Lapsias cyrboides Simon, 1900

Maddison, Wayne P., 2019, A new lapsiine jumping spider from North America, with a review of Simon's Lapsias species (Araneae, Salticidae, Spartaeinae), ZooKeys 891, pp. 17-29 : 17

Lapsias cyrboides Simon, 1900 Figs 30-40

Type material.

In MNHN, 3 males, 4 females, 3 juveniles from Colonia Tovar, Aragua State, Venezuela, in a vial with label "20924 Laps. cyrboides E.S., Tovar!" and a more recent label "det Szűts 0014". Galiano (1963) designated one male as a lectotype, in separate microvial with her label "Typus? M.E. Galiano II 1959". She mentions one female designated also as lectotype, but no female is separated and labelled as such. Because Galiano (incorrectly) designated two lectotypes, the name is not yet fixed to a single specimen. This ambiguity is resolved by designating her male lectotype as the only lectotype.

Notes.

Simon (1900) described a male and female. However, as noted by Galiano (1963), there are two species of female among the four females in the type vial, similar in body but easily distinguished by the epigyne. Two of the females ( Figs 37-40 ) have an anteriorly placed guide ( Fig. 37 ), while the other two females ( Figs 48-51 ) lack such a guide and instead show two wing-shaped atria extending laterally ( Fig. 48 ). It is reasonable to assume that these two kinds of female belong to the two smaller-bodied Lapsias at Colonia Tovar, L. cyrboides and L. tovarensis . Under L. cyrboides Simon described the female kind with anterior guide ("Plaga genitalis...longior quam latior"), but he did not justify this choice nor even mention the second form of female. Galiano followed Simon’s choice of matching female. The two forms of female are approximately the same size and carapace shape and are too faded to supply distinctive markings by which to match to the males. Nonetheless, I tentatively support Simon’s and Galiano’s matching based on an expected correlation between the form of the female’s guide and that of the male’s RTA. An anterior guide is expected to be associated with an extraordinary RTA, for instance, as in Papuamyr omhifosa Maddison & Szűts, 2019 ( Maddison and Szűts 2019). Males of L. ciliatus and L. tovarensis have small RTAs, suggesting a simple or small guide along the epigastric furrow, as is common in salticids, while the male of L. cyrboides has an unusual dorsally projecting tibial apophysis, which predicts an unusually-placed female guide. Thus, the female with anterior guide is tentatively considered that of L. cyrboides , and the female with wing-shaped atria is considered that of L. tovarensis .

Figures

Figures 30 - 40. Lapsias cyrboides lectotype male (30 - 34, 36), paralectotype male (35), and female tentatively considered of this species (37 - 40). 30 - 32 Palp 30 ventral view 31 retrolateral view of tibia 32 ventral view 33 body from above 34 body from below 35 paralectotype male body from above 36 lectotype male face 37 epigyne from below 38 vulva from above 39 female body from above 40 body from below. Scale bars: 0.1 mm (30); 1.0 mm (otherwise).

Figures 41 - 51. Lapsias tovarensis lectotype male (41 - 47) and female tentatively considered of this species (48 - 51). 41 - 43 Palp 41 ventral view 42 retrolateral view of tibia 43 ventral view 44 body from above 45 body from below 46 face 47 chelicerae from below 48 epigyne from below 49 vulva from above 50 female body from above 51 body from below. Abbreviations: c? sclerite homologous to that called the conductor in other lapsiines. Scale bars: 0.1 mm (41); 1.0 mm (otherwise).