Homoneura (Homoneura) simplicissima ( Meijere, 1910 )

Shi, Li & Yang, Ding, 2014, Supplements to species groups of the subgenus Homoneura in China (Diptera: Lauxaniidae: Homoneura), with descriptions of twenty new species, Zootaxa 3890 (1), pp. 1-117 : 41

Homoneura (Homoneura) simplicissima ( Meijere, 1910)

( Figs 260–271 )

Lauxania simplicissima Meijere, 1910: 132 . Type locality: Indonesia (Sumatra). Syntypes in HNHM (1 ♀), NNM (1 ♂, 1 ♀), ZMAN (6 ♂, 4 ♀).

Mallochomyza tagalica Frey, 1927: 38 [as syn.].

Homoneura (Homoneura) simplicissima: Malloch, 1929: 51 . Shewell in Delfinado & Hardy, 1977: 208; Sasakawa, 1992: 199.

Diagnosis. Parafacial with a row of setulae on ventral 1/2 of inner margin (top one is longest). Ray long plumose, about 1.5 times as high as 1st flagellomere. Wing with brown stripe-like spot on dm-cu. Abdomen yellow, tergites 5–6 with posterior marginal setae stretching out, almost angle 90° with tergites.

Redescription. MALE. Body length 3.2–3.6 mm, wing length 3.2–3.6 mm. FEMALE. Body length 2.5–3.8 mm, wing length 3.2–3.5 mm.

Head ( Figs 261–262 ) yellow. Frons pale brown, with yellow anterior margin and a medial stripe extending to ocellar triangle (or yellow medial stripe absent), about 1.2 times as wide as long and parallel-sided; ocellar triangle yellow, oc shorter than anterior or; anterior or shorter than posterior or. Parafacial with a row of setulae on ventral 1/2 of inner margin (top one is longest). Gena about 1/5 height of eye, with a long seta. Antenna yellow or brownish yellow, 1st flagellomere about 1.5 times as long as high; arista black except for brownish base, ray long plumose, about 1.5 times as high as 1st flagellomere. Proboscis and palpus yellow.

Thorax ( Figs 263–264 ) yellow. Mesonotum with 0+3 dc (1st post-sutural dc clearly behind transverse suture), acr in irregular 8 rows and prsc as long as 1st post-sutural dc. Scutellum yellow, apical scutellar setae crossed. Legs yellow. Fore femur with 4–5 pv, 6 pd, and ctenidium with 11–14 short setae; fore tibia with 1 long preapical ad and 1 short apv. Mid femur with 4–5 a and 1 app; mid tibia with 1 strong preapical ad and 2–3 strong apv. Hind femur with 1 weak preapical ad and a row of av; hind tibia with 1 preapical ad and 1 apv. Wing ( Fig. 266 ) with brown stripe-like spot on dm-cu; subcostal cell hyaline; costa with 2nd (between R1 and R2+3), 3rd (between R2+3 and R4+5) and 4th (between R4+5 and M1) sections in proportion of 8.7:1.4:1; r-m before middle of discal cell; ultimate and penultimate sections of M 1 in proportion of 1:1.4; ultimate section of CuA1 about 1/7 of penultimate. Halter pale yellow.

Abdomen ( Fig. 265 ) yellow, tergites 5–6 with posterior marginal setae stretching out, almost angle 90° with tergites. Male genitalia ( Figs 267–271 ): syntergosternite circular without dorsal setulae; epandrium with three pairs of long dorsal setae and broaden apically with a row of long setae; surstylus finger-like in lateral view and acute and upcurved apically in ventral view; hypandrium broad bar-like in ventral view; postgonite slender, subuliform; aedeagus gradually becoming narrow in ventral view, with a middle and distal dorsal process in lateral view; aedeagal apodeme shorter than aedeagus.

Material examined. 3 ♂, 1 ♀ ( USNM), Philippines: Puerto Princess, Palawan, R. C. McGregor (det. J. R. Malloch); 1 ♂, 1 ♀ ( USNM), Philippines: Puerto Princess, Palawan, R. C. McGregor (det. Shewell, 1987, one is marked by comp. type); 1 ♂ ( USNM), Mindanao, Kolambugan, Baker; 1 ♂ ( USNM), Philippines: Cuernos Mts. Negros, Baker.

Distribution. China ( Taiwan), Indonesia, Malaysia, Philippines.

Figures

FIGURES 260 – 266. Homoneura (Homoneura) simplicissima (Meijere, 1910). Male. 260. body, lateral view; 261, 262. head, anterior and lateral view; 263, 264. thorax, lateral and dorsal view; 265. abdomen, dorsal view; 266. wing.

FIGURES 267 – 271. Homoneura (Homoneura) simplicissima (Meijere, 1910). Male. 267. syntergosternite and epandrium, lateral view; 268. syntergosternite, anterior view; 269. epandrial complex, posterior view; 270. aedeagal complex, ventral view; 271. aedeagal complex, lateral view. Scale bar = 0.1 mm.