally on basal third; f2 & f3 black, combined length halflength of f1. Occiput convex with grey brown pubescence, several indistinct rows of black macrosetae, 33–35 setae on each side; postocciput to gena with grey pubescence and dense, long, pale, hair-like pile. Palp pale yellow with pale hair-like setae; labellum grey; prementum setae black. Thorax. Scutal chaetotaxy black (pairs): np, 4; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with thin dark grey medial line within broad grey stripe, margined by narrow yellowish brown stripes. Katepisternum with pile anterodorsally; prosternal furrow without pile; pleura grey; coxae grey with long pile admixed with black macrosetae. Wing. Hyaline with pale brown infuscation, brown veins, stigma brown; costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel pale brown, knob ventrally pale brown, dorsally yellowish brown. Legs: Fore femur 5 pd distributed over apical two-thirds, 4–5 pv, weak; middle femur 1–2 pd, 4–5 pv variable strength; hind femur 1 av macroseta; with sparse black setae admixed with pale appressed pile. All femora yellow-brown, apical dorsal surface darker at angle; all femora with short appressed pale pile admixed with short black setae, ventrally erect; tibiae and tarsi yellow-brown, apically darker. Abdomen. Integument blackish brown; anterior bands tergites 2–6 blackish brown with dark appressed pile; lateral edges with grey pubescence covered with erect pale pile; apical bands pale yellow; sternites blackish with grey pubescence, apical bands yellowish white. Terminalia. (QM_T:185508); Epandrium ( Fig. 25 A): length approximately 3/4 of its width (measured along mid-line and between widest points when viewed dorsally), narrowing posteriorly. Gonocoxite ( Fig. 25 B): onion-shaped when viewed ventrally; posteroventral edge with lobed outer gonocoxal process, the apical edge directed medially with black macrosetae along lateroposterior margin and outer gonocoxal process; posterolaterally of the ventromedial plane the gonocoxal surface is distinctly indented, gonocoxite joined by hypandrium and along the ventromedial plane by a thin membrane. Gonocoxal apodeme extends beyond the anterior margin. Inner gonocoxal process darkly sclerotised, apically rounded with patch of short pile on sub-apical inner surface directed ventromedially. Gonostylus dorsoventrally broad, basal half posteriorly directed, apical half curved dorsally, apically pointed, dorsal surface a ridge along which weak pale elongate setae are directed dorsally and slightly medially, ventral edge weak pale elongate setae ventrally directed; ventral lobe translucent and discshaped. Aedeagus ( Figs 25 C, D): distiphallus posteriorly directed, sub-apically tapered and curved dorsally, with rows of small angular spines posteriorly directed along the ventral surface, an open ventral cleft extends along full length of the distiphallus; anteroventral edge with a pair of broadly triangular projections; parameral sheath dark sclerotised, broadly triangular with apical ends projected dorsally; ventral apodeme anterior broad and flat with the apical margin slightly bilobed; lateral ejaculatory apodeme broad, band-like; ejaculatory apodeme long with subapical lateral lobes.

Variation. Female, body length: 8.0– 9.5 mm. Frons width 3.2–3.3 × ocellus. Occipital macrosetae 29–32 each side. Wing length: 7–8 mm; tergites 4–7 with black erect pile laterally. Paratype female (QM_T:185512): Sternite 8 (Fig. 27F); anterior margin with deep concavity; ‘T’-shaped depressed area medially is thinly sclerotised; long, black, setae distributed anteriorly and laterally of depressed area; posteriorly with broad bilobed apex, covered with short weak setae. Furca ( Fig. 24 E): frame oval and evenly sclerotised, middle slightly waisted; anterior beam divided, with posterior division joining anterior beam in mid-line.

Etymology. The specific epithet is to honor the Dr S.L. Winterton entomologist and collector of the type specimen.

Comments. Known from five specimens collected in November and January from the Carnarvon National Park, mid-east Queensland.

Anabarhynchus wintertoni sp. n. keys to A. tauricus Lyneborg and A. yeppoon Lyneborg at couplet 57 in Lyneborg (2001). Anabarhynchus wintertoni sp. n. is readily separated from A. tauricus and A. yeppoon , both species belonging to the flavus species-group, by the shape and position of the processes on the distiphallus. In Anabarhynchus tauricus the processes are narrow and curved posteriorly and positioned basally, while in A. yeppoon the processes are spine-like and positioned sub-apically. In Anabarhynchus wintertoni sp. n. the processes on the distiphallus are broadly triangular and positioned basally. Anabarhynchus wintertoni sp. n. appears to be most closely aligned to species to Lyneborg’s flavus species-group.

Figures

FIGURE 23. Anabarhynchus wintertoni sp. n., holotype male (QM _ T: 185508): lateral view. Scale line = 2 mm.

FIGURE 24. Anabarhynchus wintertoni sp. n., holotype male, (QM _ T: 185508): oblique view. Scale line = 2 mm.

FIGURE 25. Anabarhynchus wintertoni sp. n., holotype male terminalia (QM _ T: 185508): A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view. Paratype female (QM _ T: 185512); E, furca; F, female sternite 8, ventral surface. Scale line = 0.5 mm.